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<document id="55ED5939B3E9B818CE57CA4F6B5EA9DD" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFC3FFF7FF04FD61FE21F87D" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis tirio Machado, 2009, sp. nov." docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="27" updateTime="1698585892014" updateUser="plazi">
<mods:mods id="FC689C19B047B4500E47EBB07DD6A0AC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4B42260E33E63D0679BBC51DCA559A48">
<mods:title id="4938127DDB5EF495632379C2E86018EC">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
</mods:titleInfo>
<mods:name id="ABCED4D876F521E5D846D555356477CD" type="personal">
<mods:role id="976D46871B5DD2CE6593CC87E6ED47F6">
<mods:roleTerm id="ABF10EC9ABFA2889F09DDEACC6762E2A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B307FE950B295A1BABE732E1D0F25ACF">Machado, Angelo B. M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C90F5CFC723BC5019D3391275D64054A">text</mods:typeOfResource>
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<mods:title id="3AC9CCC04F47F6DA77AAC7FAFC61EA4C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E57DF57C2639676760EB1EAC4963F17F">
<mods:date id="F8102BC1F923137E8160FAB013DA3986">2009</mods:date>
<mods:detail id="AB56B6B7686B1FE82A970E7F3622C535" type="volume">
<mods:number id="00E98F325B5A4700037F47D1587CB030">2108</mods:number>
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<mods:classification id="E983026E7DE64D6F7F675CE9476670B0">journal article</mods:classification>
<mods:identifier id="DDDA9971C075A12F23C1014FCCDA76ED" type="DOI">10.5281/zenodo.187806</mods:identifier>
<mods:identifier id="283E2C56873EF2AC509B534EF1D66A5D" type="GBIF-Dataset">9a495f32-3eb8-4b35-bd46-adca3b5db2d2</mods:identifier>
<mods:identifier id="931B4E596FCE6E5261858ECDDE35CBEC" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="480B60AEAD101AE98A9EB5077DF353BD" type="Zenodo-Dep">187806</mods:identifier>
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<treatment id="1312774BFFC3FFF7FF04FD61FE21F87D" ID-DOI="http://doi.org/10.5281/zenodo.4391383" ID-GBIF-Taxon="119617263" ID-Zenodo-Dep="4391383" LSID="urn:lsid:plazi:treatment:1312774BFFC3FFF7FF04FD61FE21F87D" httpUri="http://treatment.plazi.org/id/1312774BFFC3FFF7FF04FD61FE21F87D" lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27">
<subSubSection id="D3A195D6FFC3FFF1FF04FD61FE48FD65" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="9B04C65DFFC3FFF1FF04FD61FE49FD06" blockId="26.[151,495,646,707]" box="[151,494,646,672]" pageId="26" pageNumber="27">
<heading id="C04C7131FFC3FFF1FF04FD61FE49FD06" bold="true" box="[151,494,646,672]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="A9CF1A4FFFC3FFF1FF04FD61FE49FD06" bold="true" box="[151,494,646,672]" pageId="26" pageNumber="27">
<taxonomicName id="5CBBBDDEFFC3FFF1FF04FD61FE2EFD06" ID-CoL="7CY9H" box="[151,393,646,672]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="tirio" status="sp. nov.">
<emphasis id="A9CF1A4FFFC3FFF1FF04FD61FE2EFD06" bold="true" box="[151,393,646,672]" italics="true" pageId="26" pageNumber="27">Tuberculobasis tirio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFC3FFF1FE03FD61FE49FD06" box="[400,494,646,672]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF04FD4EFE48FD65" blockId="26.[151,495,646,707]" box="[151,495,681,707]" pageId="26" pageNumber="27">
<figureCitation id="0380DAD8FFC3FFF1FF04FD4EFEE5FD65" box="[151,322,681,707]" captionStart="FIGURES 35 – 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 – 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="26" pageNumber="27">Figures 41–42</figureCitation>
,
<figureCitation id="0380DAD8FFC3FFF1FEDDFD4EFEC9FD65" box="[334,366,681,707]" captionStart="FIGURES 47 – 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="26" pageNumber="27">51</figureCitation>
,
<figureCitation id="0380DAD8FFC3FFF1FEE9FD4EFE61FD65" box="[378,454,681,707]" captionStart="FIGURES 78 – 86" captionStartId="27.[151,269,1886,1910]" captionTargetBox="[175,1436,638,1834]" captionTargetId="figure@27.[151,1436,572,1958]" captionTargetPageId="27" captionText="FIGURES 78 – 86. Tuberculobasis, male, anal appendages in dorsal, lateral, and posterior view: T. mammilaris, lectotype (78 – 80); T. tirio, holotype (81 – 83); T. yanomami, holotype (84), paratype (85 – 86) [78 – 80, 85 – 86 by R. W. Garrison]." httpUri="https://zenodo.org/record/187819/files/figure.png" pageId="26" pageNumber="27">81–83</figureCitation>
,
<figureCitation id="0380DAD8FFC3FFF1FE41FD4EFE48FD65" box="[466,495,681,707]" captionStart="FIGURES 87 – 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 – 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="26" pageNumber="27">92</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC3FFF1FF04FD11FD9DFC91" pageId="26" pageNumber="27" type="etymology">
