457 lines
73 KiB
XML
457 lines
73 KiB
XML
<document id="FBF957393073F2A9832FCC6D1B9548FA" ID-CLB-Dataset="32199" ID-DOI="10.5852/ejt.2017.261" ID-GBIF-Dataset="708e152e-4450-4439-84a9-3c56148cd082" ID-Zenodo-Dep="885318" ID-ZooBank="C59239D1-0BF0-446A-AF6E-71234D01B23D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1504621962129" checkinUser="plazi" docAuthor="Reboleira, Ana Sofa P. S., Fresneda, Javier & Salgado, José Maria" docDate="2017" docId="03FB5E33666AAB42B740621DFD293D2F" docLanguage="en" docName="ejt-261.pdf.imf" docOrigin="European Journal of Taxonomy 261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Speonemadus escalerai" docType="treatment" docVersion="15" lastPageNumber="18" masterDocId="FFC2264B6665AB53B56C656DFFD63B4C" masterDocTitle="A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="16" updateTime="1698467889284" updateUser="ExternalLinkService">
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<mods:title id="B8BA55EFECB8E97F632BB05473B20B94">A new species of Speonemadus from Portugal, with the revision of the escalerai-group (Coleoptera, Leiodidae)</mods:title>
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<mods:namePart id="269478E1CC7EAB0333836D02B086B0F8">Reboleira, Ana Sofa P. S.</mods:namePart>
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<mods:namePart id="A9F2419F7B3ACAA27A1A47E2226F9DA9">Fresneda, Javier</mods:namePart>
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<mods:namePart id="6A1939D611E72F7D99FEB68612E58F35">Salgado, José Maria</mods:namePart>
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<mods:title id="8448E31FF30B3CD6F84EE2DFAA1137CA">European Journal of Taxonomy</mods:title>
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<mods:part id="F8C7E252C8543CC20D558903F6B2B0DA">
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<mods:date id="C43D0EE16D049227EEE26FD327E66BC5">2017</mods:date>
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<treatment id="03FB5E33666AAB42B740621DFD293D2F" ID-DOI="http://doi.org/10.5281/zenodo.5689576" ID-GBIF-Taxon="133183538" ID-Zenodo-Dep="5689576" LSID="urn:lsid:plazi:treatment:03FB5E33666AAB42B740621DFD293D2F" httpUri="http://treatment.plazi.org/id/03FB5E33666AAB42B740621DFD293D2F" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<subSubSection id="C348BCAE666AAB5CB740621DFBD13CC7" box="[556,1031,1904,1931]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BEDEF25666AAB5CB740621DFBD13CC7" blockId="15.[556,1031,1904,1931]" box="[556,1031,1904,1931]" pageId="15" pageNumber="16">
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<heading id="D0A55849666AAB5CB740621DFBD13CC7" box="[556,1031,1904,1931]" centered="true" fontSize="11" level="3" pageId="15" pageNumber="16" reason="4">
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<treatmentCitation id="0AF3C934666AAB5CB740621DFBD13CC7" author="Uhagon" authority="(Uhagon, 1898)" authorityName="Uhagon" authorityYear="1898" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[556,1031,1904,1931]" class="Insecta" family="Leiodidae" firstAuthor="Uhagon" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" pagination="117 - 126" phylum="Arthropoda" rank="species" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." species="escalerai" treatmentTaxon="Speonemadus escalerai" type="journal article" year="1898">
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<taxonomicName id="4C5294A6666AAB5CB740621DFBD13CC7" ID-CoL="4YHYZ" authority="(Uhagon, 1898)" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[556,1031,1904,1931]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B9263337666AAB5CB740621DFC913CC7" bold="true" box="[556,839,1904,1931]" italics="true" pageId="15" pageNumber="16">Speonemadus escalerai</emphasis>
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(
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<bibRefCitation id="EFC392D4666AAB5CB63A621CFC293CC7" author="Uhagon" box="[854,1023,1905,1931]" firstAuthor="Uhagon" pageId="15" pageNumber="16" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón, 1898</bibRefCitation>
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)
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</taxonomicName>
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</treatmentCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C348BCAE666AAB43B5D162DFFABA3AB8" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="reference_group">
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<paragraph id="8BEDEF25666AAB5CB5D162DFFD483C80" blockId="15.[189,670,1970,1996]" box="[189,670,1970,1996]" pageId="15" pageNumber="16">
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<treatmentCitation id="0AF3C934666AAB5CB5D162DFFD4C3C80" author="Uhagon" authority="Uhagon, 1898: 118" authorityName="Uhagon" authorityPageNumber="118" authorityYear="1898" box="[189,666,1970,1996]" class="Insecta" family="Leiodidae" firstAuthor="Uhagon" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" pagination="117 - 126" phylum="Arthropoda" rank="species" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." species="escalerae" treatmentTaxon="Anemadus escalerae" type="journal article" year="1898">
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<taxonomicName id="4C5294A6666AAB5CB5D162DFFD4C3C80" ID-CoL="DWRM" authority="Uhagon, 1898: 118" authorityName="Uhagon" authorityPageNumber="118" authorityYear="1898" box="[189,666,1970,1996]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="escalerae">
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<emphasis id="B9263337666AAB5CB5D162DFFE793C80" box="[189,431,1970,1996]" italics="true" pageId="15" pageNumber="16">Anemadus escalerae</emphasis>
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<bibRefCitation id="EFC392D4666AAB5CB4DA62DFFD4C3C80" author="Uhagon" box="[438,666,1970,1996]" firstAuthor="Uhagon" pageId="15" pageNumber="16" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón, 1898: 118</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BEDEF256675AB43B5D16466FC813A6A" blockId="16.[189,855,267,364]" box="[189,855,267,294]" pageId="16" pageNumber="17">
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<taxonomicName id="4C5294A66675AB43B5D16466FDBD3A69" ID-CoL="DWRM" authorityName="Uhagon" authorityYear="1898" box="[189,619,267,294]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="escalerae" subGenus="Speonemadus">
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<emphasis id="B92633376675AB43B5D16466FEE13A69" box="[189,311,267,293]" italics="true" pageId="16" pageNumber="17">Anemadus</emphasis>
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(
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<emphasis id="B92633376675AB43B4246466FE3F3A69" box="[328,489,267,293]" italics="true" pageId="16" pageNumber="17">Speonemadus</emphasis>
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)
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<emphasis id="B92633376675AB43B4976466FDBD3A69" box="[507,619,267,293]" italics="true" pageId="16" pageNumber="17">escalerae</emphasis>
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</taxonomicName>
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–
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<bibRefCitation id="EFC392D46675AB43B7E46466FC853A6A" author="Jeannel" box="[648,851,267,294]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922: 58</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BEDEF256675AB43B5D16442FC943A05" blockId="16.[189,855,267,364]" box="[189,834,302,329]" pageId="16" pageNumber="17">
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<taxonomicName id="4C5294A66675AB43B5D16442FD9B3A04" ID-CoL="4YHZ8" authorityName="Jeannel" authorityYear="1936" box="[189,589,303,329]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="escalerai" subSpecies="zariquieyi">
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<emphasis id="B92633376675AB43B5D16442FD9B3A04" box="[189,589,303,329]" italics="true" pageId="16" pageNumber="17">Speonemadus escalerai zariquieyi</emphasis>
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</taxonomicName>
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– Jeannel l936: 221.
