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<document id="1B700C0F78DF5CA5BA61C0A2688A6A04" ID-DOI="10.5281/zenodo.208624" ID-GBIF-Dataset="5614569f-41f5-49cd-b66a-bbe1d807e923" ID-ISSN="1175-5326" ID-Zenodo-Dep="208624" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460518711016" checkinUser="plazi" docAuthor="Perreau, Michel" docDate="2012" docId="F32787BE0222A002FF0558EFFE5FFD9F" docLanguage="en" docName="zt03455p088.pdf" docOrigin="Zootaxa 3455" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Catops perkovskyi Perreau, 2012, sp. n." docType="treatment" docVersion="9" lastPageNumber="84" masterDocId="0F1EFFC60223A001FF925C1EFFBFFFC0" masterDocTitle="Description of a new genus and two new species of Leiodidae (Coleoptera) from Baltic amber using phase contrast synchrotron X-ray microtomography" masterLastPageNumber="88" masterPageNumber="81" pageNumber="82" updateTime="1698314275572" updateUser="plazi">
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<mods:title id="C7C98949521EE7DFC75D922EF1300F63">Description of a new genus and two new species of Leiodidae (Coleoptera) from Baltic amber using phase contrast synchrotron X-ray microtomography</mods:title>
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<mods:namePart id="3B8B310D6598730657AE56C6C81C37E0">Perreau, Michel</mods:namePart>
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<mods:date id="4CF6247590493F5105B8E9EAA7A9437C">2012</mods:date>
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<treatment id="F32787BE0222A002FF0558EFFE5FFD9F" ID-DOI="http://doi.org/10.5281/zenodo.6174245" ID-GBIF-Taxon="119524760" ID-Zenodo-Dep="6174245" LSID="urn:lsid:plazi:treatment:F32787BE0222A002FF0558EFFE5FFD9F" httpUri="http://treatment.plazi.org/id/F32787BE0222A002FF0558EFFE5FFD9F" lastPageId="3" lastPageNumber="84" pageId="1" pageNumber="82">
<subSubSection id="339465230222A000FF0558EFFE04FACA" box="[151,443,1264,1291]" pageId="1" pageNumber="82" type="nomenclature">
<paragraph id="7B3136A80222A000FF0558EFFE04FACA" blockId="1.[151,443,1264,1291]" box="[151,443,1264,1291]" pageId="1" pageNumber="82">
<heading id="207981C40222A000FF0558EFFE04FACA" bold="true" box="[151,443,1264,1291]" fontSize="11" level="1" pageId="1" pageNumber="82" reason="1">
<emphasis id="49FAEABA0222A000FF0558EFFE04FACA" bold="true" box="[151,443,1264,1291]" pageId="1" pageNumber="82">
<taxonomicName id="BC8E4D2B0222A000FF0558EFFECDFACB" ID-CoL="RVJ4" box="[151,370,1265,1291]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
<emphasis id="49FAEABA0222A000FF0558EFFECDFACB" bold="true" box="[151,370,1265,1291]" italics="true" pageId="1" pageNumber="82">Catops perkovskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52C957C10222A000FEEB58EEFE04FACA" box="[377,443,1264,1290]" pageId="1" pageNumber="82" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="339465230222A000FF055926FEE6FAB4" pageId="1" pageNumber="82" type="materials_examined">
<paragraph id="7B3136A80222A000FF055926FEE6FAB4" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">
<emphasis id="49FAEABA0222A000FF055926FE01FA90" bold="true" box="[151,446,1335,1360]" pageId="1" pageNumber="82">
<typeStatus id="A435880A0222A000FF055926FF6DFA90" box="[151,210,1336,1360]" pageId="1" pageNumber="82">Type</typeStatus>
material.
<typeStatus id="A435880A0222A000FEC35926FE01FA90" box="[337,446,1336,1360]" pageId="1" pageNumber="82" type="holotype">Holotype</typeStatus>
</emphasis>
: 3,
<emphasis id="49FAEABA0222A000FE665929FDDAFA90" bold="true" box="[500,613,1335,1360]" pageId="1" pageNumber="82">
<collectingCountry id="039976380222A000FE665929FDE0FA90" box="[500,607,1335,1360]" name="Russia" pageId="1" pageNumber="82">RUSSIA</collectingCountry>
:
</emphasis>
amber deposit of Jantarnij, near Kaliningrad (collection of M. Perreau, Paris, n°MP002).
</paragraph>
</subSubSection>
<subSubSection id="339465230222A000FF555961FE2FF818" pageId="1" pageNumber="82" type="description">
<paragraph id="7B3136A80222A000FF555961FC0BFA1F" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">
<emphasis id="49FAEABA0222A000FF555961FEE6FA58" bold="true" box="[199,345,1407,1432]" pageId="1" pageNumber="82">Description.</emphasis>
Body length: 3.0 mm. Habitus as in
<figureCitation id="E3B52A2D0222A000FC955961FCCBFA57" box="[775,884,1407,1431]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Figs. 12</figureCitation>
. Head with fine punctation. Antenna long (onethird of body length) and slender, all antennomeres longer than wide except for disk-like eighth (
<figureCitation id="E3B52A2D0222A000FB4159BAFAA7FA7C" box="[1235,1304,1444,1468]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Fig. 3</figureCitation>
). Maxillary palps with last palpomere as long as penultimate. Eyes well developed.
</paragraph>
<paragraph id="7B3136A80222A000FF5559F3FBC5F98C" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">Pronotum 1.6 times wider than long, with lateral sides widely arcuate, not sinuate near basal angles. Anterior margin 1.4 times wider than straight basal margin. Basal angles distinct and obtuse. Punctation simple, randomly distributed, punctures small with distance between them much larger than their diameter.</paragraph>
<paragraph id="7B3136A80222A000FF555A49FC09F91C" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">
Elytra 1.6 times longer than wide, with punctation similar to that of pronotum, randomly distributed, punctures small and widely spaced, with distance between them much larger than their diameter, as in extant species of
<taxonomicName id="BC8E4D2B0222A000FF055ABEFF59F977" box="[151,230,1696,1719]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="49FAEABA0222A000FF055ABEFF59F977" box="[151,230,1696,1719]" italics="true" pageId="1" pageNumber="82">Catops</emphasis>
</taxonomicName>
. Single longitudinal stria visible: parasutural stria. Tiny short setae that accompany punctures in extant species indistinct, probably due to lack of preservation or visualization.
</paragraph>
<paragraph id="7B3136A80222A000FF555AF9FE3FF887" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">
Profemur without ventral tooth (
<figureCitation id="E3B52A2D0222A000FDDE5AF9FD27F93F" box="[588,664,1767,1791]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Fig. 4</figureCitation>
). Protibia sinuate in middle of ventral side (
<figureCitation id="E3B52A2D0222A000FB2A5AF9FABBF93F" box="[1208,1284,1767,1791]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Fig. 4</figureCitation>
). All tarsi 5 segmented. Protarsi dilated, 0.75 times as wide as apex of protibia. First two mesotarsomeres dilated. Metatarsi without any dilation.
</paragraph>
<paragraph id="7B3136A80222A000FF555B4AFB85F874" blockId="1.[151,1436,1335,2008]" pageId="1" pageNumber="82">
Aedeagus very long and slender, regularly and strongly arcuate, nearly semi-circular in lateral view (
<figureCitation id="E3B52A2D0222A000FAD55B4AFA33F8AC" box="[1351,1420,1876,1900]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Fig. 5</figureCitation>
), broken inside body at level of first third of its length by undetermined process (
<figureCitation id="E3B52A2D0222A000FBD65B69FB32F84F" box="[1092,1165,1911,1935]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1908,1931]" captionTargetBox="[151,1435,235,1874]" captionTargetId="figure@2.[151,1436,235,1874]" captionTargetPageId="2" captionText="FIGURES 1 7. External and internal morphology of Catops perkovskyi sp. n. (holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view." httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="1" pageNumber="82">Fig. 6</figureCitation>
). Apex of median lobe roughly triangular. Tegmen (both basal lamella and parameres) has not been found.
</paragraph>
<paragraph id="7B3136A80222A000FF555BDEFE2FF818" blockId="1.[151,1436,1335,2008]" box="[199,400,1984,2008]" pageId="1" pageNumber="82">Female unknown.</paragraph>
</subSubSection>
<caption id="2FF166200221A003FF055B6AFEADF80F" httpUri="https://zenodo.org/record/208625/files/figure.png" pageId="2" pageNumber="83" targetBox="[151,1435,235,1874]" targetPageId="2">
<paragraph id="7B3136A80221A003FF055B6AFEADF80F" blockId="2.[151,1436,1908,1999]" pageId="2" pageNumber="83">
<emphasis id="49FAEABA0221A003FF055B6AFE83F849" bold="true" box="[151,316,1908,1931]" pageId="2" pageNumber="83">FIGURES 17.</emphasis>
External and internal morphology of
<taxonomicName id="BC8E4D2B0221A003FD5A5B6BFC3BF84A" box="[712,900,1908,1930]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="83" phylum="Arthropoda" rank="species" species="perkovskyi" status="sp. nov.">
<emphasis id="49FAEABA0221A003FD5A5B6BFC3BF84A" box="[712,900,1908,1930]" italics="true" pageId="2" pageNumber="83">Catops perkovskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="52C957C10221A003FC1F5B6BFC79F84A" box="[909,966,1909,1930]" pageId="2" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
(holotype) by PPC-SRμCT. 1, habitus dorsal view. 2, habitus lateral view. 3, antenna. 4, front leg. 5, aedeagus, lateral view. 6, aedeagus, dorsal view. 7, apex of aedeagus, dorsal view.
</paragraph>
</caption>
<subSubSection id="339465230220A002FF555C89FEE3FF14" pageId="3" pageNumber="84" type="distribution">
<paragraph id="7B3136A80220A002FF555C89FEE3FF14" blockId="3.[151,1436,151,607]" pageId="3" pageNumber="84">
<emphasis id="49FAEABA0220A002FF555C89FEDFFF70" bold="true" box="[199,352,151,176]" pageId="3" pageNumber="84">Distribution.</emphasis>
The new species is known only by the
<typeStatus id="A435880A0220A002FC855C89FCC6FF6F" box="[791,889,151,175]" pageId="3" pageNumber="84" type="holotype">holotype</typeStatus>
from Baltic amber, from the deposit of Jantarnij, near Kaliningrad.
</paragraph>
</subSubSection>
<subSubSection id="339465230220A002FF555CFEFD22FEDC" pageId="3" pageNumber="84" type="etymology">
<paragraph id="7B3136A80220A002FF555CFEFD22FEDC" blockId="3.[151,1436,151,607]" pageId="3" pageNumber="84">
<emphasis id="49FAEABA0220A002FF555CFEFEF1FF38" bold="true" box="[199,334,224,248]" pageId="3" pageNumber="84">Etymology.</emphasis>
The species is dedicated to Evgeny Perkovsky, who was the first to study fossils in
<taxonomicName id="BC8E4D2B0220A002FB6E5CC1FAD3FF37" box="[1276,1388,223,247]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="family">Leiodidae</taxonomicName>
and described the first fossil species in this family.
</paragraph>
</subSubSection>
<subSubSection id="339465230220A002FF555D39FE5FFD9F" pageId="3" pageNumber="84" type="discussion">
<paragraph id="7B3136A80220A002FF555D39FE5FFD9F" blockId="3.[151,1436,151,607]" pageId="3" pageNumber="84">
<emphasis id="49FAEABA0220A002FF555D39FEF3FE80" bold="true" box="[199,332,295,320]" pageId="3" pageNumber="84">Discussion.</emphasis>
This species is placed in the genus
<taxonomicName id="BC8E4D2B0220A002FD4A5D36FC98FEFF" box="[728,807,296,319]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="49FAEABA0220A002FD4A5D36FC98FEFF" box="[728,807,296,319]" italics="true" pageId="3" pageNumber="84">Catops</emphasis>
</taxonomicName>
because of the absence of epistomal suture, the two first mesotarsomeres dilated, and the shape of the aedeagus: median lobe narrow and arcuate, and parameres thin. Moreover, the extremely long and arcuate median lobe of the aedeagus is characteristic of the “
<emphasis id="49FAEABA0220A002FB465D6EFA23FE48" box="[1236,1436,368,392]" italics="true" pageId="3" pageNumber="84">
<taxonomicName id="BC8E4D2B0220A002FB465D6EFA2BFE48" box="[1236,1428,368,392]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="longulus">Catops longulus</taxonomicName>
</emphasis>
species group. The elongated antennae with all antennomeres (except the eighth) longer than wide, and the absence of tubercles on the ventral side of profemora, are also consistent with a placement in this species group.
<taxonomicName id="BC8E4D2B0220A002FADE5DA6FEB0FE34" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="perkovskyi">
<emphasis id="49FAEABA0220A002FADE5DA6FEB0FE34" italics="true" pageId="3" pageNumber="84">Catops perkovskyi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="BC8E4D2B0220A002FE325DC3FD9EFE34" box="[416,545,476,500]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="49FAEABA0220A002FE325DC3FD9EFE34" box="[416,545,476,500]" italics="true" pageId="3" pageNumber="84">C. longulus</emphasis>
</taxonomicName>
by a triangular apex of the aedeagus that is shorter (compared to the total length of the aedeagus), by the sinuate internal side of the protibia (straight in
<taxonomicName id="BC8E4D2B0220A002FC5E5E1EFBF1FDD8" box="[972,1102,512,536]" class="Insecta" family="Leiodidae" genus="Catops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="49FAEABA0220A002FC5E5E1EFBF1FDD8" box="[972,1102,512,536]" italics="true" pageId="3" pageNumber="84">C. longulus</emphasis>
</taxonomicName>
) and by its smaller size. It is clear that the sinuate protibia is not an artifact due to bad preservation in amber, given that it is exactly symmetrical in the right and left protibiae.
</paragraph>
</subSubSection>
</treatment>
</document>