<paragraph id="9B04C65DFFC3FFF1FF04FD11FD9DFC91" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF04FD11FE8FFCB6" bold="true" box="[151,296,758,784]" pageId="26" pageNumber="27">Etymology:</emphasis>
The name refers to the
<taxonomicName id="5CBBBDDEFFC3FFF1FDD5FD11FD27FCB6" box="[582,640,758,784]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="34" phylum="Arthropoda" rank="species" species="tirio">Tirio</taxonomicName>
indian people, who helped the author’s collecting activities during his one- month visit to their village.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC3FFF1FF55FCA4FCEFFC0D" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="9B04C65DFFC3FFF1FF55FCA4FCEFFC0D" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FCA4FE2CFCFB" bold="true" box="[198,395,835,861]" pageId="26" pageNumber="27">
<typeStatus id="440078FFFFC3FFF1FF55FCA4FEA4FCFB" box="[198,259,835,861]" pageId="26" pageNumber="27">Type</typeStatus>
specimens
</emphasis>
:
<emphasis id="A9CF1A4FFFC3FFF1FE08FCA4FDAAFCFB" bold="true" box="[411,525,835,861]" pageId="26" pageNumber="27">
<typeStatus id="440078FFFFC3FFF1FE08FCA4FDAAFCFB" box="[411,525,835,861]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
</emphasis>
(3) and
<emphasis id="A9CF1A4FFFC3FFF1FDE9FCA4FD7AFCFB" bold="true" box="[634,733,835,861]" pageId="26" pageNumber="27">allotype</emphasis>
(Ƥ):
<collectingCountry id="E3AC86CDFFC3FFF1FCB2FCA3FC29FCF8" box="[801,910,836,862]" name="Brazil" pageId="26" pageNumber="27">BRAZIL</collectingCountry>
, Pará State, Tumucumaque Indian Park, two days walk east from the
<taxonomicName id="5CBBBDDEFFC3FFF1FE2EFC8DFE50FC22" box="[445,503,874,900]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="26" pageNumber="34" phylum="Arthropoda" rank="species" species="tirio">Tirió</taxonomicName>
Indian village (temporary pool in savannah
<date id="EF05E09DFFC3FFF1FB9EFC8DFBCDFC22" box="[1037,1130,874,900]" pageId="26" pageNumber="27" value="1963-02">II-1963</date>
), Machado &amp; Acewa leg. (
<collectionCode id="FDAA5E98FFC3FFF1FF0CFC76FEA4FC0D" box="[159,259,913,939]" pageId="26" pageNumber="27">ABMM</collectionCode>
).
<emphasis id="A9CF1A4FFFC3FFF1FE85FC77FE2FFC0C" bold="true" box="[278,392,912,938]" pageId="26" pageNumber="27">
<typeStatus id="440078FFFFC3FFF1FE85FC77FE2FFC0C" box="[278,392,912,938]" pageId="26" pageNumber="27" type="paratype">Paratype</typeStatus>
</emphasis>
(1 3) same data as
<typeStatus id="440078FFFFC3FFF1FDE2FC76FD70FC0D" box="[625,727,913,939]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
(
<collectionCode id="FDAA5E98FFC3FFF1FD74FC76FC9BFC0D" box="[743,828,913,939]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="26" pageNumber="27">MNRJ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC3FFF0FF55FC50FD58FE3C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="9B04C65DFFC3FFF1FF55FC50FDBFFC74" blockId="26.[151,1437,758,2025]" box="[198,536,951,978]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FC50FEF9FC77" bold="true" box="[198,350,951,977]" pageId="26" pageNumber="27">Description.</emphasis>
Male
<typeStatus id="440078FFFFC3FFF1FE3AFC5FFDB4FC74" box="[425,531,952,978]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55FC39FE10FBE0" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FC39FEAEFC5E" bold="true" box="[198,265,990,1016]" pageId="26" pageNumber="27">Head</emphasis>
. Labium yellowish white. Genae and base of mandibles dark brown. Labrum, anteclypeus, postclypeus, and antefrons black. Top of head black. Postocular spot and area behind eyes orange brown. Rear of head yellowish white.
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55FBB5FF42FA97" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FBB5FE8DFBCA" bold="true" box="[198,298,1106,1132]" pageId="26" pageNumber="27">Thorax.</emphasis>
Prothorax. Pronotum brownish yellow. Propleuron greyish blue. Pterothorax with bluish grey or silver stripes as follows: one on the mesepisternum adjacent to the humeral suture, another on metepisternum continuing on to mesinfrepisternum and a third one on metepimeron. Legs yellowish, wings hyaline, venation brown, pterostigma light brown surrounded with yellow. Px in FW 11, in HW 9–10 R
<quantity id="5C436BB8FFC3FFF1FBEDFB21FB0EFB46" box="[1150,1193,1222,1248]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="26" pageNumber="27" unit="in" value="3.0">3 in</quantity>
FW originating near Px 5, in HW near 4.. Petiolation distal to Ac by a distance equal (50%) or ½ as long (50%) as Ac in both wings.
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55FADAFCBDFAD8" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FADAFEEFFAF1" bold="true" box="[198,328,1341,1367]" pageId="26" pageNumber="27">Abdomen.</emphasis>
S1–2 dorsally brown laterally bluish gray. S3–7 dorsally brown ventro-laterally brownish yellow. S8–10, cercus, and paraproct brownish orange.
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55FA6DFAC1F9C0" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55FA6DFE72FA02" bold="true" box="[198,469,1418,1444]" pageId="26" pageNumber="27">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFC3FFF1FE75FA6DFD86FA02" box="[486,545,1418,1444]" name="India" pageId="26" pageNumber="27">Hind</collectingCountry>
prothoracic lobe with well-developed lateral lobes (
<figureCitation id="0380DAD8FFC3FFF1FB04FA6DFB56FA02" box="[1175,1265,1418,1444]" captionStart="FIGURES 35 – 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 – 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="26" pageNumber="27">Fig. 41</figureCitation>
); median lobe projected caudally, subtriangular, with borders two-lipped. Mesepisternal tubercles low (0.64 mm), in dorsal view rounded, with bases well separated from mid-dorsal carina (
<figureCitation id="0380DAD8FFC3FFF1FC33FA3FFC5EFA54" box="[928,1017,1496,1522]" captionStart="FIGURES 35 – 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 – 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="26" pageNumber="27">Fig. 41</figureCitation>
). Cercus subquadrangular in dorsal view (
<figureCitation id="0380DAD8FFC3FFF1FF73FA18FE90F9BF" box="[224,311,1535,1561]" captionStart="FIGURES 78 – 86" captionStartId="27.[151,269,1886,1910]" captionTargetBox="[175,1436,638,1834]" captionTargetId="figure@27.[151,1436,572,1958]" captionTargetPageId="27" captionText="FIGURES 78 – 86. Tuberculobasis, male, anal appendages in dorsal, lateral, and posterior view: T. mammilaris, lectotype (78 – 80); T. tirio, holotype (81 – 83); T. yanomami, holotype (84), paratype (85 – 86) [78 – 80, 85 – 86 by R. W. Garrison]." httpUri="https://zenodo.org/record/187819/files/figure.png" pageId="26" pageNumber="27">Fig. 81</figureCitation>
); ventral process in posterior view with medial margin straight (
<figureCitation id="0380DAD8FFC3FFF1FBBEFA18FB23F9BF" box="[1069,1156,1535,1561]" captionStart="FIGURES 78 – 86" captionStartId="27.[151,269,1886,1910]" captionTargetBox="[175,1436,638,1834]" captionTargetId="figure@27.[151,1436,572,1958]" captionTargetPageId="27" captionText="FIGURES 78 – 86. Tuberculobasis, male, anal appendages in dorsal, lateral, and posterior view: T. mammilaris, lectotype (78 – 80); T. tirio, holotype (81 – 83); T. yanomami, holotype (84), paratype (85 – 86) [78 – 80, 85 – 86 by R. W. Garrison]." httpUri="https://zenodo.org/record/187819/files/figure.png" pageId="26" pageNumber="27">Fig. 83</figureCitation>
), in lateral view shaped like an isosceles triangle tapering into a narrow, straight tip (
<figureCitation id="0380DAD8FFC3FFF1FCFBF9C2FC66F999" box="[872,961,1573,1599]" captionStart="FIGURES 78 – 86" captionStartId="27.[151,269,1886,1910]" captionTargetBox="[175,1436,638,1834]" captionTargetId="figure@27.[151,1436,572,1958]" captionTargetPageId="27" captionText="FIGURES 78 – 86. Tuberculobasis, male, anal appendages in dorsal, lateral, and posterior view: T. mammilaris, lectotype (78 – 80); T. tirio, holotype (81 – 83); T. yanomami, holotype (84), paratype (85 – 86) [78 – 80, 85 – 86 by R. W. Garrison]." httpUri="https://zenodo.org/record/187819/files/figure.png" pageId="26" pageNumber="27">Fig. 82</figureCitation>
). Paraproct subequal to cercus. Penis as described for genus, but filiform process of internal fold extends beyond lateral border of penis (
<figureCitation id="0380DAD8FFC3FFF1FA93F9ABFAFEF9C0" box="[1280,1369,1612,1638]" captionStart="FIGURES 87 – 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 – 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="26" pageNumber="27">Fig. 92</figureCitation>
).
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55F995FB7EF92A" blockId="26.[151,1437,758,2025]" box="[198,1241,1650,1676]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55F995FE0CF92A" bold="true" box="[198,427,1650,1676]" pageId="26" pageNumber="27">Dimensions (mm).</emphasis>
Abdomen 26.77–27.50 (mean 26.91); HW 14.8–15.73 (mean 15.32).
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55F97EFD84F915" blockId="26.[151,1437,758,2025]" box="[198,547,1689,1715]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55F97EFEF9F915" bold="true" box="[198,350,1689,1715]" pageId="26" pageNumber="27">Description.</emphasis>
Female
<typeStatus id="440078FFFFC3FFF1FE2FF97EFD84F915" box="[444,547,1689,1715]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55F927FD61F83D" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55F927FEB7F97C" bold="true" box="[198,272,1728,1754]" pageId="26" pageNumber="27">Head.</emphasis>
Labium yellowish. Labrum brown surrounded by yellow. Base of mandibles and genae yellow; anteclypeus and postclypeus black; antefrons and top of head dominantly brownish orange, with a black band adjoining the eye dilated posteriorly to form a large subtriangular black area connected with that of the opposite side by a narrow black stripe at occipital bar. A rounded blue postocular spot within black area. A transverse orange stripe at postocular area connecting with that of the opposite side through occipital bar. Antennae brown, the two basal segments black.
</paragraph>
<paragraph id="9B04C65DFFC3FFF1FF55F84FFB74F84F" blockId="26.[151,1437,758,2025]" pageId="26" pageNumber="27">
<emphasis id="A9CF1A4FFFC3FFF1FF55F84FFE8DF864" bold="true" box="[198,298,1960,1986]" pageId="26" pageNumber="27">Thorax.</emphasis>
Color including legs and wings, as in the male. Px in FW 12, in HW 11; R
<quantity id="5C436BB8FFC3FFF1FB22F84FFB79F864" box="[1201,1246,1960,1986]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="26" pageNumber="27" unit="in" value="3.0">3 in</quantity>
FW originating near Px 6, in HW near 5. Petiolation distal to Ac by a distance equal to length in both wings.
</paragraph>
<paragraph id="9B04C65DFFC2FFF0FF55FF70FF4BFF59" blockId="27.[151,1436,151,564]" pageId="27" pageNumber="28">
<emphasis id="A9CF1A4FFFC2FFF0FF55FF70FEE7FF17" bold="true" box="[198,320,151,177]" pageId="27" pageNumber="28">Abdomen</emphasis>
. S1 dorsally orange, laterally bluish grey, S2 and proximal 1/3 of S3 dorsally brown, laterally bluish grey. Distal part of S3, and S4–8 dorsally brown, ventro-laterally brownish yellow. S9–10 brownish yellow.
</paragraph>
<paragraph id="9B04C65DFFC2FFF0FF55FEECFB21FED5" blockId="27.[151,1436,151,564]" pageId="27" pageNumber="28">
<emphasis id="A9CF1A4FFFC2FFF0FF55FEECFE73FE83" bold="true" box="[198,468,267,293]" pageId="27" pageNumber="28">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFC2FFF0FE76FEEBFD87FE80" box="[485,544,268,294]" name="India" pageId="27" pageNumber="28">Hind</collectingCountry>
prothoracic lobe with a semicircular and depressed median lobe separated from lateral lobes by two deep concavities (
<figureCitation id="0380DAD8FFC2FFF0FD09FED5FD55FEEA" box="[666,754,306,332]" captionStart="FIGURES 47 – 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="27" pageNumber="28">Fig. 51</figureCitation>
).
<collectingCountry id="E3AC86CDFFC2FFF0FC94FED5FCE5FEEA" box="[775,834,306,332]" name="India" pageId="27" pageNumber="28">Hind</collectingCountry>
margin of mesostigmal plate curved anteriorly and then laterally forming a C-shaped structure (
<figureCitation id="0380DAD8FFC2FFF0FD0DFEBEFD51FED5" box="[670,758,345,371]" captionStart="FIGURES 47 – 54" captionStartId="20.[151,272,1755,1779]" captionTargetBox="[188,1374,193,1754]" captionTargetId="figure@20.[180,1408,117,1857]" captionTargetPageId="20" captionText="FIGURES 47 – 54. Tuberculobasis female thorax, in dorsal view: T. cardinalis, holotype (47); T. costalimai, allotype (48); T. inversa, Peru, Loreto (49); T. mammilaris, Venezuela (50); T. tirio, allotype (51); T. williamsoni, allotype (52); T. yanomami, paratype (53), and in dorso-lateral view: Tuberculobasis sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels]." httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="27" pageNumber="28">Fig. 51</figureCitation>
). Mesepisternal tubercles lacking.
</paragraph>
<paragraph id="9B04C65DFFC2FFF0FF55FE98FD58FE3C" blockId="27.[151,1436,151,564]" box="[198,767,383,410]" pageId="27" pageNumber="28">
<emphasis id="A9CF1A4FFFC2FFF0FF55FE98FE0DFE3F" bold="true" box="[198,426,383,409]" pageId="27" pageNumber="28">Dimensions (mm).</emphasis>
Abdomen 30.71; HW 18.82.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFC2FFF7FF55FE41FE21F87D" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
<paragraph id="9B04C65DFFC2FFF0FF55FE41FB06FD92" blockId="27.[151,1436,151,564]" pageId="27" pageNumber="28">
<emphasis id="A9CF1A4FFFC2FFF0FF55FE41FE9FFE66" bold="true" box="[198,312,422,448]" pageId="27" pageNumber="28">Remarks</emphasis>
.
<taxonomicName id="5CBBBDDEFFC2FFF0FED8FE40FE3CFE66" box="[331,411,423,448]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tirio">
<emphasis id="A9CF1A4FFFC2FFF0FED8FE40FE3CFE66" box="[331,411,423,448]" italics="true" pageId="27" pageNumber="28">T. tirio</emphasis>
</taxonomicName>
belongs in the
<taxonomicName id="5CBBBDDEFFC2FFF0FDF3FE40FD4BFE66" box="[608,748,423,448]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mammilaris">
<emphasis id="A9CF1A4FFFC2FFF0FDF3FE40FD4BFE66" box="[608,748,423,448]" italics="true" pageId="27" pageNumber="28">mammilaris</emphasis>
</taxonomicName>
species group together with
<emphasis id="A9CF1A4FFFC2FFF0FBC3FE40FE8AFE40" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="5CBBBDDEFFC2FFF0FBC3FE40FB7FFE66" box="[1104,1240,423,448]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="geijskesi">T. geijskesi</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC2FFF0FB7BFE40FAD0FE66" box="[1256,1399,423,448]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="karitiana">T. karitiana</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC2FFF0FA15FE40FE8AFE40" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
</emphasis>
, and
<taxonomicName id="5CBBBDDEFFC2FFF0FEE6FE2AFDBFFE40" box="[373,536,461,486]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yanomami">
<emphasis id="A9CF1A4FFFC2FFF0FEE6FE2AFDBFFE40" box="[373,536,461,486]" italics="true" pageId="27" pageNumber="28">T. yanomami</emphasis>
</taxonomicName>
. Male is very close to
<emphasis id="A9CF1A4FFFC2FFF0FCD3FE2AFBA7FE40" box="[832,1024,461,486]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="5CBBBDDEFFC2FFF0FCD3FE2AFC5CFE40" box="[832,1019,461,486]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
,
</emphasis>
but female is more similar to
<taxonomicName id="5CBBBDDEFFC2FFF0FA16FE2AFEB5FDAB" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yanomami">
<emphasis id="A9CF1A4FFFC2FFF0FA16FE2AFEB5FDAB" italics="true" pageId="27" pageNumber="28">T. yanomami</emphasis>
</taxonomicName>
. Differences are given in the keys. Specimens were collected at temporary pools in the savannah. The ecological significance of this habitat is discussed under Ecological Considerations.
</paragraph>
<caption id="CFC496D5FFC2FFF0FF04F8B9FAD3F83E" httpUri="https://zenodo.org/record/187819/files/figure.png" pageId="27" pageNumber="28" targetBox="[175,1436,638,1834]" targetPageId="27">
<paragraph id="9B04C65DFFC2FFF0FF04F8B9FAD3F83E" blockId="27.[151,1436,1886,1944]" pageId="27" pageNumber="28">
<emphasis id="A9CF1A4FFFC2FFF0FF04F8B9FEFAF8D0" bold="true" box="[151,349,1886,1910]" pageId="27" pageNumber="28">FIGURES 78–86.</emphasis>
<emphasis id="A9CF1A4FFFC2FFF0FEF0F8B9FDABF8D3" box="[355,524,1886,1909]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="5CBBBDDEFFC2FFF0FEF0F8B9FDAFF8D3" box="[355,520,1886,1909]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">Tuberculobasis</taxonomicName>
,
</emphasis>
male, anal appendages in dorsal, lateral, and posterior view:
<emphasis id="A9CF1A4FFFC2FFF0FB07F8B9FA93F8D3" box="[1172,1332,1886,1909]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="5CBBBDDEFFC2FFF0FB07F8B9FA97F8D3" box="[1172,1328,1886,1909]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
,
</emphasis>
lectotype (78–80);
<emphasis id="A9CF1A4FFFC2FFF0FF6AF866FEEFF83E" box="[249,328,1921,1944]" italics="true" pageId="27" pageNumber="28">
<taxonomicName id="5CBBBDDEFFC2FFF0FF6AF866FEE3F83E" box="[249,324,1921,1944]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tirio">T. tirio</taxonomicName>
,
</emphasis>
holotype (81–83);
<taxonomicName id="5CBBBDDEFFC2FFF0FD86F866FD39F83E" box="[533,670,1921,1944]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="yanomami">
<emphasis id="A9CF1A4FFFC2FFF0FD86F866FD39F83E" box="[533,670,1921,1944]" italics="true" pageId="27" pageNumber="28">T. yanomami</emphasis>
</taxonomicName>
, holotype (84), paratype (85–86) [78–80, 85–86 by R.W. Garrison].
</paragraph>
</caption>
<caption id="CFC496D5FFC5FFF7FF04FF70FB79FF09" box="[151,1246,151,175]" pageId="28" pageNumber="29">
<paragraph id="9B04C65DFFC5FFF7FF04FF70FB79FF09" blockId="28.[151,1437,151,2011]" box="[151,1246,151,175]" pageId="28" pageNumber="29">
<emphasis id="A9CF1A4FFFC5FFF7FF04FF70FEAFFF09" bold="true" box="[151,264,151,175]" pageId="28" pageNumber="29">TABLE 3.</emphasis>
Diagnostic characters for
<taxonomicName id="5CBBBDDEFFC5FFF7FDB0FF7FFD62FF09" box="[547,709,152,175]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="A9CF1A4FFFC5FFF7FDB0FF7FFD62FF09" box="[547,709,152,175]" italics="true" pageId="28" pageNumber="29">Tuberculobasis</emphasis>
</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC5FFF7FD41FF7FFCD5FF09" box="[722,882,152,175]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="A9CF1A4FFFC5FFF7FD41FF7FFCD5FF09" box="[722,882,152,175]" italics="true" pageId="28" pageNumber="29">Denticulobasis</emphasis>
</taxonomicName>
,
<taxonomicName id="5CBBBDDEFFC5FFF7FC13FF7FFC54FF09" box="[896,1011,152,175]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="A9CF1A4FFFC5FFF7FC13FF7FFC54FF09" box="[896,1011,152,175]" italics="true" pageId="28" pageNumber="29">Leptobasis</emphasis>
</taxonomicName>
and
<taxonomicName id="5CBBBDDEFFC5FFF7FBB4FF7FFB77FF09" box="[1063,1232,152,175]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="34" phylum="Arthropoda" rank="genus">Metaleptobasis</taxonomicName>
*.
</paragraph>
</caption>
<paragraph id="9B04C65DFFC5FFF7FF0CFF59FA94FF73" blockId="28.[151,1437,151,2011]" box="[159,1331,190,213]" pageId="28" pageNumber="29">
Characters
<emphasis id="A9CF1A4FFFC5FFF7FE16FF59FA94FF73" box="[389,1331,190,213]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FE16FF59FD80FF73" box="[389,551,190,213]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Tuberculobasis</taxonomicName>
<taxonomicName id="5CBBBDDEFFC5FFF7FD0FFF59FB85FF73" authority="Metaleptobasis" authorityName="Metaleptobasis" box="[668,1058,190,213]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Leptobasis Metaleptobasis</taxonomicName>
<taxonomicName id="5CBBBDDEFFC5FFF7FB00FF59FA94FF73" box="[1171,1331,190,213]" class="Insecta" family="Coenagrionidae" genus="Denticulobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Denticulobasis</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFF07FB54FF51" blockId="28.[151,1437,151,2011]" box="[159,1267,224,247]" pageId="28" pageNumber="29">Frons Rounded Rounded Angulated Rounded</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFEE6FA81FEBE" blockId="28.[151,1437,151,2011]" box="[159,1318,257,280]" pageId="28" pageNumber="29">Pale postocular Present 97.7%, absent Present 53.2%, Absent 100% Present 100%</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFEC3FD46FEFB" blockId="28.[151,1437,151,2011]" pageId="28" pageNumber="29">spots* 2.23% absent 46.8% n = 25 n = 7 n = 45 n = 32</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFE8FFAC0FED9" blockId="28.[151,1437,151,2011]" box="[159,1383,360,383]" pageId="28" pageNumber="29">Median lobe of Distinguishable and Poorly Mostly distinguishable Distinguishable and</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFE6DFAF7FE07" blockId="28.[151,1437,151,2011]" box="[159,1360,394,417]" pageId="28" pageNumber="29">hind prothoracic projected caudally distinguishable and but not projected projectedcaudally</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFE4AFC7EFE62" blockId="28.[151,1437,151,2011]" box="[159,985,429,452]" pageId="28" pageNumber="29">lobe in males not projected caudally</paragraph>
<paragraph id="9B04C65DFFC5FFF7FD0FFE37FD51FE41" blockId="28.[151,1437,151,2011]" box="[668,758,464,487]" pageId="28" pageNumber="29">caudally</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFE16FAC8FDAE" blockId="28.[151,1437,151,2011]" box="[159,1391,497,520]" pageId="28" pageNumber="29">Mesepisternal Present in males and Absent Present in males and Present in males and</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFDF3FB41FD8D" blockId="28.[151,1437,151,2011]" box="[159,1254,532,555]" pageId="28" pageNumber="29">processes most females females females</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFDD2FA9CFDEA" blockId="28.[151,1437,151,2011]" box="[159,1339,565,588]" pageId="28" pageNumber="29">
<typeStatus id="440078FFFFC5FFF7FF0CFDD2FF73FDEA" box="[159,212,565,588]" pageId="28" pageNumber="29">Type</typeStatus>
of Mostly horns in males, Horns in males,
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFDBFFACFFDC9" blockId="28.[151,1437,151,2011]" box="[159,1384,600,623]" pageId="28" pageNumber="29">mesepisternal Tubercles No processes tubercles in females tubercles in females</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFD9DFEA2FD37" blockId="28.[151,1437,151,2011]" box="[159,261,634,657]" pageId="28" pageNumber="29">processes</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFD7BFCCCFD15" blockId="28.[151,1437,151,2011]" box="[159,875,668,691]" pageId="28" pageNumber="29">Supplementary Well-developed but</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFD59FA9CFD73" blockId="28.[151,1437,151,2011]" box="[159,1339,702,725]" pageId="28" pageNumber="29">tooth of tarsal claws Well-developed small Absent Well-developed</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFD07FACCFD51" blockId="28.[151,1437,151,2011]" box="[159,1387,736,759]" pageId="28" pageNumber="29">Petiolation in At or slightly distal: At or slightly Distal by ½ to twice Distal by ½ to twice</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFCE5FAD5FCBF" blockId="28.[151,1437,151,2011]" box="[159,1394,770,793]" pageId="28" pageNumber="29">relation to Ac ** 11.74%, distal by ½ to distal: 100% length of Ac (100%) length of Ac l: 100%</paragraph>
<paragraph id="9B04C65DFFC5FFF7FE16FCC2FE7FFCF9" blockId="28.[151,1437,151,2011]" pageId="28" pageNumber="29">twice length of Ac: n = 133 n = 90 n = 28 88.25%</paragraph>
<paragraph id="9B04C65DFFC5FFF7FE16FC8DFE70FC27" blockId="28.[151,1437,151,2011]" box="[389,471,874,897]" pageId="28" pageNumber="29">n = 315</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFC6BFE97FC05" blockId="28.[151,1437,151,2011]" box="[159,304,908,931]" pageId="28" pageNumber="29">Pair of lateral</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFC49FB41FC63" blockId="28.[151,1437,151,2011]" box="[159,1254,942,965]" pageId="28" pageNumber="29">chitinized spines on Absent Present Absent Absent</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFC36FEF2FC4E" blockId="28.[151,1437,151,2011]" box="[159,341,977,1000]" pageId="28" pageNumber="29">distal segment of</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFC13FF7FFBAD" blockId="28.[151,1437,151,2011]" box="[159,216,1012,1035]" pageId="28" pageNumber="29">penis</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFBF2FB4FFB8A" blockId="28.[151,1437,151,2011]" box="[159,1256,1045,1068]" pageId="28" pageNumber="29">Penis internal fold Present Absent Absent Present</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFBD1FECCFBEB" blockId="28.[151,1437,151,2011]" box="[159,363,1078,1101]" pageId="28" pageNumber="29">Filiform process of</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFBBEFB4FFBD6" blockId="28.[151,1437,151,2011]" box="[159,1256,1113,1136]" pageId="28" pageNumber="29">penis internal fold Present Absent Absent Present</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFB9DFEC9FB37" blockId="28.[151,1437,151,2011]" box="[159,366,1146,1169]" pageId="28" pageNumber="29">Length of cercus in</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFB7AFB5AFB12" blockId="28.[151,1437,151,2011]" box="[159,1277,1181,1204]" pageId="28" pageNumber="29">relation to S10*** Much shorter Much shorter Subequal Subequal</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFB59FB41FB73" blockId="28.[151,1437,151,2011]" box="[159,1254,1214,1237]" pageId="28" pageNumber="29">Ventral process of Present Present Absent Absent</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFB06FF44FB5E" blockId="28.[151,1437,151,2011]" box="[159,227,1249,1272]" pageId="28" pageNumber="29">cercus</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFAE5FE8BFABF" blockId="28.[151,1437,151,2011]" box="[159,300,1282,1305]" pageId="28" pageNumber="29">Membranous</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFAC2FB4FFA9A" blockId="28.[151,1437,151,2011]" box="[159,1256,1317,1340]" pageId="28" pageNumber="29">depression at dorsal Absent Absent Absent Present</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFAAFFEF4FAF9" blockId="28.[151,1437,151,2011]" box="[159,339,1352,1375]" pageId="28" pageNumber="29">surface of cercus</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFA8EFA8DFA26" blockId="28.[151,1437,151,2011]" box="[159,1322,1385,1408]" pageId="28" pageNumber="29">Vulvar spine**** Present 98.2%, absent Present 73.3%, Absent 100% Replaced by a</paragraph>
<paragraph id="9B04C65DFFC5FFF7FE16FA6BFB60FA63" blockId="28.[151,1437,151,2011]" pageId="28" pageNumber="29">0.98% absent 26.7% n = 20 denticulated vulvar n = 57 n = 30 plate</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CFA37FAD5FA41" blockId="28.[151,1437,151,2011]" box="[159,1394,1488,1511]" pageId="28" pageNumber="29">Ovipositor Not surpassing tip of Surpassing tip of Not surpassing tip of Not surpassing tip of</paragraph>
<paragraph id="9B04C65DFFC5FFF7FE16FA15FB70F9AF" blockId="28.[151,1437,151,2011]" box="[389,1239,1522,1545]" pageId="28" pageNumber="29">cercus cercus cercus cercus</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CF9F3FA37F98D" blockId="28.[151,1437,151,2011]" box="[159,1424,1556,1579]" pageId="28" pageNumber="29">Denticles of ventral Subtriangular, high and Subtriangular, low Subrectagular, high and</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CF9D1FA9AF9EB" blockId="28.[151,1437,151,2011]" box="[159,1341,1590,1613]" pageId="28" pageNumber="29">border of ovipositor directed ventro- and directed directed ventro-</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CF9BEFAD6F9D6" blockId="28.[151,1437,151,2011]" box="[159,1393,1625,1648]" pageId="28" pageNumber="29">lateral valve in medially. Associated posteriorly. Not studied with SEM medially. Associated</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF0CF99BFACFF935" blockId="28.[151,1437,151,2011]" box="[159,1384,1660,1683]" pageId="28" pageNumber="29">S.E.M. seta long and fine tipped Associated seta seta short with apex</paragraph>
<paragraph id="9B04C65DFFC5FFF7FD0FF979FD78F97E" blockId="28.[151,1437,151,2011]" pageId="28" pageNumber="29">long and fine truncated tipped</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F917FA3AF8A1" blockId="28.[151,1437,151,2011]" box="[151,1437,1776,1799]" pageId="28" pageNumber="29">
* n = number of specimens herein described, those from the literature plus
<specimenCount id="8DBD0DD4FFC5FFF7FC75F917FBDBF8A1" box="[998,1148,1776,1799]" pageId="28" pageNumber="29" type="generic">30 specimens</specimenCount>
of
<emphasis id="A9CF1A4FFFC5FFF7FB37F917FA9CF8A1" box="[1188,1339,1776,1799]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FB37F917FA90F8A1" box="[1188,1335,1776,1799]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
,
</emphasis>
32 of
<emphasis id="A9CF1A4FFFC5FFF7FA15F917FA3AF8A1" box="[1414,1437,1776,1799]" italics="true" pageId="28" pageNumber="29">L.</emphasis>
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F8F2FC86F88A" blockId="28.[151,1437,151,2011]" box="[151,801,1813,1836]" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FF04F8F2FF50F88A" box="[151,247,1813,1836]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A9CF1A4FFFC5FFF7FF04F8F2FF50F88A" box="[151,247,1813,1836]" italics="true" pageId="28" pageNumber="29">vacillans</emphasis>
</taxonomicName>
, and
<specimenCount id="8DBD0DD4FFC5FFF7FEA1F8F2FE65F88A" box="[306,450,1813,1836]" pageId="28" pageNumber="29" type="generic">25 specimens</specimenCount>
of 10 species of
<emphasis id="A9CF1A4FFFC5FFF7FDE4F8F2FC86F88A" box="[631,801,1813,1836]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FDE4F8F2FCBAF88A" box="[631,797,1813,1836]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">Metaleptobasis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F8DEFA3BF8F6" blockId="28.[151,1437,151,2011]" box="[151,1436,1849,1872]" pageId="28" pageNumber="29">
** n = number of wings of species here described plus 80 wings of
<emphasis id="A9CF1A4FFFC5FFF7FCFAF8DEFC5EF8F6" box="[873,1017,1849,1872]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FCFAF8DEFC52F8F6" box="[873,1013,1849,1872]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
,
</emphasis>
30 of
<taxonomicName id="5CBBBDDEFFC5FFF7FBADF8DEFB1CF8F6" box="[1086,1211,1849,1872]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="A9CF1A4FFFC5FFF7FBADF8DEFB1CF8F6" box="[1086,1211,1849,1872]" italics="true" pageId="28" pageNumber="29">L. vacillans</emphasis>
</taxonomicName>
and 90 of 10 species
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F8BBFEFAF8D5" blockId="28.[151,1437,151,2011]" box="[151,349,1884,1907]" pageId="28" pageNumber="29">
of
<taxonomicName id="5CBBBDDEFFC5FFF7FF27F8BBFEF0F8D5" box="[180,343,1884,1907]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="A9CF1A4FFFC5FFF7FF27F8BBFEF0F8D5" box="[180,343,1884,1907]" italics="true" pageId="28" pageNumber="29">Metaleptobasis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F899FE0CF833" blockId="28.[151,1437,151,2011]" box="[151,427,1918,1941]" pageId="28" pageNumber="29">*** excluding distal lobe.</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F846FA3BF81E" blockId="28.[151,1437,151,2011]" box="[151,1436,1953,1976]" pageId="28" pageNumber="29">
****n = number of female specimens of species here described, those from literature plus
<specimenCount id="8DBD0DD4FFC5FFF7FBE7F846FB4DF81E" box="[1140,1258,1953,1976]" pageId="28" pageNumber="29" type="female">40 females</specimenCount>
of
<taxonomicName id="5CBBBDDEFFC5FFF7FA83F846FA3BF81E" box="[1296,1436,1953,1976]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="A9CF1A4FFFC5FFF7FA83F846FA3BF81E" box="[1296,1436,1953,1976]" italics="true" pageId="28" pageNumber="29">T. costalimai</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="9B04C65DFFC5FFF7FF04F823FE21F87D" blockId="28.[151,1437,151,2011]" box="[151,390,1988,2011]" pageId="28" pageNumber="29">
and 30 of
<emphasis id="A9CF1A4FFFC5FFF7FE91F823FE21F87D" box="[258,390,1988,2011]" italics="true" pageId="28" pageNumber="29">
<taxonomicName id="5CBBBDDEFFC5FFF7FE91F823FE26F87D" box="[258,385,1988,2011]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="vacillans">L. vacillans</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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