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</paragraph>
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<paragraph id="8BEDEF256675AB43B5D1643FFD133A20" blockId="16.[189,855,267,364]" box="[189,709,338,364]" pageId="16" pageNumber="17">
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<taxonomicName id="4C5294A66675AB43B5D1643FFE043A20" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,466,338,364]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376675AB43B5D1643FFE043A20" box="[189,466,338,364]" italics="true" pageId="16" pageNumber="17">Speonemadus escalerai</emphasis>
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</taxonomicName>
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– Jeannel l936: 221.
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</paragraph>
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<paragraph id="8BEDEF256675AB43B5D164FEFABA3AB8" blockId="16.[189,1399,403,500]" pageId="16" pageNumber="17">
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The specifc epithet proposed by
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<bibRefCitation id="EFC392D46675AB43B75164FEFD393AE1" author="Uhagon" box="[573,751,403,429]" firstAuthor="Uhagon" pageId="16" pageNumber="17" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón (1898)</bibRefCitation>
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in the description was
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<taxonomicName id="4C5294A66675AB43B69564FEFBBF3AE1" ID-CoL="DWRM" authorityName="Uhagon" authorityYear="1898" box="[1017,1129,403,429]" class="Insecta" family="Leiodidae" genus="Anemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="escalerae">
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<emphasis id="B92633376675AB43B69564FEFBBF3AE1" box="[1017,1129,403,429]" italics="true" pageId="16" pageNumber="17">escalerae</emphasis>
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</taxonomicName>
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. However, the spelling
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<taxonomicName id="4C5294A66675AB43B5D164DBFEFF3A9C" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,297,438,464]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376675AB43B5D164DBFEFF3A9C" box="[189,297,438,464]" italics="true" pageId="16" pageNumber="17">escalerai</emphasis>
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</taxonomicName>
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has mostly been used in the entomological literature; since its use is prevalent, attributed to the original author, according to the
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<bibRefCitation id="EFC392D46675AB43B75064B7FD073ABF" author="ICZN" box="[572,721,473,500]" firstAuthor="ICZN" journalOrPublisher="International Code of Zoological Nomenclature. [online]." pageId="16" pageNumber="17" publicationUrl="http://www.iczn. org/iczn/index.jsp" refId="ref12229" refString="ICZN. 2016. International Code of Zoological Nomenclature. [online]. Available from http: // www. iczn. org / iczn / index. jsp [accessed 4 Feb. 2016]." type="journal article" year="2016">ICZN (2016</bibRefCitation>
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: Article 33.3.1), we propose to continue to use
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<taxonomicName id="4C5294A66675AB43B19664B7FAB03AB8" ID-CoL="4YHYZ" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[1274,1382,474,500]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="escalerai">
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<emphasis id="B92633376675AB43B19664B7FAB03AB8" box="[1274,1382,474,500]" italics="true" pageId="16" pageNumber="17">escalerai</emphasis>
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||
</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
|
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<subSubSection id="C348BCAE6675AB43B5D16776FC823E7C" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8BEDEF256675AB43B5D16776FE8B3979" blockId="16.[189,1202,539,612]" box="[189,349,539,565]" pageId="16" pageNumber="17">
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<heading id="D0A558496675AB43B5D16776FE8B3979" bold="true" box="[189,349,539,565]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="6">
|
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<emphasis id="B92633376675AB43B5D16776FE8B3979" bold="true" box="[189,349,539,565]" pageId="16" pageNumber="17">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D16727FB643928" blockId="16.[189,1202,539,612]" box="[189,1202,585,612]" pageId="16" pageNumber="17">
|
||
“Cuevas de la Zarza, del Seguret y del Encomat, cerca de Bocairente” (
|
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<bibRefCitation id="EFC392D46675AB43B16C6724FB733928" author="Uhagon" box="[1024,1189,585,612]" firstAuthor="Uhagon" pageId="16" pageNumber="17" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón 1898</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D167E6FE7C39E9" blockId="16.[189,426,651,677]" box="[189,426,651,677]" pageId="16" pageNumber="17">
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<heading id="D0A558496675AB43B5D167E6FE7C39E9" bold="true" box="[189,426,651,677]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
|
||
<emphasis id="B92633376675AB43B5D167E6FE7C39E9" bold="true" box="[189,426,651,677]" pageId="16" pageNumber="17">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D167D7FD513871" blockId="16.[189,1399,696,830]" pageId="16" pageNumber="17">
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<materialsCitation id="3B3AE5786675AB43B5D167D7FD513871" ID-GBIF-Occurrence="1609370446" collectingDate="1976-05-25" collectingDateMax="2005-04-06" collectingDateMin="1976-05-25" collectionCode="CMCN, CZULE" collectorName="de Barratxina & Escola & Ortuno & Zaragoza & Enguera & Sima de la Carrasquilla & Montagud" country="Spain" county="Alicante" location="Cova-Avenc de Barratxina" municipality="Xixona" pageId="16" pageNumber="17" specimenCount="11" specimenCount-female="11" stateProvince="Valencia">
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<collectingCountry id="F345AFB56675AB43B5D167D7FEC23998" box="[189,276,698,724]" name="Spain" pageId="16" pageNumber="17">SPAIN</collectingCountry>
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:
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<collectingCounty id="628C97A96675AB43B47267D4FE50399F" box="[286,390,697,723]" pageId="16" pageNumber="17">Alicante</collectingCounty>
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: 3 ♁♁,
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<specimenCount id="9D5424AC6675AB43B48A67D7FDFD3998" box="[486,555,698,724]" pageId="16" pageNumber="17" type="female">2 ♀♀</specimenCount>
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,
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<collectingMunicipality id="6B89755F6675AB43B75667D4FD45399F" box="[570,659,697,723]" pageId="16" pageNumber="17">Xixona</collectingMunicipality>
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,
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|
||
Cova-Avenc
|
||
<collectorName id="26A78AF36675AB43B65767D4FC08399F" box="[827,990,697,723]" pageId="16" pageNumber="17">de Barratxina</collectorName>
|
||
</location>
|
||
,
|
||
<date id="FFECC9E56675AB43B68767D4FB5B399F" box="[1003,1165,697,724]" pageId="16" pageNumber="17" value="1976-05-25">
|
||
<collectingDate id="EFA8300D6675AB43B68767D4FB5B399F" box="[1003,1165,697,724]" pageId="16" pageNumber="17" value="1976-05-25">25 May 1976</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8E8DB9FE6675AB43B1F767D7FB3A3998" LSID="urn:lsid:plazi:treatment:03FB5E33666AAB42B740621DFD293D2F:8E8DB9FE6675AB43B1F767D7FB3A3998" box="[1179,1260,698,724]" country="Spain" county="Alicante" municipality="Xixona" name="Comas" pageId="16" pageNumber="17" stateProvince="Valencia">Comas</location>
|
||
and
|
||
<collectorName id="26A78AF36675AB43B04467D4FAA0399F" box="[1320,1398,697,723]" pageId="16" pageNumber="17">Escolà</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED4377E06675AB43B59767B0FE8E39BB" box="[251,344,733,759]" pageId="16" pageNumber="17">CMCN</collectionCode>
|
||
); 5 ♁♁,
|
||
<specimenCount id="9D5424AC6675AB43B4A967B0FDDD39BB" box="[453,523,733,759]" pageId="16" pageNumber="17" type="female">7 ♀♀</specimenCount>
|
||
, Xixona, Peñas Roset,
|
||
<date id="FFECC9E56675AB43B64B67B0FC6C39B4" box="[807,954,733,760]" pageId="16" pageNumber="17" value="2005-04-06">
|
||
<collectingDate id="EFA8300D6675AB43B64B67B0FC6C39B4" box="[807,954,733,760]" pageId="16" pageNumber="17" value="2005-04-06">6 Apr. 2005</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A78AF36675AB43B6A467B0FBCD39BB" box="[968,1051,733,759]" pageId="16" pageNumber="17">Ortuño</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED4377E06675AB43B10F67B0FB1E39BB" box="[1123,1224,733,759]" pageId="16" pageNumber="17">CZULE</collectionCode>
|
||
). —
|
||
<collectingRegion id="499621C76675AB43B06A67B0FAA539BB" box="[1286,1395,733,759]" country="Spain" name="Comunitat Valenciana" pageId="16" pageNumber="17">Valencia</collectingRegion>
|
||
: 3 ♁♁,
|
||
<specimenCount id="9D5424AC6675AB43B478666DFE923856" box="[276,324,768,794]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
|
||
, Bocairent, Cova Sarsa,
|
||
<collectorName id="26A78AF36675AB43B71F666DFD363856" box="[627,736,768,794]" pageId="16" pageNumber="17">Zaragoza</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED4377E06675AB43B645666DFC503856" box="[809,902,768,794]" pageId="16" pageNumber="17">CMCN</collectionCode>
|
||
);
|
||
<specimenCount id="9D5424AC6675AB43B6F2666DFC1B3856" box="[926,973,768,794]" pageId="16" pageNumber="17" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectorName id="26A78AF36675AB43B6B3666DFB923856" box="[991,1092,768,794]" pageId="16" pageNumber="17">Enguera</collectorName>
|
||
,
|
||
<collectorName id="26A78AF36675AB43B13F666DFAA43856" box="[1107,1394,768,794]" pageId="16" pageNumber="17">Sima de la Carrasquilla</collectorName>
|
||
,
|
||
<date id="FFECC9E56675AB43B5D1664EFE983871" box="[189,334,803,830]" pageId="16" pageNumber="17" value="1993-03-08">
|
||
<collectingDate id="EFA8300D6675AB43B5D1664EFE983871" box="[189,334,803,830]" pageId="16" pageNumber="17" value="1993-03-08">8 Mar. 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A78AF36675AB43B436664EFE053871" box="[346,467,803,829]" pageId="16" pageNumber="17">Montagud</collectorName>
|
||
leg. (
|
||
<collectionCode id="ED4377E06675AB43B779664EFDAC3871" box="[533,634,803,829]" pageId="16" pageNumber="17">CZULE</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D16609FE593833" blockId="16.[189,399,868,895]" box="[189,399,868,895]" pageId="16" pageNumber="17">
|
||
<heading id="D0A558496675AB43B5D16609FE593833" bold="true" box="[189,399,868,895]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
|
||
<emphasis id="B92633376675AB43B5D16609FE593833" bold="true" box="[189,399,868,895]" pageId="16" pageNumber="17">Previous records</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D166FEFC823E7C" blockId="16.[189,1399,915,1328]" pageId="16" pageNumber="17">
|
||
<collectingCountry id="F345AFB56675AB43B5D166FEFEC238E1" box="[189,276,915,941]" name="Spain" pageId="16" pageNumber="17">SPAIN</collectingCountry>
|
||
: Alicante: Albaida, Castalla, Cova Melxor (
|
||
<bibRefCitation id="EFC392D46675AB43B67766FEFC4038E1" author="Blas M." box="[795,918,915,941]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B6C866FEFC3738E1" author="Blas M." box="[932,993,915,941]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B68366FEFBFB38E1" author="Blas M." box="[1007,1069,915,941]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B15666F9FABF38E1" author="Zaragoza & Sendra" box="[1082,1385,915,942]" firstAuthor="Zaragoza" pageId="16" pageNumber="17" refId="ref13879" refString="Zaragoza J. A. & Sendra A. 1988. Fauna cavernicola de la provincia de Alicante. Volumen 3: Fauna, Flora, Ciencia y Medicina. Instituto de Estudios Juan Gil-Albert. Ed. Diputacion Provincial de Alicante." type="journal volume" year="1988">Zaragoza & Sendra 1988</bibRefCitation>
|
||
); Banyeres, Avenc de Vinalogó (
|
||
<bibRefCitation id="EFC392D46675AB43B77366DBFD41389C" author="Blas M." box="[543,663,950,976]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B7CD66DBFD08389C" author="Blas M." box="[673,734,950,976]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B78466DBFCF0389C" author="Blas M." box="[744,806,950,976]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B65D66DAFB83389C" author="Zaragoza & Sendra" box="[817,1109,950,977]" firstAuthor="Zaragoza" pageId="16" pageNumber="17" refId="ref13879" refString="Zaragoza J. A. & Sendra A. 1988. Fauna cavernicola de la provincia de Alicante. Volumen 3: Fauna, Flora, Ciencia y Medicina. Instituto de Estudios Juan Gil-Albert. Ed. Diputacion Provincial de Alicante." type="journal volume" year="1988">Zaragoza & Sendra 1988</bibRefCitation>
|
||
); Onil, Cova-Avenc de la Foieta (= Cova de la Foieta, Cueva-Sima de la Foieta) (
|
||
<bibRefCitation id="EFC392D46675AB43B65766B4FC0F38BF" author="Jeannel" box="[827,985,985,1011]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B68966B4FB8838B8" author="Blas M." box="[997,1118,985,1012]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">Blas 1979</bibRefCitation>
|
||
); Onil, Cova de Galiano (
|
||
<bibRefCitation id="EFC392D46675AB43B5A96690FE2F3F5B" author="Zaragoza & Sendra" box="[197,505,1021,1047]" firstAuthor="Zaragoza" pageId="16" pageNumber="17" refId="ref13879" refString="Zaragoza J. A. & Sendra A. 1988. Fauna cavernicola de la provincia de Alicante. Volumen 3: Fauna, Flora, Ciencia y Medicina. Instituto de Estudios Juan Gil-Albert. Ed. Diputacion Provincial de Alicante." type="journal volume" year="1988">Zaragoza & Sendra 1988</bibRefCitation>
|
||
); Onil, Cova de Garbina (
|
||
<bibRefCitation id="EFC392D46675AB43B6586690FC003F5B" author="Jeannel" box="[820,982,1021,1047]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B6896690FBB43F5B" author="Blas M." box="[997,1122,1021,1047]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">Blas 1979</bibRefCitation>
|
||
); Onil, Cova Porrasses (
|
||
<bibRefCitation id="EFC392D46675AB43B5A9614DFE953F76" author="Blas M." box="[197,323,1056,1082]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B43E614DFE593F76" author="Blas M." box="[338,399,1056,1082]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B4F3614DFE0B3F76" author="Blas M." box="[415,477,1056,1082]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B481614DFCF73F76" author="Zaragoza & Sendra" box="[493,801,1056,1082]" firstAuthor="Zaragoza" pageId="16" pageNumber="17" refId="ref13879" refString="Zaragoza J. A. & Sendra A. 1988. Fauna cavernicola de la provincia de Alicante. Volumen 3: Fauna, Flora, Ciencia y Medicina. Instituto de Estudios Juan Gil-Albert. Ed. Diputacion Provincial de Alicante." type="journal volume" year="1988">Zaragoza & Sendra 1988</bibRefCitation>
|
||
); Xixona, Cova-Avenc de Barratxina (
|
||
<bibRefCitation id="EFC392D46675AB43B198614DFAA73F76" author="Blas M." box="[1268,1393,1056,1082]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B5D1612EFF2C3F11" author="Blas M." box="[189,250,1091,1117]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B46A612EFE923F11" author="Blas M." box="[262,324,1091,1117]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B43D6129FDAA3F11" author="Zaragoza & Sendra" box="[337,636,1091,1118]" firstAuthor="Zaragoza" pageId="16" pageNumber="17" refId="ref13879" refString="Zaragoza J. A. & Sendra A. 1988. Fauna cavernicola de la provincia de Alicante. Volumen 3: Fauna, Flora, Ciencia y Medicina. Instituto de Estudios Juan Gil-Albert. Ed. Diputacion Provincial de Alicante." type="journal volume" year="1988">Zaragoza & Sendra 1988</bibRefCitation>
|
||
).
|
||
<collectingRegion id="499621C76675AB43B7FC6129FD3C3F17" box="[656,746,1092,1118]" country="Spain" name="Murcia" pageId="16" pageNumber="17">MURCIA</collectingRegion>
|
||
, Jumilla, Sierra del Carche, MSS (
|
||
<bibRefCitation id="EFC392D46675AB43B1EF612EFB293F11" author="Blas M." box="[1155,1279,1091,1117]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B067612EFEA43FCC" author="Giachino P. M. & Vailati D." firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana" pageId="16" pageNumber="17" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino & Vailati 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B4ED610BFDAA3FCC" author="Fresneda J. & Cardenas A. M. & Castro A. & Lencina J. L. & Lopez-Colon J. I. & Baena. M." box="[385,636,1126,1153]" etAl="et al." firstAuthor="Fresneda" journalOrPublisher="Boletin de la Asociacion espanola de entomologia" pageId="16" pageNumber="17" pagination="187 - 214" part="31" refId="ref11915" refString="Fresneda J., Cardenas A. M., Castro A., Lencina J. L., Lopez-Colon J. I. & Baena. M. 2007. Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera). Boletin de la Asociacion espanola de entomologia 31 (3 - 4): 187 - 214." title="Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera)" type="journal article" year="2007">
|
||
Fresneda
|
||
<emphasis id="B92633376675AB43B499610AFDFD3FCC" box="[501,555,1126,1152]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2007
|
||
</bibRefCitation>
|
||
); Jumilla, Cueva Los Cachorros (
|
||
<bibRefCitation id="EFC392D46675AB43B17C610BFB593FCC" author="Blas M." box="[1040,1167,1126,1152]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
|
||
); Jumilla, Sima los Cachorros, (
|
||
<bibRefCitation id="EFC392D46675AB43B42761E4FDE83FE8" author="Fresneda J. & Cardenas A. M. & Castro A. & Lencina J. L. & Lopez-Colon J. I. & Baena. M." box="[331,574,1161,1188]" etAl="et al." firstAuthor="Fresneda" journalOrPublisher="Boletin de la Asociacion espanola de entomologia" pageId="16" pageNumber="17" pagination="187 - 214" part="31" refId="ref11915" refString="Fresneda J., Cardenas A. M., Castro A., Lencina J. L., Lopez-Colon J. I. & Baena. M. 2007. Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera). Boletin de la Asociacion espanola de entomologia 31 (3 - 4): 187 - 214." title="Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera)" type="journal article" year="2007">
|
||
Fresneda
|
||
<emphasis id="B92633376675AB43B4D161E7FE213FEF" box="[445,503,1161,1187]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
); Jumilla, Cueva Los Tiestos (
|
||
<bibRefCitation id="EFC392D46675AB43B6F761E4FBC03FE8" author="Blas M." box="[923,1046,1161,1188]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
|
||
). —
|
||
<collectingRegion id="499621C76675AB43B12061E4FB603FEF" box="[1100,1206,1161,1187]" country="Spain" name="Comunitat Valenciana" pageId="16" pageNumber="17">Valencia</collectingRegion>
|
||
: Bocairent (
|
||
<bibRefCitation id="EFC392D46675AB43B02D61E4FF2A3F8B" author="Blas M." firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">Blas 1979</bibRefCitation>
|
||
); Bocairent, Cueva del Encomat (
|
||
<bibRefCitation id="EFC392D46675AB43B7E861C0FCFE3F8B" author="Uhagon" box="[644,808,1197,1223]" firstAuthor="Uhagon" pageId="16" pageNumber="17" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B65861C0FC063F8B" author="Jeannel" box="[820,976,1197,1223]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B6B061C0FBCC3F8B" author="Jeannel" box="[988,1050,1197,1223]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B14961C0FB483F8B" author="Blas M." box="[1061,1182,1197,1223]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B1C561C0FB3E3F8B" author="Blas M." box="[1193,1256,1197,1223]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
); Bocairent, Cueva del Seguret (
|
||
<bibRefCitation id="EFC392D46675AB43B4F261BDFD973FA6" author="Uhagon" box="[414,577,1232,1258]" firstAuthor="Uhagon" pageId="16" pageNumber="17" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B72761BDFD303FA6" author="Jeannel" box="[587,742,1232,1258]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B79C61BDFCF83FA6" author="Jeannel" box="[752,814,1232,1258]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B65561BDFC663FA6" author="Blas M." box="[825,944,1232,1258]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B6D661BDFC213FA6" author="Blas M." box="[954,1015,1232,1258]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B16E61BDFB963FA6" author="Blas M." box="[1026,1088,1232,1258]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B12761BDFABF3FA6" author="Giachino P. M. & Vailati D." box="[1099,1385,1232,1258]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana" pageId="16" pageNumber="17" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino & Vailati 1993</bibRefCitation>
|
||
); Bocairent, Cueva de la Zarza (
|
||
<bibRefCitation id="EFC392D46675AB43B74A619EFD1A3E41" author="Uhagon" box="[550,716,1267,1293]" firstAuthor="Uhagon" pageId="16" pageNumber="17" pagination="117 - 126" refId="ref13828" refString="Uhagon S. 1898. Adiciones a mi " Ensayo sobre las especies espanolas del grupo " Cholevae ". Anales de la Sociedad espanola de Historia natural, 7 (2), 27 ". Actas de la Sociedad espanola de Historia natural 1: 117 - 126." type="journal article" year="1898">Uhagón 1898</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B7B5619EFCA03E41" author="Jeannel" box="[729,886,1267,1293]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1 - 98" refId="ref12267" refString="Jeannel R. 1922. Silphidae Catopinae (Coleopteres) (deuxieme serie) avec une etude phylogenique et paleogeographique de la sous-famille. Archives de Zoologie experimentale et generale 61 (1): 1 - 98." type="journal article" year="1922">Jeannel 1922</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B6EF619EFC173E41" author="Jeannel" box="[899,961,1267,1293]" firstAuthor="Jeannel" pageId="16" pageNumber="17" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">1936</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B6A3619EFB9F3E41" author="Blas M." box="[975,1097,1267,1293]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="16" pageNumber="17" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas 1977</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B13A619EFB453E41" author="Blas M." box="[1110,1171,1267,1293]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B1CD619EFB093E41" author="Blas M." box="[1185,1247,1267,1293]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC392D46675AB43B180619EFE863E7C" author="Giachino P. M. & Vailati D." firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana" pageId="16" pageNumber="17" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino & Vailati 1993</bibRefCitation>
|
||
); Bocairent, Cova Sarsa, (
|
||
<bibRefCitation id="EFC392D46675AB43B7ED607BFD2D3E7C" author="Blas M." box="[641,763,1302,1328]" firstAuthor="Blas" journalOrPublisher="PhD Thesis, University of Barcelona, Barcelona" pageId="16" pageNumber="17" refId="ref11559" refString="Blas M. 1979. Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea). PhD Thesis, University of Barcelona, Barcelona." title="Contribucion al conocimiento de los Catopidae (excepto la subfam. Bathysciinae) de la Peninsula Iberica (Coleoptera Staphylinoidea)" type="book" year="1979">Blas 1979</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46675AB43B664607BFC913E7C" author="Blas M." box="[776,839,1302,1328]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="16" pageNumber="17" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">1989</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348BCAE6675AB42B5D16035FEDF38B6" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BEDEF256675AB43B5D16035FEBD3E3E" blockId="16.[189,1398,1368,1511]" box="[189,363,1368,1394]" pageId="16" pageNumber="17">
|
||
<heading id="D0A558496675AB43B5D16035FEBD3E3E" bold="true" box="[189,363,1368,1394]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="6">
|
||
<emphasis id="B92633376675AB43B5D16035FEBD3E3E" bold="true" box="[189,363,1368,1394]" pageId="16" pageNumber="17">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D160EAFBAA3EAB" blockId="16.[189,1398,1368,1511]" pageId="16" pageNumber="17">
|
||
BODY. Length 4.5–5.0 mm (males) and
|
||
<quantity id="4CAA42C06675AB43B7F360EBFCE63EED" box="[671,816,1414,1441]" metricMagnitude="-3" metricUnit="m" metricValue="4.65" metricValueMax="4.9" metricValueMin="4.4" pageId="16" pageNumber="17" unit="mm" value="4.65" valueMax="4.9" valueMin="4.4">4.4–4.9 mm</quantity>
|
||
(females); body width
|
||
<quantity id="4CAA42C06675AB43B12360EBFB093EED" box="[1103,1247,1414,1441]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="16" pageNumber="17" unit="mm" value="1.45" valueMax="1.6" valueMin="1.3">1.3–1.6 mm</quantity>
|
||
(males) and
|
||
<quantity id="4CAA42C06675AB43B5D160C4FE9A3E88" box="[189,332,1449,1476]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="16" pageNumber="17" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
|
||
(females). Body shape long and slender (
|
||
<figureCitation id="1369F3A06675AB43B62E60C4FC5A3E88" box="[834,908,1449,1476]" captionStart="Figs 1 – 5" captionStartId="3.[189,241,1892,1918]" captionTargetBox="[189,1395,1323,1850]" captionTargetId="figure@3.[189,1398,1323,1850]" captionTargetPageId="3" captionText="Figs 1 – 5. Speonemadus of the escalera i- group habitus. 1. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 2. S. angusticollis (Kraatz, 1870) (Rute, Spain). 3. S. bolivari (Jeannel, 1922) (Ardales, Spain). 4. S. breuili (Jeannel, 1922) (Motillas, Spain). 5. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 1 mm." httpUri="https://zenodo.org/record/889348/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
|
||
). Integument reddish brown; pubescence fairly long, golden and lying. Head structure similar to other species of the group.
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D16363FD8F3D23" blockId="16.[189,1399,1548,1647]" pageId="16" pageNumber="17">
|
||
ANTENNAE. Long and slender, when oriented back reaching sixth basal part of elytra; 3rd antennomere slightly longer than 2nd one; 2nd, 4th, 5th and 7th ones are similar in length and longer than the 6th one; 8th is 1.5 × longer than wide (
|
||
<tableCitation id="C6D0DA9E6675AB43B49C6339FD9E3D23" box="[496,584,1620,1647]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="16" pageNumber="17" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D163FBFAA13C55" blockId="16.[189,1399,1686,1818]" pageId="16" pageNumber="17">
|
||
PRONOTUM. Slightly transverse, maximum width / length ratio = 1.19–1.30 (males) and 1.13– 1.20 (females); hexagonal in dorsal view, with maximum width in middle; sides arcuate and almost linear converging towards the base (
|
||
<figureCitation id="1369F3A06675AB43B74863B1FDAA3DBA" box="[548,636,1756,1782]" captionStart="Figs 6 – 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 – 15. Speonemadus of the escalerai - group pronotum. 6 – 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 – 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 – 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 – 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 – 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
|
||
); posterior angles obtuse with rounded apex. Base of pronotum almost as wide as base of elytra, gently curved sideways and fnely beaded. Median granularity sculpture.
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256675AB43B5D1622FFAA13CEE" blockId="16.[189,1399,1857,1954]" pageId="16" pageNumber="17">ELYTRA. Elliptical and very elongated, maximum length / width ratio = 2.06–2.18 (males) and 1.96– 2.06 (female), with rounded apical areas; elytral disc slightly convex and depressed along suture in its half basal part; elytral suture well-marked with thin transverse striae, distant and perpendicular to suture.</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5866466FEFC3A69" blockId="17.[189,1399,267,434]" box="[234,298,267,293]" pageId="17" pageNumber="18">
|
||
<heading id="D0A558496674AB42B5866466FEFC3A69" bold="true" box="[234,298,267,293]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="6">
|
||
<emphasis id="B92633376674AB42B5866466FEFC3A69" bold="true" box="[234,298,267,293]" pageId="17" pageNumber="18">Male</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16443FC213AFE" blockId="17.[189,1399,267,434]" pageId="17" pageNumber="18">
|
||
Aedeagus and structures of inner sac are similar to those described for
|
||
<taxonomicName id="4C5294A66674AB42B1796443FB443A04" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1045,1170,302,328]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bolivari">
|
||
<emphasis id="B92633376674AB42B1796443FB443A04" box="[1045,1170,302,328]" italics="true" pageId="17" pageNumber="18">S. bolivari</emphasis>
|
||
</taxonomicName>
|
||
, although larger in shape, length
|
||
<quantity id="4CAA42C06674AB42B432643CFE3C3A20" box="[350,490,337,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="17" pageNumber="18" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
|
||
(measured to the apex of the parameres); in dorsal view very similar to all the species of the group but in close slimmer apical area (
|
||
<figureCitation id="1369F3A06674AB42B65C6419FC513AC3" box="[816,903,372,399]" captionStart="Figs 21 – 25" captionStartId="7.[189,241,1930,1956]" captionTargetBox="[252,1351,550,1857]" captionTargetId="figure@7.[189,1398,543,1890]" captionTargetPageId="7" captionText="Figs 21 – 25. Speonemadus of the escalerai - group, aedeagus. 21. Paratype of S. algarvensis sp. nov. (Gruta do Vale Telheiro). 22. S. angusticollis (Kraatz, 1870) (Rute). 23. S. bolivari (Jeannel, 1922) (Ardales). 24. S. breuili (Jeannel, 1922) (Motillas). 25. S. escalerai (Uhagón, 1898) (Jumilla). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889354/files/figure.png" pageId="17" pageNumber="18">Fig. 25</figureCitation>
|
||
). Parameres also similar to
|
||
<taxonomicName id="4C5294A66674AB42B1A56418FA953AC2" baseAuthorityName="Jeannel" baseAuthorityYear="1922" box="[1225,1347,372,399]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bolivari">
|
||
<emphasis id="B92633376674AB42B1A56418FA953AC2" box="[1225,1347,372,399]" italics="true" pageId="17" pageNumber="18">S. bolivari</emphasis>
|
||
</taxonomicName>
|
||
, but with more pronounced steeper area of outer margin in third distal part.
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D164B4FB253AB8" blockId="17.[189,1267,473,500]" box="[189,1267,473,500]" pageId="17" pageNumber="18">
|
||
GENITAL SEGMENT STRUCTURE. Similar to that described for
|
||
<taxonomicName id="4C5294A66674AB42B60D64B4FB473ABF" authority="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado, 2017" authorityName="Ana Sofa P.S. Reboleira & Javier Fresneda & José Maria Salgado" authorityYear="2017" box="[865,1169,473,499]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="algarvensis" status="sp. nov.">
|
||
<emphasis id="B92633376674AB42B60D64B4FB473ABF" box="[865,1169,473,499]" italics="true" pageId="17" pageNumber="18">Speonemadus algarvensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2158E4C6674AB42B1F464B7FB253AB8" box="[1176,1267,474,500]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16776FEA4398D" blockId="17.[189,1398,538,705]" pageId="17" pageNumber="18">
|
||
LEGS. Relatively long and slender; protibia widened in apical area and with a long keel running through the two basal thirds of the margin, showing two sharp protrusions, separated by a slight depression, anterior is sharp and much higher and the second is fuzzy (
|
||
<figureCitation id="1369F3A06674AB42B60B670CFC683937" box="[871,958,609,635]" captionStart="Figs 16 – 20" captionStartId="6.[189,241,1892,1918]" captionTargetBox="[190,1399,880,1816]" captionTargetId="figure@6.[190,1399,879,1852]" captionTargetPageId="6" captionText="Figs 16 – 20. Speonemadus of the escalerai - group male protibia. 16. Speonemadus algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal). 17. S. angusticollis (Kraatz, 1870) (Rute, Spain). 18. S. bolivari (Jeannel, 1922) (Ardales, Spain). 19. S. breuili (Jeannel, 1922) (Motillas, Spain). 20. S. escalerai (Uhagón, 1898) (Jumilla, Spain). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/889352/files/figure.png" pageId="17" pageNumber="18">Fig. 20</figureCitation>
|
||
); frst three protarsomeres are dilated, the frst being slightly narrower than the anterior tip of protibia; mesotibiae arched and metatibiae slightly arched.
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5866784FE93384F" blockId="17.[189,1399,745,1018]" box="[234,325,745,771]" pageId="17" pageNumber="18">
|
||
<heading id="D0A558496674AB42B5866784FE93384F" bold="true" box="[234,325,745,771]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="6">
|
||
<emphasis id="B92633376674AB42B5866784FE93384F" bold="true" box="[234,325,745,771]" pageId="17" pageNumber="18">Female</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16661FEDF38B6" blockId="17.[189,1399,745,1018]" pageId="17" pageNumber="18">
|
||
Antennae long, reaching the basal third of elytra, with antennomeres generally slightly longer and transverse (
|
||
<tableCitation id="C6D0DA9E6674AB42B42B6642FE723805" box="[327,420,815,841]" captionStart="Table 1" captionStartId="4.[188,257,267,293]" captionTargetBox="[193,1384,385,1378]" captionTargetPageId="4" captionText="Table 1. Mean measurements (in millimeters) of the antennomers of Speonemadus of the escalerai - group (L) length; (W) width." httpUri="http://table.plazi.org/id/DF2DBFAD6661AB57B5D06466FDD93A04" pageId="17" pageNumber="18" tableUuid="DF2DBFAD6661AB57B5D06466FDD93A04">Table 1</tableCitation>
|
||
). Pronotum not so transverse; in dorsal view subrectangular, with side margins almost straight before maximum width; which is situated towards the third part of basal area (
|
||
<figureCitation id="1369F3A06674AB42B07D663FFABE3821" box="[1297,1384,850,877]" captionStart="Figs 6 – 15" captionStartId="5.[189,241,1857,1883]" captionTargetBox="[189,1398,686,1816]" captionTargetId="figure@5.[189,1398,686,1816]" captionTargetPageId="5" captionText="Figs 6 – 15. Speonemadus of the escalerai - group pronotum. 6 – 7. S. algarvensis sp. nov. (Gruta do Vale Telheiro, Portugal), respectively ♁ and ♀. 8 – 9. S. angusticollis (Kraatz, 1870) (Rute, Spain) respectively ♁ and ♀. 10 – 11. S. bolivari (Jeannel, 1922) (Ardales, Spain) respectively ♁ and ♀. 12 – 13. S. breuili (Jeannel, 1922) (Motillas, Spain) respectively ♁ and ♀. 14 – 15. S. escalerai (Uhagón, 1898) (Jumilla, Spain) respectively ♁ and ♀. Scale bars = 1 mm." httpUri="https://zenodo.org/record/889350/files/figure.png" pageId="17" pageNumber="18">Fig. 15</figureCitation>
|
||
). Elytra elliptical and elongate, with notched and serrated apical areas. Sexual dimorphism very clear in pronotum, protibiae and apical part of elytra. Females are generally less robust and with less transverse pronotum than males. The uroventrites 7th and 8th, and spermatheca follows the overall model for the group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348BCAE6674AB42B5D1614CFEFC3F9F" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="8BEDEF256674AB42B5D1614CFF2E3F77" blockId="17.[189,248,1057,1083]" box="[189,248,1057,1083]" pageId="17" pageNumber="18">
|
||
<heading id="D0A558496674AB42B5D1614CFF2E3F77" bold="true" box="[189,248,1057,1083]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
|
||
<emphasis id="B92633376674AB42B5D1614CFF2E3F77" bold="true" box="[189,248,1057,1083]" pageId="17" pageNumber="18">Note</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16122FEFC3F9F" blockId="17.[189,1399,1103,1235]" pageId="17" pageNumber="18">
|
||
The great variability of this species led to the description of the subspecies
|
||
<treatmentCitation id="0AF3C9346674AB42B121613DFAA73F25" author="Jeannel" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1101,1393,1103,1129]" class="Insecta" family="Leiodidae" firstAuthor="Jeannel" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" pagination="1" phylum="Arthropoda" rank="subSpecies" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." species="escalerai" subSpecies="zariquieyi" treatmentTaxon="zariquieyi" type="journal article" year="1936">
|
||
<taxonomicName id="4C5294A66674AB42B121613DFAA73F25" ID-CoL="4YHZ8" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1101,1393,1103,1129]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="subSpecies" species="escalerai" subSpecies="zariquieyi">
|
||
<emphasis id="B92633376674AB42B121613DFB143F25" box="[1101,1218,1104,1129]" italics="true" pageId="17" pageNumber="18">zariquieyi</emphasis>
|
||
<bibRefCitation id="EFC392D46674AB42B1A06122FAA73F25" author="Jeannel" box="[1228,1393,1103,1129]" firstAuthor="Jeannel" pageId="17" pageNumber="18" pagination="1" refId="ref12306" refString="Jeannel R. 1936. Monographie des Catopides. Memoires du Museum national d'Histoire naturelle 1 (1). Museum national d'Histoire naturelle, Paris." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, later synonymized by
|
||
<treatmentCitation id="0AF3C9346674AB42B4AE611FFD993FC0" author="Blas M." box="[450,591,1138,1164]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="17" pageNumber="18" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" treatmentTaxon="zariquieyi" type="book chapter" year="1977">
|
||
<bibRefCitation id="EFC392D46674AB42B4AE611FFD993FC0" author="Blas M." box="[450,591,1138,1164]" firstAuthor="Blas" journalOrPublisher="Comunicacions 6 e Simposium d'Espeleologia" pageId="17" pageNumber="18" pagination="125 - 130" refId="ref11528" refString="Blas M. 1977. Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae). Comunicacions 6 e Simposium d'Espeleologia: 125 - 130." title="Contribucion al conocimiento de losAnemadinae de la Peninsula Iberica. (Col. Catopidae)" type="book chapter" year="1977">Blas (1977)</bibRefCitation>
|
||
</treatmentCitation>
|
||
, who observed that the differences correspond to intermediate forms and, moreover, the proximity of the different caves does not represent any real evidence of geographic isolation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348BCAE6674AB42B5D16197FEA43E83" pageId="17" pageNumber="18" type="biology_ecology">
|
||
<paragraph id="8BEDEF256674AB42B5D16197FE6A3E58" blockId="17.[189,444,1274,1300]" box="[189,444,1274,1300]" pageId="17" pageNumber="18">
|
||
<heading id="D0A558496674AB42B5D16197FE6A3E58" bold="true" box="[189,444,1274,1300]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
|
||
<emphasis id="B92633376674AB42B5D16197FE6A3E58" bold="true" box="[189,444,1274,1300]" pageId="17" pageNumber="18">Biology and Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16044FEA43E83" blockId="17.[189,1399,1321,1488]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="4C5294A66674AB42B5D16044FE073E0F" baseAuthorityName="Uhagon" baseAuthorityYear="1898" box="[189,465,1321,1347]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="escalerai">
|
||
<emphasis id="B92633376674AB42B5D16044FE073E0F" box="[189,465,1321,1347]" italics="true" pageId="17" pageNumber="18">Speonemadus escalerai</emphasis>
|
||
</taxonomicName>
|
||
is a subterranean species and, therefore, common in caves. It is mostly found in areas with small accumulations of bat guano, but also under stones or stalagmitic foors and in deep parts of caves (
|
||
<bibRefCitation id="EFC392D46674AB42B4406002FE713EC5" author="Blas M." box="[300,423,1391,1417]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="17" pageNumber="18" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas 1989</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFC392D46674AB42B4D66002FD903EC5" author="Blas M." box="[442,582,1391,1417]" firstAuthor="Blas" journalOrPublisher="Memoires de Biospeologie" pageId="17" pageNumber="18" pagination="135 - 140" part="16" refId="ref11653" refString="Blas M. 1989. El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae). Memoires de Biospeologie 16: 135 - 140." title="El genero Speonemadus Jeannel y su distribucion geografca. (Col. Catopidae)" type="journal article" year="1989">Blas (1989)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC392D46674AB42B73C6002FC573EC5" author="Giachino P. M. & Vailati D." box="[592,897,1391,1417]" firstAuthor="Giachino" journalOrPublisher="Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana" pageId="17" pageNumber="18" pagination="1 - 314" part="18" refId="ref12035" refString="Giachino P. M. & Vailati D. 1993. Revisione degli Anemadinae. Museo Civico di Scienze Naturali di Brescia. Monografe di " Natura Bresciana " 18: 1 - 314." title="Revisione degli Anemadinae" type="journal article" year="1993">Giachino & Vailati (1993)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFC392D46674AB42B6D46002FB603EC5" author="Fresneda J. & Cardenas A. M. & Castro A. & Lencina J. L. & Lopez-Colon J. I. & Baena. M." box="[952,1206,1391,1417]" etAl="et al." firstAuthor="Fresneda" journalOrPublisher="Boletin de la Asociacion espanola de entomologia" pageId="17" pageNumber="18" pagination="187 - 214" part="31" refId="ref11915" refString="Fresneda J., Cardenas A. M., Castro A., Lencina J. L., Lopez-Colon J. I. & Baena. M. 2007. Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera). Boletin de la Asociacion espanola de entomologia 31 (3 - 4): 187 - 214." title="Nuevos datos de los Cholevidae en la Peninsula Iberica (Coleoptera)" type="journal article" year="2007">
|
||
Fresneda
|
||
<emphasis id="B92633376674AB42B144601DFBB43EC5" box="[1064,1122,1391,1417]" italics="true" pageId="17" pageNumber="18">et al.</emphasis>
|
||
(2007)
|
||
</bibRefCitation>
|
||
also pointed out its presence in the Superfcial Subterranean Habitat (MSS) (
|
||
<emphasis id="B92633376674AB42B61060FEFC6B3EE0" box="[892,957,1427,1452]" italics="true" pageId="17" pageNumber="18">sensu</emphasis>
|
||
<bibRefCitation id="EFC392D46674AB42B6A860FFFB333EE0" author="Giachino & Vailati" box="[964,1253,1426,1452]" firstAuthor="Giachino" pageId="17" pageNumber="18" pagination="3" refId="ref12117" refString="Giachino P. M. & Vailati D. 2010. The subterranean environment. Hypogean life, concepts and collecting techniques. WBA Handbooks 3, World Biodiversity Association, Verona." type="book chapter" year="2010">Giachino & Vailati 2010</bibRefCitation>
|
||
) in the karst area of
|
||
<collectingRegion id="499621C76674AB42B47960D8FEB83E83" box="[277,366,1461,1487]" country="Spain" name="Murcia" pageId="17" pageNumber="18">Murcia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C348BCAE6674AB42B5D1609AFD293D2F" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="8BEDEF256674AB42B5D1609AFE803D5D" blockId="17.[189,342,1527,1553]" box="[189,342,1527,1553]" pageId="17" pageNumber="18">
|
||
<heading id="D0A558496674AB42B5D1609AFE803D5D" bold="true" box="[189,342,1527,1553]" fontSize="11" level="2" pageId="17" pageNumber="18" reason="2">
|
||
<emphasis id="B92633376674AB42B5D1609AFE803D5D" bold="true" box="[189,342,1527,1553]" pageId="17" pageNumber="18">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEDEF256674AB42B5D16348FD293D2F" blockId="17.[189,1398,1573,1635]" pageId="17" pageNumber="18">
|
||
<materialsCitation id="3B3AE5786674AB42B5D16348FD293D2F" ID-GBIF-Occurrence="1609370441" country="Spain" location="Murcia" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Murcia">
|
||
This species is distributed in the southeast Iberian Peninsula and colonizes caves of the provinces of Alicante,
|
||
<collectingRegion id="499621C76674AB42B4436324FE523D2F" box="[303,388,1609,1635]" country="Spain" name="Murcia" pageId="17" pageNumber="18">Murcia</collectingRegion>
|
||
and Valencia in
|
||
<collectingCountry id="F345AFB56674AB42B72B6324FD5C3D2F" box="[583,650,1609,1635]" name="Spain" pageId="17" pageNumber="18">Spain</collectingCountry>
|
||
(
|
||
<figureCitation id="1369F3A06674AB42B7F66324FD273D2F" box="[666,753,1609,1635]" captionStart="Fig. 32" captionStartId="18.[189,232,1955,1981]" captionTargetBox="[189,1398,986,1883]" captionTargetId="figure@18.[189,1398,986,1914]" captionTargetPageId="18" captionText="Fig. 32. Distribution of Speonemadus of the escalerai - group: S. algarvensis sp. nov., S. angusticollis (Kraatz, 1870), S. bolivari (Jeannel, 1922), S. breuili (Jeannel, 1922) and S. escalerai (Uhagón, 1898)." httpUri="https://zenodo.org/record/889358/files/figure.png" pageId="17" pageNumber="18">Fig. 32</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |