422 lines
36 KiB
XML
422 lines
36 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.60072" ID-GBIF-Dataset="a8375122-971c-4804-92a6-fe16e10e3224" ID-PMC="PMC8222295" ID-Pensoft-Pub="1313-2970-1044-427" ID-Pensoft-UUID="8320077CEFA5571D971EFEA66B0C6268" ID-PubMed="34183882" ID-ZooBank="9AA72654B6CC4081B557CD0BAE3939A3" ModsDocID="1313-2970-1044-427" checkinTime="1623875398087" checkinUser="pensoft" docAuthor="Allegro, Gianni & Giachino, Pier Mauro" docDate="2021" docId="DC82C0062D235FA08F17B1D83A9DD9EB" docLanguage="en" docName="ZooKeys 1044: 427-448" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.60072" docTitle="Diplocheila (Diplocheila) erwini Allegro & Giachino 2021, sp. nov." docType="treatment" docUuid="108A8C27-E70A-4322-AF94-73E2CE64DEAB" docUuidSource="ZooBank" docVersion="4" id="8320077CEFA5571D971EFEA66B0C6268" lastPageNumber="427" masterDocId="8320077CEFA5571D971EFEA66B0C6268" masterDocTitle="The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)" masterLastPageNumber="448" masterPageNumber="427" pageNumber="427" updateTime="1668150474377" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Diplocheila Brulle, 1834 in Cambodia, with descriptions of two new species (Coleoptera, Carabidae, Licinini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Allegro, Gianni</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5079-6671</mods:nameIdentifier>
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<mods:affiliation>Strada Patro 11, I- 14036 Moncalvo (AT), Italy</mods:affiliation>
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<mods:nameIdentifier type="email">gianni.allegro54@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Giachino, Pier Mauro</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1167-5447</mods:nameIdentifier>
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<mods:affiliation>Via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1044</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>427</mods:start>
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<mods:end>448</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1044.60072</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1044.60072</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1044-427</mods:identifier>
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<mods:identifier type="ZooBank">9AA72654B6CC4081B557CD0BAE3939A3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">8320077CEFA5571D971EFEA66B0C6268</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182754008" LSID="urn:lsid:zoobank.org:act:108A8C27-E70A-4322-AF94-73E2CE64DEAB" httpUri="http://treatment.plazi.org/id/DC82C0062D235FA08F17B1D83A9DD9EB" lastPageNumber="427" pageId="0" pageNumber="427">
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<subSubSection pageId="0" pageNumber="427" type="nomenclature">
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<paragraph pageId="0" pageNumber="427">
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<taxonomicName LSID="http://zoobank.org/108A8C27-E70A-4322-AF94-73E2CE64DEAB" authority="Allegro & Giachino, 2021" authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila) erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini" status="sp. nov." subGenus="Diplocheila">Diplocheila (Diplocheila) erwini</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="427">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diplocheila erwini sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427">Figures 5</figureCitation>
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<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">, 11</figureCitation>
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<figureCitation captionStart="Figures 12–17" captionStartId="F7" captionText="Figures 12 - 17. Labrum of Diplocheila species 12 D. laevigata HT 13 D. laevigotoides HT 14 D. laevigata from Cambodia 15 Diplocheila sp. from Thailand 16 D. walterrossii sp. nov. HT 17 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427">, 17</figureCitation>
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<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">, 23</figureCitation>
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<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">, 30</figureCitation>
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<figureCitation captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">, 33</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="type locality">
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<paragraph pageId="0" pageNumber="427">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="427">Cambodia, Kampong Chhnang, Khom Domnatpopol, Tonle Sap Lake.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="materials_examined">
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<paragraph pageId="0" pageNumber="427">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<materialsCitation collectingDate="2018-05-21" collectingMethod="light trap" collectorName="Rossi, Bernardi, Kong" country="Cambodia" latitude="12.237223" location="Tonle Sap Lake" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" specimenCount="♂" stateProvince="Kampong Chhnang Province" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount>♂</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang Province</collectingRegion>
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,
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<collectingMunicipality>Khom Domnatpopol</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:4A7326C6677D03AB590E5636C48D3351" country="Cambodia" latitude="12.237223" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" name="Tonle Sap Lake" stateProvince="Kampong Chhnang Province">Tonle Sap Lake</location>
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,
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<geoCoordinate degrees="12" direction="north" minutes="14" orientation="latitude" precision="15" seconds="14" value="12.237223">12°14'14"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="41" orientation="longitude" precision="15" seconds="15" value="104.6875">104°41'15"E</geoCoordinate>
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, (
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<collectingMethod>light trap</collectingMethod>
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),
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<collectingDate value="2018-05-21">21.V.2018</collectingDate>
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,
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<collectorName>Rossi</collectorName>
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,
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<collectorName>Bernardi</collectorName>
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and
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<collectorName>Kong</collectorName>
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leg. (CAl).
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</materialsCitation>
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</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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: (
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<specimenCount type="male">1 ♂</specimenCount>
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and
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<specimenCount type="female">3 ♀♀</specimenCount>
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);
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Rossi & Bernardi & Kong" country="Cambodia" latitude="12.237223" location="Tonle Sap Lake" longLatPrecision="21" longitude="104.6875" municipality="Khom Domnatpopol" specimenCount="1" specimenCount-female="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="female">1 ♀</specimenCount>
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same data as holotype
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</materialsCitation>
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;
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<materialsCitation collectingDate="2018-11-07" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" county="Rolea" latitude="12.225277" location="Toulkrolanh Village" longLatPrecision="21" longitude="104.66389" municipality="District" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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,
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<collectingCounty>Rolea</collectingCounty>
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<normalizedToken originalValue="B’ier">B'ier</normalizedToken>
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<collectingMunicipality>District</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:B511AA93BB25AF21D0B8B2631DEF3FAE" country="Cambodia" county="Rolea" latitude="12.225277" longLatPrecision="21" longitude="104.66389" municipality="District" name="Toulkrolanh Village" stateProvince="Kampong Chhnang">Toulkrolanh Village</location>
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,
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<geoCoordinate degrees="12" direction="north" minutes="13" orientation="latitude" precision="15" seconds="31" value="12.225277">12°13'31"N</geoCoordinate>
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,
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<geoCoordinate degrees="104" direction="east" minutes="39" orientation="longitude" precision="15" seconds="50" value="104.66389">104°39'50"E</geoCoordinate>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2018-11-07">7.XI.2018</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
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and
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<collectorName>V. Kong</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation collectingDate="2019-05-17" collectingMethod="light trap" collectorName="W. Rossi, V. Kong" country="Cambodia" location="Tonle Sap Lake" specimenCount="2" specimenCount-female="2" stateProvince="Kampong Chhnang" typeStatus="paratype">
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<specimenCount type="female">2 ♀♀</specimenCount>
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,
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<collectingCountry name="Cambodia">Cambodia</collectingCountry>
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,
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<collectingRegion country="Cambodia" name="Kampong Chhnang">Kampong Chhnang</collectingRegion>
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, banks of
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<location LSID="urn:lsid:plazi:treatment:DC82C0062D235FA08F17B1D83A9DD9EB:9712BFCB0D04CF69E72E5185E37AA987" country="Cambodia" name="Tonle Sap Lake" stateProvince="Kampong Chhnang">Tonle Sap Lake</location>
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,
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<collectingMethod>light trap</collectingMethod>
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,
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<collectingDate value="2019-05-17">17.V.2019</collectingDate>
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,
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<collectorName>W. Rossi</collectorName>
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and
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<collectorName>V. Kong</collectorName>
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leg. (CAl, CGi)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="diagnosis">
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<paragraph pageId="0" pageNumber="427">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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A medium-sized to small
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<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
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</taxonomicName>
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(ABL: 12-14 mm) of the
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<taxonomicName baseAuthorityName="Fabricius" baseAuthorityYear="1792" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila polita" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="polita">
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<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila polita</emphasis>
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</taxonomicName>
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group in the subgenus
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<taxonomicName authority="Diplocheila" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila (Diplocheila)" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="subGenus" subGenus="Diplocheila">Diplocheila Diplocheila</taxonomicName>
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(sensu
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<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
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). It is easily distinguished from the other species of the group with a sexsetose labrum (
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<taxonomicName lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
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<emphasis italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
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<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
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<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
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</taxonomicName>
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, and
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<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
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<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
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</taxonomicName>
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sp. nov.) by the smaller body size (≤14 mm), the narrower and almost quadrangular pronotum (transverse in the four other species), with hind angles not protruding (protruding in
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<taxonomicName lsidName="D. indus" pageId="0" pageNumber="427" rank="species" species="indus">
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<emphasis italics="true" pageId="0" pageNumber="427">D. indus</emphasis>
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</taxonomicName>
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), and by the morphology of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="427" type="description">
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<paragraph pageId="0" pageNumber="427">Description.</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Habitus</emphasis>
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: ABL: 12-14 mm (HT ♂ 13.5 mm). Body parallel-sided, moderately shiny, black with antennae and palpi piceous-brown (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Diplocheila erwini sp. nov.: habitus of holotype." figureDoi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427">5</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1044.60072.figure5" httpUri="https://binary.pensoft.net/fig/554096" pageId="0" pageNumber="427" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" pageId="0" pageNumber="427">Figure 5.</emphasis>
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<taxonomicName authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini">
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<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila erwini</emphasis>
|
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</taxonomicName>
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sp. nov.: habitus of holotype.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Head</emphasis>
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: almost quadrangular, glabrous except for the supraorbital setae, narrow in comparison with pronotum. Eyes markedly convex; a single supraorbital seta on each side. Dorsum with microsculpture not evident, only with scattered punctures visible at>100
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification; frontal impressions short and superficial. Labrum symmetrically and deeply (LR = 0.78) emarginate, with six setigerous punctures on anterior margin (4 medial equidistant + 2 lateral on lobes). Clypeus trapezoid, distinctly concave anteriorly, with 1 seta on each side at anterolateral corner. Antennae moderately long, densely pubescent from segment 4, with terminal 2 articles surpassing base of pronotum; segments elongate, the second one short, as long as a half of first. Mandibles elongate, broad, approximately similar each another (the left with apical cutting edge more concave), with scrobe well-defined and glabrous and apex blunt; terebral tooth triangular and prominent. Labial and maxillary palps fusiform, with apices narrowly truncate.
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</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Thorax</emphasis>
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: pronotum smooth, with very faint microsculpture evident at>200
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification and with scattered punctures, subquadrate (PW/PL = 1.18), widest at middle (Fig.
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<figureCitation captionStart="Figures 6–11" captionStartId="F6" captionText="Figures 6 - 11. Pronotum of Diplocheila species 6 D. laevigata HT 7 D. laevigotoides HT 8 D. laevigata from Cambodia 9 Diplocheila sp. from Thailand 10 D. walterrossii sp. nov. HT 11 D. erwini sp. nov. HT." figureDoi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427">11</figureCitation>
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). Disk moderately convex. Sides from rectilinear to hardly rounded in anterior half; rectilinear or very slightly sinuate backwards. Hind angles rounded obtuse, provided with a postero-lateral seta. Posterior margin rectilinear between basal impressions, which are linear and markedly impressed; anterior margin with front angles nearly obsolete. A single lateral seta on each side just above middle. Lateral bead continuous, separated from the discal area by a narrow groove, only scarcely dilated before hind angles. Medial longitudinal impression fine, nearly reaching anterior and posterior margins; anterior transversal impression absent.
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</paragraph>
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<paragraph pageId="0" pageNumber="427">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Elytra</emphasis>
|
||
: moderately long (EL/EW = 1.59), parallel-sided, slightly convex and flattened on disk, widest at middle, with rounded shoulders and sides delicately sinuate before apex. Surface moderately shiny; microsculpture evident only at high magnification (>100
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), consisting of fine, slightly transverse meshes. Epipleura without any distinct external plicae ("uncrossed epipleura"). Intervals moderately convex, smooth; striae deeply impressed on the whole length, distinctly punctate. Parascutellar stria present; scutellar setigerous pore present at base of stria 1, just before conjunction with stria 2. Basal margin complete. Discal setigerous punctures absent; umbilicate series of setigerous punctures continuous, not interrupted at middle. Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Ventral surface</emphasis>
|
||
(thorax and abdomen)
|
||
</emphasis>
|
||
: prosternum and proepisterna glabrous and impunctate (only with very fine punctures). Metepisterna as long as twice the width of anterior side; metepimera large, broadly rounded. Prosternal intercoxal process widely rounded and bordered at apex. Abdominal ventrites IV-VI shiny but shagreened at sides, glabrous except one pair subapical central setae; males with 2, females with 4 marginal setae at apex of ventrite VII.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Legs</emphasis>
|
||
: moderately slender. Posterior face of femora with 1 seta in profemora, 2 in mesofemora and metafemora. Metatrochanters glabrous and slightly shorter than half length of metafemora. Protibial antennal cleaning organ well developed, with 2 clip setae. Protibiae robust, with 4 or 5 outer apical spines; mesotibiae with a group of setae at middle of inner face; metatibiae longitudinally furrowed at inner face. Dorsal face of tarsomeres smooth. Male protarsomeres 1-3 distinctly dilated, slightly asymmetrical; meso- and metatarsomeres not dilated in both sexes; tarsomere 5 ventrally glabrous, dorsally with 2 apical setae; claws smooth.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="427">Male genitalia</emphasis>
|
||
: median lobe of aedeagus short and markedly swollen before apex in lateral view (Fig.
|
||
<figureCitation captionStart="Figures 20–25" captionStartId="F9" captionText="Figures 20 - 25. Aedeagus in lateral view of Diplocheila species 20 D. laevigata from Cambodia 21 Diplocheila sp. from Thailand 22 D. walterrossii sp. nov. HT 23 D. erwini sp. nov. HT 24 D. laevis from Cambodia 25 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures20-25" httpUri="https://binary.pensoft.net/fig/554100" pageId="0" pageNumber="427">23</figureCitation>
|
||
); the apical lamella shortly triangular in dorsal view, with blunt tip (Fig.
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">30</figureCitation>
|
||
), apex very slightly bent downwards in lateral view. Ostium long, in dorsal position. Right paramere oval; left paramere conchoid.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="etymology">
|
||
<paragraph pageId="0" pageNumber="427">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
This species is named, as a token of our esteem, after our late colleague Terry Erwin, a world-renowned specialist in world and tropical
|
||
<taxonomicName authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="427" rank="family">Carabidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="distribution">
|
||
<paragraph pageId="0" pageNumber="427">Distribution and ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<taxonomicName genus="Geographical" lsidName="Geographical distribution" pageId="0" pageNumber="427" rank="species" species="distribution">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Geographical distribution</emphasis>
|
||
</taxonomicName>
|
||
: this species is recorded only from the extreme south banks of the Tonle Sap Lake, Kampong Chhnang Province, Cambodia. It seems to have a more restricted
|
||
<taxonomicName genus="Geographical" lsidName="Geographical distribution" pageId="0" pageNumber="427" rank="species" species="distribution">Geographical distribution</taxonomicName>
|
||
than
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., which has been recorded from the same site as well as from other two localities in north-western Cambodia (Fig.
|
||
<figureCitation captionStart="Figure 33" captionStartId="F11" captionText="Figure 33. Distribution map of Diplocheila walterrossii sp. nov. and D. erwini sp. nov. in Cambodia." figureDoi="10.3897/zookeys.1044.60072.figure33" httpUri="https://binary.pensoft.net/fig/554102" pageId="0" pageNumber="427">33</figureCitation>
|
||
).
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Life habits</emphasis>
|
||
: the specimens of the type series were collected on lake banks by light trapping. No other data are available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="427" type="remarks">
|
||
<paragraph pageId="0" pageNumber="427">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="427">
|
||
It is difficult to assess the closest relatives of
|
||
<taxonomicName lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. It belongs to the
|
||
<taxonomicName lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
group (sensu
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
), as it shares the characters distinguishing this group and, moreover, its aedeagus is very similar to that of
|
||
<taxonomicName lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
(see pl. V, fig. 72a in
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Allegro & Giachino" authorityYear="2021" class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila erwini" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., on the other hand, is easily distinguished from
|
||
<taxonomicName lsidName="D. polita" pageId="0" pageNumber="427" rank="species" species="polita">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. polita</emphasis>
|
||
</taxonomicName>
|
||
which has a quadrisetose labrum and pronotum more transverse (PW/PL = 1.31 according to Fig.
|
||
<figureCitation captionStart="Figures 26–32" captionStartId="F10" captionText="Figures 26 - 32. Aedeagus in dorsal view of Diplocheila species 26 D. laevigata HT 27 D. laevigata from Cambodia 28 Diplocheila sp. from Thailand 29 D. walterrossii sp. nov. HT 30 D. erwini sp. nov. HT 31 D. laevis from Cambodia 32 D. latifrons from Cambodia." figureDoi="10.3897/zookeys.1044.60072.figures26-32" httpUri="https://binary.pensoft.net/fig/554101" pageId="0" pageNumber="427">26</figureCitation>
|
||
in Plate II in
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
), as well as a larger body size (ABL = 13.4-18.4) (
|
||
<bibRefCitation author="Ball, GE" journalOrPublisher="Memoirs of the American Entomological Society" pageId="0" pageNumber="427" pagination="1 - 258" refId="B5" refString="Ball, GE, 1959. A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera). Memoirs of the American Entomological Society 16: 1 - 258" title="A taxonomic study of the North American Licinini, with notes on the Old World species of the genus Diplocheila Brulle (Coleoptera)." volume="16" year="1959">Ball 1959</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1044.60072.figures6-11" httpUri="https://binary.pensoft.net/fig/554097" pageId="0" pageNumber="427" start="Figures 6–11" startId="F6">
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">Figures 6-11.</emphasis>
|
||
Pronotum of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis bold="true" pageId="0" pageNumber="427">6</emphasis>
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">7</emphasis>
|
||
<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">8</emphasis>
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis bold="true" pageId="0" pageNumber="427">9</emphasis>
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
sp. from Thailand
|
||
<emphasis bold="true" pageId="0" pageNumber="427">10</emphasis>
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">11</emphasis>
|
||
<taxonomicName lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1044.60072.figures12-17" httpUri="https://binary.pensoft.net/fig/554098" pageId="0" pageNumber="427" start="Figures 12–17" startId="F7">
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">Figures 12-17.</emphasis>
|
||
Labrum of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis bold="true" pageId="0" pageNumber="427">12</emphasis>
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">13</emphasis>
|
||
<taxonomicName lsidName="D. laevigotoides" pageId="0" pageNumber="427" rank="species" species="laevigotoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigotoides</emphasis>
|
||
</taxonomicName>
|
||
HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">14</emphasis>
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis bold="true" pageId="0" pageNumber="427">15</emphasis>
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
sp. from Thailand
|
||
<emphasis bold="true" pageId="0" pageNumber="427">16</emphasis>
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT
|
||
<emphasis bold="true" pageId="0" pageNumber="427">17</emphasis>
|
||
<taxonomicName lsidName="D. erwini" pageId="0" pageNumber="427" rank="species" species="erwini">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. erwini</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1044.60072.figures18-19" httpUri="https://binary.pensoft.net/fig/554099" pageId="0" pageNumber="427" start="Figures 18–19" startId="F8">
|
||
<paragraph pageId="0" pageNumber="427">
|
||
<emphasis bold="true" pageId="0" pageNumber="427">Figures 18-19.</emphasis>
|
||
Male protarsi of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Diplocheila" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocheila" order="Coleoptera" pageId="0" pageNumber="427" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">Diplocheila</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
<emphasis bold="true" pageId="0" pageNumber="427">18</emphasis>
|
||
<taxonomicName lsidName="D. laevigata" pageId="0" pageNumber="427" rank="species" species="laevigata">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. laevigata</emphasis>
|
||
</taxonomicName>
|
||
from Cambodia
|
||
<emphasis bold="true" pageId="0" pageNumber="427">19</emphasis>
|
||
<taxonomicName lsidName="D. walterrossii" pageId="0" pageNumber="427" rank="species" species="walterrossii">
|
||
<emphasis italics="true" pageId="0" pageNumber="427">D. walterrossii</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. HT.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |