443 lines
59 KiB
XML
443 lines
59 KiB
XML
<document id="A8A547C35DBA19A7832C6AC233815571" ID-CLB-Dataset="24438" ID-DOI="10.11646/zootaxa.4718.1.7" ID-GBIF-Dataset="66ccf05a-746e-41a8-b2a6-85901c55d154" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602258" ID-ZooBank="A4308225-3D68-4F4F-BF39-35EC103B3029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578564328630" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03E5EE7F4B062317F3EDFB430BF2FDC4" docLanguage="en" docName="zootaxa.4718.1.7.pdf" docOrigin="Zootaxa 4718 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clidicus minilankanus Jałoszyński 2020, sp. n." docType="treatment" docVersion="6" lastPageNumber="93" masterDocId="FFDC96074B072311F37AFFB70819FFA7" masterDocTitle="Clidicus minilankanus sp. n., with notes on remaining Sri Lankan Clidicus species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="94" masterPageNumber="87" pageNumber="88" updateTime="1698776474274" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="40E4A33E31365943DF91E915686C5344">Clidicus minilankanus sp. n., with notes on remaining Sri Lankan Clidicus species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:name id="ED449362E3D3E3519D168FC25D9DE8C0" type="personal">
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<mods:namePart id="38A65D779C71BCA5F2D4C4FA159B45A0">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="BD2C11432368A8E35BE8712C57A882DD">Zootaxa</mods:title>
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<mods:part id="FDAA38E283F6C208624790BA8102E438">
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<mods:date id="770798AA8FBAAFE0A3DC1B984C0E210B">2020</mods:date>
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<mods:number id="4192B7E05223E2C47709EBD91D29C67F">2020-01-03</mods:number>
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<mods:number id="32F02B8295DBEB7A7DE29FEBB219D977">4718</mods:number>
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<mods:identifier id="F2ADE97010CAB2C8D93084AFD52194CD" type="DOI">10.11646/zootaxa.4718.1.7</mods:identifier>
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<treatment id="03E5EE7F4B062317F3EDFB430BF2FDC4" ID-GBIF-Taxon="161670736" LSID="urn:lsid:plazi:treatment:03E5EE7F4B062317F3EDFB430BF2FDC4" httpUri="http://treatment.plazi.org/id/03E5EE7F4B062317F3EDFB430BF2FDC4" lastPageId="6" lastPageNumber="93" pageId="1" pageNumber="88">
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<subSubSection id="C3560CE24B062310F3EDFB4309EAFAA8" box="[151,499,1268,1295]" pageId="1" pageNumber="88" type="nomenclature">
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<paragraph id="8BF35F694B062310F3EDFB4309EAFAA8" blockId="1.[151,499,1268,1331]" box="[151,499,1268,1295]" pageId="1" pageNumber="88">
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<heading id="D0BBE8054B062310F3EDFB4309EAFAA8" bold="true" box="[151,499,1268,1295]" fontSize="11" level="1" pageId="1" pageNumber="88" reason="1">
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<emphasis id="B938837B4B062310F3EDFB4309EAFAA8" bold="true" box="[151,499,1268,1295]" pageId="1" pageNumber="88">
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<taxonomicName id="4C4C24EA4B062310F3EDFB4309B3FAA8" ID-CoL="8XXT9" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,426,1268,1295]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="minilankanus" status="sp. nov.">
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<emphasis id="B938837B4B062310F3EDFB4309B3FAA8" bold="true" box="[151,426,1268,1295]" italics="true" pageId="1" pageNumber="88">Clidicus minilankanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B3E004B062310F2CBFB4209EAFAA8" box="[433,499,1269,1295]" pageId="1" pageNumber="88" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560CE24B062310F3EDFAAF09BCFA94" box="[151,421,1304,1331]" pageId="1" pageNumber="88" type="description">
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<paragraph id="8BF35F694B062310F3EDFAAF09BCFA94" blockId="1.[151,499,1268,1331]" box="[151,421,1304,1331]" pageId="1" pageNumber="88">
|
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(
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<figureCitation id="137743EC4B062310F3E5FAAF091DFA94" box="[159,260,1304,1331]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="1" pageNumber="88">Figs 1–2</figureCitation>
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,
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<figureCitation id="137743EC4B062310F26AFAAE0939FA94" box="[272,288,1305,1331]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1900,1925]" captionTargetBox="[158,1430,409,1880]" captionTargetId="figure@3.[158,1430,409,1880]" captionTargetPageId="3" captionText="FIGURES 7–12. Clidicus of Sri Lanka; head and pronotum in dorsal (7–10, 12) and posterodorsal (11) views. Clidicus mi- nilankanus sp. n., holotype male (7); C. loebli Besuchet, holotype female (8); C. monstrosus (Walker), lectotype female (9); C. mussardi Besuchet, holotype male (10–11; arrow showing transverse groove demarcating pronotal collar); C. quadricollis Besuchet, holotype female (12)." figureDoi="http://doi.org/10.5281/zenodo.3602264" httpUri="https://zenodo.org/record/3602264/files/figure.png" pageId="1" pageNumber="88">7</figureCitation>
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,
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<figureCitation id="137743EC4B062310F256FAAE0953FA94" box="[300,330,1305,1331]" captionStart="FIGURES 13–17" captionStartId="4.[151,264,1936,1961]" captionTargetBox="[151,1436,181,1871]" captionTargetId="figure@4.[151,1436,181,1871]" captionTargetPageId="4" captionText="FIGURES 13–17. Clidicus of Sri Lanka; head and thorax in ventral view. Clidicus minilankanus sp. n., paratype female (13); C. loebli Besuchet, holotype female (14); C. monstrosus (Walker), lectotype female (15); C. mussardi Besuchet, holotype male (16); C. quadricollis Besuchet, holotype female (17)." figureDoi="http://doi.org/10.5281/zenodo.3602266" httpUri="https://zenodo.org/record/3602266/files/figure.png" pageId="1" pageNumber="88">13</figureCitation>
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,
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<figureCitation id="137743EC4B062310F22CFAAE0984FA94" box="[342,413,1305,1331]" captionStart="FIGURES 18–21" captionStartId="5.[151,264,1737,1762]" captionTargetBox="[164,1419,922,1711]" captionTargetId="figure@5.[151,1436,913,1718]" captionTargetPageId="5" captionText="FIGURES 18–21. Clidicus minilankanus sp. n. Aedeagus in ventral (18) and lateral (19) views; sperm pump (20); ovipositor with spermatheca (21)." figureDoi="http://doi.org/10.5281/zenodo.3602268" httpUri="https://zenodo.org/record/3602268/files/figure.png" pageId="1" pageNumber="88">18–21</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560CE24B062310F3EDFAD70B4DFA40" pageId="1" pageNumber="88" type="materials_examined">
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<paragraph id="8BF35F694B062310F3EDFAD70B4DFA40" blockId="1.[151,1436,1376,2015]" pageId="1" pageNumber="88">
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<emphasis id="B938837B4B062310F3EDFAD7097EFADC" bold="true" box="[151,359,1376,1403]" pageId="1" pageNumber="88">Material studied.</emphasis>
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<materialsCitation id="3B2455344B062310F20AFAD70CF7FA64" ID-GBIF-Occurrence="2545448161" collectingDate="2016-09-17" collectionCode="SRI, LANKA, EUMJ" collectorName="Sk. Yamane" country="Sri Lanka" location="Western Prov." pageId="1" pageNumber="88" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B938837B4B062310F20AFAD70A01FADC" bold="true" box="[368,536,1376,1403]" pageId="1" pageNumber="88">
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<typeStatus id="54F7E1CB4B062310F20AFAD70A0AFADC" box="[368,531,1376,1403]" pageId="1" pageNumber="88" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D4A94E04B062310F15BFAD60A2FFADC" box="[545,566,1377,1403]" pageId="1" pageNumber="88" type="male">♂</specimenCount>
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:
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<emphasis id="B938837B4B062310F133FAD70AFFFADC" bold="true" box="[585,742,1376,1403]" pageId="1" pageNumber="88">
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<collectingCountry id="F35B1FF94B062310F133FAD70AFFFADC" box="[585,742,1376,1403]" name="Sri Lanka" pageId="1" pageNumber="88">SRI LANKA</collectingCountry>
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</emphasis>
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, with three labels: “[
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<collectionCode id="ED5DC7AC4B062310F099FAD60C16FADC" box="[995,1039,1377,1403]" country="Tanzania" httpUri="http://grbio.org/cool/z5p4-dpar" name="Serengetti Research Institute" pageId="1" pageNumber="88">SRI</collectionCode>
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<collectionCode id="ED5DC7AC4B062310F762FAD60C61FADC" box="[1048,1144,1377,1403]" pageId="1" pageNumber="88">LANKA</collectionCode>
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] /
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<location id="8E9309B24B062310F7DFFAD60D5FFADC" LSID="urn:lsid:plazi:treatment:03E5EE7F4B062317F3EDFB430BF2FDC4:8E9309B24B062310F7DFFAD60D5FFADC" box="[1189,1350,1377,1403]" country="Sri Lanka" name="Western Prov." pageId="1" pageNumber="88">Western Prov.</location>
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, / Keg- alle distr. / Salgala For. Resv.” [pink, printed], “
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<date id="FFF279A94B062310F1CBFA320B21FA39" box="[689,824,1412,1439]" pageId="1" pageNumber="88" value="2016-09-17">
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<collectingDate id="EFB680414B062310F1CBFA320B21FA39" box="[689,824,1412,1439]" pageId="1" pageNumber="88" value="2016-09-17">17. ix. 2016</collectingDate>
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</date>
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/
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<collectorName id="26B93ABF4B062310F035FA330BCDFA38" box="[847,980,1412,1439]" pageId="1" pageNumber="88">Sk. Yamane</collectorName>
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leg. / ex leaf litter / general coll.” [pink, printed], “
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<taxonomicName id="4C4C24EA4B062310F270FA1F097EFA65" box="[266,359,1448,1474]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="genus">
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<emphasis id="B938837B4B062310F270FA1F097EFA65" box="[266,359,1448,1474]" italics="true" pageId="1" pageNumber="88">Clidicus</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="4C4C24EA4B062310F207FA1F0A0DFA65" authorityName="Jałoszyński" authorityYear="2020" box="[381,532,1448,1474]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="88" phylum="Arthropoda" rank="species" species="minilankanus">
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<emphasis id="B938837B4B062310F207FA1F0A0DFA65" box="[381,532,1448,1474]" italics="true" pageId="1" pageNumber="88">minilankanus</emphasis>
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</taxonomicName>
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m. /
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<typeStatus id="54F7E1CB4B062310F136FA1E0AECFA64" box="[588,757,1449,1475]" pageId="1" pageNumber="88" type="holotype">HOLOTYPUS</typeStatus>
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, P.
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<bibRefCitation id="EFDD22984B062310F061FA1F0BFDFA64" author="Jaloszynski, P. & Cheng, Z. - F. & Brajczewska, N. & Yin, Z. - W." box="[795,996,1448,1475]" pageId="1" pageNumber="88" pagination="571 - 584" refId="ref3660" refString="Jaloszynski, P., Cheng, Z. - F., Brajczewska, N. & Yin, Z. - W. (2019) Hidden diversity of giant scydmaenines: five new species and new records of Clidicus in the Philippines (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4658 (3), 571 - 584. https: // doi. org / 10.11646 / zootaxa. 4658.3.8" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
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” [red, printed] (
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<collectionCode id="ED5DC7AC4B062310F7EDFA1E0CFFFA64" box="[1175,1254,1449,1475]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="1" pageNumber="88">EUMJ</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B2455344B062310F782FA1F0B49FA40" ID-GBIF-Occurrence="2545448160" collectingDate="2016-09-17" collectionCode="EUMJ" collectorName="Sk. Yamane" country="Sri Lanka" location="Western Prov." pageId="1" pageNumber="88" specimenCount="8" specimenCount-female="5" specimenCount-male="3" typeStatus="paratype">
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<emphasis id="B938837B4B062310F782FA1F0D85FA64" bold="true" box="[1272,1436,1448,1475]" pageId="1" pageNumber="88">
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||
<typeStatus id="54F7E1CB4B062310F782FA1F0D85FA64" box="[1272,1436,1448,1475]" pageId="1" pageNumber="88" type="paratype">PARATYPES</typeStatus>
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||
</emphasis>
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(8 exx.):
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<specimenCount id="9D4A94E04B062310F387FA7A0927FA40" box="[253,318,1485,1511]" pageId="1" pageNumber="88" type="male">3 ♂♂</specimenCount>
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,
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<specimenCount id="9D4A94E04B062310F230FA7B0993FA40" box="[330,394,1484,1511]" pageId="1" pageNumber="88" type="female">5 ♀♀</specimenCount>
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, same data as for holotype (
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<collectionCode id="ED5DC7AC4B062310F1BEFA7A0B08FA40" box="[708,785,1485,1511]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="1" pageNumber="88">EUMJ</collectionCode>
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, cPJ)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3560CE24B062310F3BDFA470D7AF919" pageId="1" pageNumber="88" type="diagnosis">
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||
<paragraph id="8BF35F694B062310F3BDFA470D7AF919" blockId="1.[151,1436,1376,2015]" pageId="1" pageNumber="88">
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<emphasis id="B938837B4B062310F3BDFA47095BF9AC" bold="true" box="[199,322,1520,1547]" pageId="1" pageNumber="88">Diagnosis.</emphasis>
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Body length <
|
||
<quantity id="4CB4F28C4B062310F28BFA460A2CF9AC" box="[497,565,1521,1547]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="88" unit="mm" value="4.0">4 mm</quantity>
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; pronotum in dorsal view subquadrangular and distinctly flattened, with shallow and diffuse subtriangular posteromedian impression, with posterior ‘collar’ reduced to barely discernible short median stripe dorsally demarcated from disc by shallow biarcuate transverse groove; elytra shield-shaped, broadest just behind distinct, obtuse-angled humeri; male protrochanter modified, with anteroventral subtriangular projection; aedeagus symmetrical, broadest near base, with apices of parameres projecting distad beyond apex of median lobe, strongly curved in lateral view; sperm pump long and slender, with shallow distal and proximal funnels.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3560CE24B062317F3BDF97F0965FF70" lastPageId="6" lastPageNumber="93" pageId="1" pageNumber="88" type="description">
|
||
<paragraph id="8BF35F694B062310F3BDF97F0BFEF8A1" blockId="1.[151,1436,1376,2015]" pageId="1" pageNumber="88">
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||
<emphasis id="B938837B4B062310F3BDF97F0940F944" bold="true" box="[199,345,1736,1763]" pageId="1" pageNumber="88">Description.</emphasis>
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Body of male (
|
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<figureCitation id="137743EC4B062310F174F97F0A49F944" box="[526,592,1736,1763]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="1" pageNumber="88">Fig. 1</figureCitation>
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) relatively stout, distinctly flattened, BL
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<quantity id="4CB4F28C4B062310F750F97F0CD6F944" box="[1066,1231,1736,1763]" metricMagnitude="-3" metricUnit="m" metricValue="3.79" metricValueMax="3.83" metricValueMin="3.75" pageId="1" pageNumber="88" unit="mm" value="3.79" valueMax="3.83" valueMin="3.75">3.75–3.83 mm</quantity>
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||
(mean
|
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<quantity id="4CB4F28C4B062310F65CF97E0D96F944" box="[1318,1423,1737,1763]" metricMagnitude="-3" metricUnit="m" metricValue="3.79" pageId="1" pageNumber="88" unit="mm" value="3.79">3.79 mm</quantity>
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); cuticle glossy, pigmentation uniformly light to dark brown, setae yellowish.
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</paragraph>
|
||
<paragraph id="8BF35F694B062312F3BDF8A70ACAFE2C" blockId="1.[151,1436,1376,2015]" lastBlockId="3.[151,1437,153,395]" lastPageId="3" lastPageNumber="90" pageId="1" pageNumber="88">
|
||
Head (
|
||
<figureCitation id="137743EC4B062310F269F8A7094EF88C" box="[275,343,1808,1835]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1900,1925]" captionTargetBox="[158,1430,409,1880]" captionTargetId="figure@3.[158,1430,409,1880]" captionTargetPageId="3" captionText="FIGURES 7–12. Clidicus of Sri Lanka; head and pronotum in dorsal (7–10, 12) and posterodorsal (11) views. Clidicus mi- nilankanus sp. n., holotype male (7); C. loebli Besuchet, holotype female (8); C. monstrosus (Walker), lectotype female (9); C. mussardi Besuchet, holotype male (10–11; arrow showing transverse groove demarcating pronotal collar); C. quadricollis Besuchet, holotype female (12)." figureDoi="http://doi.org/10.5281/zenodo.3602264" httpUri="https://zenodo.org/record/3602264/files/figure.png" pageId="1" pageNumber="88">Fig. 7</figureCitation>
|
||
) subtrapezoidal, broadest just in front of posterior margins of tempora, HL
|
||
<quantity id="4CB4F28C4B062310F7C9F8A70D03F88C" box="[1203,1306,1808,1835]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="88" unit="mm" value="0.65">0.65 mm</quantity>
|
||
, HW 1.00–
|
||
<quantity id="4CB4F28C4B062310F3EDF88208E5F8E8" box="[151,252,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="1" pageNumber="88" unit="mm" value="1.03">1.03 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B062310F229F88209A5F8E8" box="[339,444,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="88" unit="mm" value="1.01">1.01 mm</quantity>
|
||
); tempora in dorsal view 3 times as long as eyes, nearly straight and strongly divergent posterad, their posterolateral portions subtriangular and strongly projecting; vertex with short, shallow and diffuse subtriangular posteromedian impression bearing pair of distinct posterior pits marking sites where dorsal tentorial arms are fused with dorsal wall of head. Sides of vertex and frons evenly convex, frons convex also between weakly elevated supraantennal tubercles; clypeus transverse and weakly convex, demarcated from frons by deep and distinct frontoclypeal groove; compound eyes small, nearly round, strongly convex but in dorsal view weakly projecting from head silhouette, finely faceted. Punctures on vertex and frons distinct, small and shallow but sharply marked, unevenly distributed, separated by spaces 0.5–2 × as wide as diameters of punctures. Setae long, sparse, erect and mostly directed dorsoanterad; a pair of longer but similarly thin setae can be seen on posterior margin of vertex, close to posteromedian impression. Antennae (
|
||
<figureCitation id="137743EC4B042312F07AFE9E0B5BFEE4" box="[768,834,297,323]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="3" pageNumber="90">Fig. 1</figureCitation>
|
||
) short and compact, AnL
|
||
<quantity id="4CB4F28C4B042312F70AFE9F0D0DFEE4" box="[1136,1300,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="3" pageNumber="90" unit="mm" value="1.9" valueMax="1.95" valueMin="1.85">1.85–1.95 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B042312F611FE9E08DEFEC0" metricMagnitude="-3" metricUnit="m" metricValue="1.89" pageId="3" pageNumber="90" unit="mm" value="1.89">1.89 mm</quantity>
|
||
), scape slightly shorter than head, antennomere II weakly elongate, III–VII each about as long as broad, VIII–X each slightly transverse, XI about as long as broad.
|
||
</paragraph>
|
||
<caption id="DF330FE14B052313F3EDF8E908A2F844" ID-DOI="http://doi.org/10.5281/zenodo.3602262" ID-Zenodo-Dep="3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="2" pageNumber="89" startId="2.[151,264,1886,1911]" targetBox="[157,1427,188,1830]" targetPageId="2">
|
||
<paragraph id="8BF35F694B052313F3EDF8E908A2F844" blockId="2.[151,1437,1886,2019]" pageId="2" pageNumber="89">
|
||
<emphasis id="B938837B4B052313F3EDF8E90922F8D0" bold="true" box="[151,315,1886,1911]" pageId="2" pageNumber="89">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C4C24EA4B052313F238F8E8098FF8D0" box="[322,406,1887,1911]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938837B4B052313F238F8E8098FF8D0" box="[322,406,1887,1911]" italics="true" pageId="2" pageNumber="89">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2).
|
||
<taxonomicName id="4C4C24EA4B052313F7E9F8E80D61F8D0" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[1171,1400,1887,1911]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="minilankanus" status="sp. nov.">
|
||
<emphasis id="B938837B4B052313F7E9F8E80D61F8D0" box="[1171,1400,1887,1911]" italics="true" pageId="2" pageNumber="89">Clidicus minilankanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938837B4B052313F604F8E808B5F83C" bold="true" pageId="2" pageNumber="89">
|
||
<taxonomicNameLabel id="A20B3E004B052313F604F8E808B5F83C" pageId="2" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (1) and paratype male (2; arrow showing modified protrochanter);
|
||
<taxonomicName id="4C4C24EA4B052313F096F8330CB1F83C" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[1004,1192,1922,1947]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B938837B4B052313F096F8330C5AF83C" box="[1004,1091,1923,1947]" italics="true" pageId="2" pageNumber="89">C. loebli</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (3);
|
||
<taxonomicName id="4C4C24EA4B052313F6FFF8330975F818" authority="(Walker)" authorityName="" baseAuthorityName="Walker" baseAuthorityYear="1858" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="monstrosus">
|
||
<emphasis id="B938837B4B052313F6FFF8330910F818" italics="true" pageId="2" pageNumber="89">C. monstrosus</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
, lectotype female (4);
|
||
<taxonomicName id="4C4C24EA4B052313F136F81F0B33F818" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[588,810,1958,1983]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="mussardi">
|
||
<emphasis id="B938837B4B052313F136F81F0ADCF818" box="[588,709,1959,1983]" italics="true" pageId="2" pageNumber="89">C. mussardi</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype male (5);
|
||
<taxonomicName id="4C4C24EA4B052313F089F81F0CF6F818" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[1011,1263,1958,1983]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B052313F089F81F0C93F818" box="[1011,1162,1959,1983]" italics="true" pageId="2" pageNumber="89">C. quadricollis</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (6).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF330FE14B042312F3EDF8DB09C5F857" ID-DOI="http://doi.org/10.5281/zenodo.3602264" ID-Zenodo-Dep="3602264" httpUri="https://zenodo.org/record/3602264/files/figure.png" pageId="3" pageNumber="90" startId="3.[151,264,1900,1925]" targetBox="[158,1430,409,1880]" targetPageId="3">
|
||
<paragraph id="8BF35F694B042312F3EDF8DB09C5F857" blockId="3.[151,1437,1900,2033]" pageId="3" pageNumber="90">
|
||
<emphasis id="B938837B4B042312F3EDF8DB0950F822" bold="true" box="[151,329,1900,1925]" pageId="3" pageNumber="90">FIGURES 7–12.</emphasis>
|
||
<taxonomicName id="4C4C24EA4B042312F22BF8DB09BCF823" box="[337,421,1900,1924]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938837B4B042312F22BF8DB09BCF823" box="[337,421,1900,1924]" italics="true" pageId="3" pageNumber="90">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
of Sri Lanka; head and pronotum in dorsal (7–10, 12) and posterodorsal (11) views.
|
||
<emphasis id="B938837B4B042312F664F8DB091FF80F" italics="true" pageId="3" pageNumber="90">
|
||
<taxonomicName id="4C4C24EA4B042312F664F8DB0D6BF823" box="[1310,1394,1900,1924]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">Clidicus</taxonomicName>
|
||
mi- nilankanus
|
||
</emphasis>
|
||
<emphasis id="B938837B4B042312F274F8270950F80F" bold="true" box="[270,329,1936,1960]" pageId="3" pageNumber="90">sp. n.</emphasis>
|
||
, holotype male (7);
|
||
<taxonomicName id="4C4C24EA4B042312F164F8260AC7F80E" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[542,734,1936,1961]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B938837B4B042312F164F8260A6EF80F" box="[542,631,1936,1960]" italics="true" pageId="3" pageNumber="90">C. loebli</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (8);
|
||
<taxonomicName id="4C4C24EA4B042312F0BEF8260CA3F80E" authority="(Walker)" authorityName="" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[964,1210,1936,1961]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="monstrosus">
|
||
<emphasis id="B938837B4B042312F0BEF8260C4FF80F" box="[964,1110,1937,1960]" italics="true" pageId="3" pageNumber="90">C. monstrosus</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
, lectotype female (9);
|
||
<taxonomicName id="4C4C24EA4B042312F3EDF8020960F86A" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[151,377,1972,1997]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="mussardi">
|
||
<emphasis id="B938837B4B042312F3EDF802090BF86B" box="[151,274,1972,1996]" italics="true" pageId="3" pageNumber="90">C. mussardi</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype male (10–11; arrow showing transverse groove demarcating pronotal collar);
|
||
<taxonomicName id="4C4C24EA4B042312F679F80208EFF856" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="90" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B042312F679F8020D85F86B" box="[1283,1436,1972,1996]" italics="true" pageId="3" pageNumber="90">C. quadricollis</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (12).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF330FE14B032315F3EDF8270AABF857" ID-DOI="http://doi.org/10.5281/zenodo.3602266" ID-Zenodo-Dep="3602266" httpUri="https://zenodo.org/record/3602266/files/figure.png" pageId="4" pageNumber="91" startId="4.[151,264,1936,1961]" targetBox="[151,1436,181,1871]" targetPageId="4">
|
||
<paragraph id="8BF35F694B032315F3EDF8270AABF857" blockId="4.[151,1437,1936,2033]" pageId="4" pageNumber="91">
|
||
<emphasis id="B938837B4B032315F3EDF827094DF80E" bold="true" box="[151,340,1936,1961]" pageId="4" pageNumber="91">FIGURES 13–17.</emphasis>
|
||
<taxonomicName id="4C4C24EA4B032315F221F82709B6F80F" box="[347,431,1936,1960]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938837B4B032315F221F82709B6F80F" box="[347,431,1936,1960]" italics="true" pageId="4" pageNumber="91">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
of Sri Lanka; head and thorax in ventral view.
|
||
<taxonomicName id="4C4C24EA4B032315F0EBF8270C6CF80F" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[913,1141,1936,1960]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="minilankanus" status="sp. nov.">
|
||
<emphasis id="B938837B4B032315F0EBF8270C6CF80F" box="[913,1141,1936,1960]" italics="true" pageId="4" pageNumber="91">Clidicus minilankanus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B938837B4B032315F701F8270CADF80F" bold="true" box="[1147,1204,1936,1960]" pageId="4" pageNumber="91">
|
||
<taxonomicNameLabel id="A20B3E004B032315F701F8270CADF80F" box="[1147,1204,1936,1960]" pageId="4" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype female (13);
|
||
<taxonomicName id="4C4C24EA4B032315F3EDF802094DF86A" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[151,340,1972,1997]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B938837B4B032315F3EDF80208F6F86B" box="[151,239,1972,1996]" italics="true" pageId="4" pageNumber="91">C. loebli</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (14);
|
||
<taxonomicName id="4C4C24EA4B032315F13AF8020B2AF86A" authority="(Walker)" authorityName="" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[576,819,1972,1997]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="monstrosus">
|
||
<emphasis id="B938837B4B032315F13AF8020AC9F86B" box="[576,720,1973,1996]" italics="true" pageId="4" pageNumber="91">C. monstrosus</emphasis>
|
||
(Walker)
|
||
</taxonomicName>
|
||
, lectotype female (15);
|
||
<taxonomicName id="4C4C24EA4B032315F759F8020D18F86A" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[1059,1281,1972,1997]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="mussardi">
|
||
<emphasis id="B938837B4B032315F759F8020C85F86B" box="[1059,1180,1972,1996]" italics="true" pageId="4" pageNumber="91">C. mussardi</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype male (16);
|
||
<taxonomicName id="4C4C24EA4B032315F3B5F86E09D5F856" authority="Besuchet" authorityName="Besuchet" authorityYear="1971" box="[207,460,2008,2033]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="91" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B032315F3B5F86E097FF857" box="[207,358,2008,2032]" italics="true" pageId="4" pageNumber="91">C. quadricollis</emphasis>
|
||
Besuchet
|
||
</taxonomicName>
|
||
, holotype female (17).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF35F694B022314F3BDFF2E0AADFF14" blockId="5.[151,1437,153,899]" box="[199,692,153,179]" pageId="5" pageNumber="92">Labrum with indistinct anteromedian notch.</paragraph>
|
||
<paragraph id="8BF35F694B022314F3BDFF0A0B35FE74" blockId="5.[151,1437,153,899]" pageId="5" pageNumber="92">
|
||
Pronotum subquadrangular, broadest near anterior margin; PL
|
||
<quantity id="4CB4F28C4B022314F0FDFF0B0C32FF70" box="[903,1067,188,215]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="9.8" metricValueMin="9.5" pageId="5" pageNumber="92" unit="mm" value="0.965" valueMax="0.98" valueMin="0.95">0.95–0.98 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F705FF0A0CF1FF70" box="[1151,1256,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="92" unit="mm" value="0.96">0.96 mm</quantity>
|
||
), PW
|
||
<quantity id="4CB4F28C4B022314F656FF0A08DAFF5C" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.13" metricValueMin="1.08" pageId="5" pageNumber="92" unit="mm" value="1.105" valueMax="1.13" valueMin="1.08">1.08–1.13 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F267FF560991FF5C" box="[285,392,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="5" pageNumber="92" unit="mm" value="1.11">1.11 mm</quantity>
|
||
); anterior and posterior corners obtuse-angled and blunt, but relatively well-defined; sides nearly straight and strongly convergent posterad; pronotal base with shallow and short diffuse median impression; posterolateral portions of pronotum protruding dorsoposterad (
|
||
<figureCitation id="137743EC4B022314F026FE9E0BB9FEE4" box="[860,928,297,323]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="5" pageNumber="92">Fig. 2</figureCitation>
|
||
) and overhanging posterior pronotal margin, interrupting transverse groove demarcating vestigial posterior collar, the latter reduced to short and narrow transverse area in posteromedian pronotal impression; lateral portions of transverse groove run along sides of pronotum, not visible in dorsal view. Punctures on disc similar to those on head; setae long, sparse and suberect. Cuticle between punctures on pronotal disc as glossy as that on head.
|
||
</paragraph>
|
||
<paragraph id="8BF35F694B022314F3BDFE6A0C13FD20" blockId="5.[151,1437,153,899]" pageId="5" pageNumber="92">
|
||
Elytra shield-shaped, broadest just behind humeri, EL
|
||
<quantity id="4CB4F28C4B022314F045FE6A0BFFFE50" box="[831,998,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.165" metricValueMax="2.2" metricValueMin="2.13" pageId="5" pageNumber="92" unit="mm" value="2.165" valueMax="2.2" valueMin="2.13">2.13–2.20 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F73AFE6A0CB2FE50" box="[1088,1195,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="92" unit="mm" value="2.18">2.18 mm</quantity>
|
||
), EW
|
||
<quantity id="4CB4F28C4B022314F78CFE6A0D84FE50" box="[1270,1437,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.665" metricValueMax="1.7" metricValueMin="1.63" pageId="5" pageNumber="92" unit="mm" value="1.665" valueMax="1.7" valueMin="1.63">1.63–1.70 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F39EFDB60955FDBC" box="[228,332,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="5" pageNumber="92" unit="mm" value="1.66">1.66 mm</quantity>
|
||
), EI 1.29–1.33; humeri well-defined, obtuse-angled; elytral apices unmodified, subtriangular; each elytron with five dorsal and one lateral rows of large and deep, densely and almost evenly distributed pits, cuticle between pits glossy, between rows with fine and sparse evenly distributed setiferous punctures; setae much longer and slightly thicker than those on head and pronotum, moderately dense, erect.
|
||
</paragraph>
|
||
<paragraph id="8BF35F694B022314F3BDFD260986FD0C" blockId="5.[151,1437,153,899]" box="[199,415,657,683]" pageId="5" pageNumber="92">Hind wings absent.</paragraph>
|
||
<paragraph id="8BF35F694B022314F3BDFD020D74FD68" blockId="5.[151,1437,153,899]" box="[199,1389,693,719]" pageId="5" pageNumber="92">
|
||
Legs long and slender; protrochanter (
|
||
<figureCitation id="137743EC4B022314F109FD020AAFFD68" box="[627,694,693,719]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="5" pageNumber="92">Fig. 2</figureCitation>
|
||
) modified, with robust anteroventral subtriangular projection.
|
||
</paragraph>
|
||
<paragraph id="8BF35F694B022314F3BDFD6E0A70FC24" blockId="5.[151,1437,153,899]" pageId="5" pageNumber="92">
|
||
Aedeagus (
|
||
<figureCitation id="137743EC4B022314F23CFD6E0982FD54" box="[326,411,728,755]" captionStart="FIGURES 18–21" captionStartId="5.[151,264,1737,1762]" captionTargetBox="[164,1419,922,1711]" captionTargetId="figure@5.[151,1436,913,1718]" captionTargetPageId="5" captionText="FIGURES 18–21. Clidicus minilankanus sp. n. Aedeagus in ventral (18) and lateral (19) views; sperm pump (20); ovipositor with spermatheca (21)." figureDoi="http://doi.org/10.5281/zenodo.3602268" httpUri="https://zenodo.org/record/3602268/files/figure.png" pageId="5" pageNumber="92">Figs 18</figureCitation>
|
||
–519) strongly elongate but moderately slender, in lateral view strongly curved, AeL
|
||
<quantity id="4CB4F28C4B022314F611FD6E08DFFCB0" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="92" unit="mm" value="0.8">0.80 mm</quantity>
|
||
; parameres long, symmetrical, in ventral view each paramere with subtriangular apex slightly curved mesad; flagellum differentiated from ductus ejaculatorius by a rapid broadening of walls near base of median lobe and gradually narrowing distad. 'Sperm pump' (
|
||
<figureCitation id="137743EC4B022314F102FCF20AD0FCF8" box="[632,713,837,863]" captionStart="FIGURES 18–21" captionStartId="5.[151,264,1737,1762]" captionTargetBox="[164,1419,922,1711]" captionTargetId="figure@5.[151,1436,913,1718]" captionTargetPageId="5" captionText="FIGURES 18–21. Clidicus minilankanus sp. n. Aedeagus in ventral (18) and lateral (19) views; sperm pump (20); ovipositor with spermatheca (21)." figureDoi="http://doi.org/10.5281/zenodo.3602268" httpUri="https://zenodo.org/record/3602268/files/figure.png" pageId="5" pageNumber="92">Fig. 20</figureCitation>
|
||
)
|
||
<quantity id="4CB4F28C4B022314F1ADFCF30B22FCF8" box="[727,827,836,863]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="92" unit="mm" value="0.52">0.52 mm</quantity>
|
||
in length, long and slender, with two shallow 'funnels', distal funnel narrower than proximal one.
|
||
</paragraph>
|
||
<caption id="DF330FE14B022314F3EDF97E0999F8A1" ID-DOI="http://doi.org/10.5281/zenodo.3602268" ID-Zenodo-Dep="3602268" httpUri="https://zenodo.org/record/3602268/files/figure.png" pageId="5" pageNumber="92" startId="5.[151,264,1737,1762]" targetBox="[164,1419,922,1711]" targetPageId="5">
|
||
<paragraph id="8BF35F694B022314F3EDF97E0999F8A1" blockId="5.[151,1437,1737,1798]" pageId="5" pageNumber="92">
|
||
<emphasis id="B938837B4B022314F3EDF97E094CF945" bold="true" box="[151,341,1737,1762]" pageId="5" pageNumber="92">FIGURES 18–21.</emphasis>
|
||
<taxonomicName id="4C4C24EA4B022314F226F97D0A59F945" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[348,576,1738,1762]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="92" phylum="Arthropoda" rank="species" species="minilankanus" status="sp. nov.">
|
||
<emphasis id="B938837B4B022314F226F97D0A59F945" box="[348,576,1738,1762]" italics="true" pageId="5" pageNumber="92">Clidicus minilankanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B3E004B022314F13DF97D0A98F945" box="[583,641,1738,1762]" pageId="5" pageNumber="92" rank="species">
|
||
<emphasis id="B938837B4B022314F13DF97D0A98F945" bold="true" box="[583,641,1738,1762]" pageId="5" pageNumber="92">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Aedeagus in ventral (18) and lateral (19) views; sperm pump (20); ovipositor with spermatheca (21).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF35F694B022314F3BDF8980D48F812" blockId="5.[151,1437,1839,2046]" pageId="5" pageNumber="92">
|
||
<emphasis id="B938837B4B022314F3BDF898093CF8EE" bold="true" box="[199,293,1839,1865]" pageId="5" pageNumber="92">Female.</emphasis>
|
||
Externally very similar to male, on average indistinctly smaller and with broader elytra; protrochanters unmodified; antennae slightly shorter in relation to BL; BL
|
||
<quantity id="4CB4F28C4B022314F01BF8E40C1DF8CA" box="[865,1028,1875,1901]" metricMagnitude="-3" metricUnit="m" metricValue="3.705" metricValueMax="3.73" metricValueMin="3.68" pageId="5" pageNumber="92" unit="mm" value="3.705" valueMax="3.73" valueMin="3.68">3.68–3.73 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F72DF8E40CA7F8CA" box="[1111,1214,1875,1901]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="5" pageNumber="92" unit="mm" value="3.7">3.70 mm</quantity>
|
||
); HL
|
||
<quantity id="4CB4F28C4B022314F787F8E50D7AF8CA" box="[1277,1379,1874,1901]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="92" unit="mm" value="0.65">0.65 mm</quantity>
|
||
, HW
|
||
<quantity id="4CB4F28C4B022314F3EDF8C00924F836" box="[151,317,1911,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" metricValueMax="1.1" metricValueMin="1.08" pageId="5" pageNumber="92" unit="mm" value="1.09" valueMax="1.1" valueMin="1.08">1.08–1.10 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F2EFF8C009E6F836" box="[405,511,1911,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="5" pageNumber="92" unit="mm" value="1.09">1.09 mm</quantity>
|
||
), AnL
|
||
<quantity id="4CB4F28C4B022314F134F8C10AEDF836" box="[590,756,1910,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" metricValueMax="1.75" metricValueMin="1.73" pageId="5" pageNumber="92" unit="mm" value="1.74" valueMax="1.75" valueMin="1.73">1.73–1.75 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F036F8C00BAFF836" box="[844,950,1911,1937]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="5" pageNumber="92" unit="mm" value="1.74">1.74 mm</quantity>
|
||
); PL
|
||
<quantity id="4CB4F28C4B022314F08FF8C10C82F836" box="[1013,1179,1910,1937]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="9.8" metricValueMin="9.5" pageId="5" pageNumber="92" unit="mm" value="0.965" valueMax="0.98" valueMin="0.95">0.95–0.98 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F789F8C00D44F836" box="[1267,1373,1911,1937]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="92" unit="mm" value="0.96">0.96 mm</quantity>
|
||
), PW
|
||
<quantity id="4CB4F28C4B022314F3EDF82C0923F812" box="[151,314,1947,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.13" metricValueMin="1.08" pageId="5" pageNumber="92" unit="mm" value="1.105" valueMax="1.13" valueMin="1.08">1.08–1.13 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F2F4F82C09EFF812" box="[398,502,1947,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="92" unit="mm" value="1.1">1.10 mm</quantity>
|
||
); EL
|
||
<quantity id="4CB4F28C4B022314F148F82D0ACEF812" box="[562,727,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="2.09" metricValueMax="2.13" metricValueMin="2.05" pageId="5" pageNumber="92" unit="mm" value="2.09" valueMax="2.13" valueMin="2.05">2.05–2.13 mm</quantity>
|
||
, EW
|
||
<quantity id="4CB4F28C4B022314F06EF82C0BAEF812" box="[788,951,1947,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.64" metricValueMax="1.68" metricValueMin="1.6" pageId="5" pageNumber="92" unit="mm" value="1.64" valueMax="1.68" valueMin="1.6">1.60–1.68 mm</quantity>
|
||
(mean
|
||
<quantity id="4CB4F28C4B022314F771F82D0C6AF812" box="[1035,1139,1946,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="92" unit="mm" value="1.65">1.65 mm</quantity>
|
||
), EI
|
||
<quantity id="4CB4F28C4B022314F7DDF82C0D54F812" box="[1191,1357,1947,1973]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" metricValueMax="1.28" metricValueMin="1.24" pageId="5" pageNumber="92" unit="mm" value="1.26" valueMax="1.28" valueMin="1.24">1.24–1.28 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF35F694B022317F3BDF8080965FF70" blockId="5.[151,1437,1839,2046]" lastBlockId="6.[151,1437,153,611]" lastPageId="6" lastPageNumber="93" pageId="5" pageNumber="92">
|
||
Spermatheca (
|
||
<figureCitation id="137743EC4B022314F21DF80809B3F87E" box="[359,426,1983,2009]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="5" pageNumber="92">Fig. 2</figureCitation>
|
||
) with oval capsular part with asymmetrical, subconical and long projection that surrounds insertion site of ductus spermathecae, width (diameter) of capsular part
|
||
<quantity id="4CB4F28C4B022314F0BBF8550C31F85A" box="[961,1064,2018,2045]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="92" unit="mm" value="0.5">0.50 mm</quantity>
|
||
; accessory gland inserted at base of subconical projection; ductus spermathecae relatively short and weakly entangled; bursa copulatrix short, tubular, forming a semi-coil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3560CE24B012317F3BDFF570A26FF5C" box="[199,575,224,251]" pageId="6" pageNumber="93" type="distribution">
|
||
<paragraph id="8BF35F694B012317F3BDFF570A26FF5C" blockId="6.[151,1437,153,611]" box="[199,575,224,251]" pageId="6" pageNumber="93">
|
||
<emphasis id="B938837B4B012317F3BDFF570978FF5C" bold="true" box="[199,353,224,251]" pageId="6" pageNumber="93">Distribution.</emphasis>
|
||
Western
|
||
<collectingCountry id="F35B1FF94B012317F2B3FF560A22FF5C" box="[457,571,225,251]" name="Sri Lanka" pageId="6" pageNumber="93">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3560CE24B012317F3BDFEB208C2FEE4" pageId="6" pageNumber="93" type="etymology">
|
||
<paragraph id="8BF35F694B012317F3BDFEB208C2FEE4" blockId="6.[151,1437,153,611]" pageId="6" pageNumber="93">
|
||
<emphasis id="B938837B4B012317F3BDFEB20957FEB8" bold="true" box="[199,334,261,287]" pageId="6" pageNumber="93">Etymology.</emphasis>
|
||
The name refers to the smallest body among all
|
||
<collectingRegion id="4988918B4B012317F0E9FEB20BADFEB8" box="[915,948,261,287]" country="United Kingdom" name="Surrey" pageId="6" pageNumber="93">Sri</collectingRegion>
|
||
Lankan
|
||
<taxonomicName id="4C4C24EA4B012317F767FEB20C63FEB8" box="[1053,1146,261,287]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938837B4B012317F767FEB20C63FEB8" box="[1053,1146,261,287]" italics="true" pageId="6" pageNumber="93">Clidicus</emphasis>
|
||
</taxonomicName>
|
||
species, and the country name.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3560CE24B012317F3BDFEFA0BF2FDC4" pageId="6" pageNumber="93" type="discussion">
|
||
<paragraph id="8BF35F694B012317F3BDFEFA0BF2FDC4" blockId="6.[151,1437,153,611]" pageId="6" pageNumber="93">
|
||
<emphasis id="B938837B4B012317F3BDFEFA0922FEC0" bold="true" box="[199,315,333,359]" pageId="6" pageNumber="93">Remarks.</emphasis>
|
||
<taxonomicName id="4C4C24EA4B012317F239FEFA0A26FEC0" authorityName="Jałoszyński" authorityYear="2020" box="[323,575,333,359]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="minilankanus">
|
||
<emphasis id="B938837B4B012317F239FEFA0A26FEC0" box="[323,575,333,359]" italics="true" pageId="6" pageNumber="93">Clidicus minilankanus</emphasis>
|
||
</taxonomicName>
|
||
can easily be distinguished from its sympatric congeners on the basis of the smallest body (the largest specimen examined is
|
||
<quantity id="4CB4F28C4B012317F1BFFEC60B32FE2C" box="[709,811,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="3.83" pageId="6" pageNumber="93" unit="mm" value="3.83">3.83 mm</quantity>
|
||
long;
|
||
<taxonomicName id="4C4C24EA4B012317F00FFEC60BE4FE2C" authorityName="Besuchet" authorityYear="1971" box="[885,1021,369,395]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="mussardi">
|
||
<emphasis id="B938837B4B012317F00FFEC60BE4FE2C" box="[885,1021,369,395]" italics="true" pageId="6" pageNumber="93">C. mussardi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C24EA4B012317F74CFEC60CC6FE2C" authorityName="Besuchet" authorityYear="1971" box="[1078,1247,369,395]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B012317F74CFEC60CC6FE2C" box="[1078,1247,369,395]" italics="true" pageId="6" pageNumber="93">C. quadricollis</emphasis>
|
||
</taxonomicName>
|
||
exceed
|
||
<quantity id="4CB4F28C4B012317F644FEC60D81FE2C" box="[1342,1432,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="93" unit="mm" value="4.8">4.8 mm</quantity>
|
||
, adults of
|
||
<taxonomicName id="4C4C24EA4B012317F387FE220944FE08" authorityName="Besuchet" authorityYear="1971" box="[253,349,405,431]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="loebli">
|
||
<emphasis id="B938837B4B012317F387FE220944FE08" box="[253,349,405,431]" italics="true" pageId="6" pageNumber="93">C. loebli</emphasis>
|
||
</taxonomicName>
|
||
are nearly
|
||
<quantity id="4CB4F28C4B012317F2ACFE230A32FE08" box="[470,555,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="93" unit="mm" value="6.5">6.5 mm</quantity>
|
||
long, and those of
|
||
<taxonomicName id="4C4C24EA4B012317F186FE220B83FE08" authorityName="" baseAuthorityName="Walker" baseAuthorityYear="1858" box="[764,922,405,431]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="monstrosus">
|
||
<emphasis id="B938837B4B012317F186FE220B83FE08" box="[764,922,405,431]" italics="true" pageId="6" pageNumber="93">C. monstrosus</emphasis>
|
||
</taxonomicName>
|
||
exceed
|
||
<quantity id="4CB4F28C4B012317F089FE220C55FE08" box="[1011,1100,405,431]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="93" unit="mm" value="7.7">7.7 mm</quantity>
|
||
). It is the only
|
||
<collectingRegion id="4988918B4B012317F797FE220D17FE08" box="[1261,1294,405,431]" country="United Kingdom" name="Surrey" pageId="6" pageNumber="93">Sri</collectingRegion>
|
||
Lankan species with well-defined, obtuse-angled humeri (
|
||
<figureCitation id="137743EC4B012317F1E5FE0E0AC6FE74" box="[671,735,441,467]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="6" pageNumber="93">Fig. 1</figureCitation>
|
||
); all remaining species have the humeri broadly rounded (
|
||
<figureCitation id="137743EC4B012317F610FE0E08DCFE5F" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="6" pageNumber="93">Figs 3–6</figureCitation>
|
||
).
|
||
<taxonomicName id="4C4C24EA4B012317F3A2FE6A09CAFE50" authorityName="Jałoszyński" authorityYear="2020" box="[216,467,477,503]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="minilankanus">
|
||
<emphasis id="B938837B4B012317F3A2FE6A09CAFE50" box="[216,467,477,503]" italics="true" pageId="6" pageNumber="93">Clidicus minilankanus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C4C24EA4B012317F1DAFE6A0B5EFE50" authorityName="Besuchet" authorityYear="1971" box="[672,839,477,503]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B012317F1DAFE6A0B5EFE50" box="[672,839,477,503]" italics="true" pageId="6" pageNumber="93">C. quadricollis</emphasis>
|
||
</taxonomicName>
|
||
, with which it shares a similar shape of the pronotum with the unique posteromedian impression (
|
||
<figureCitation id="137743EC4B012317F1F3FDB60AE6FDBC" box="[649,767,513,539]" captionStart="FIGURES 7–12" captionStartId="3.[151,264,1900,1925]" captionTargetBox="[158,1430,409,1880]" captionTargetId="figure@3.[158,1430,409,1880]" captionTargetPageId="3" captionText="FIGURES 7–12. Clidicus of Sri Lanka; head and pronotum in dorsal (7–10, 12) and posterodorsal (11) views. Clidicus mi- nilankanus sp. n., holotype male (7); C. loebli Besuchet, holotype female (8); C. monstrosus (Walker), lectotype female (9); C. mussardi Besuchet, holotype male (10–11; arrow showing transverse groove demarcating pronotal collar); C. quadricollis Besuchet, holotype female (12)." figureDoi="http://doi.org/10.5281/zenodo.3602264" httpUri="https://zenodo.org/record/3602264/files/figure.png" pageId="6" pageNumber="93">Figs 7, 12</figureCitation>
|
||
), and a similar shape of the elytra (
|
||
<figureCitation id="137743EC4B012317F7EEFDB60CE5FDBB" box="[1172,1276,513,540]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1886,1911]" captionTargetBox="[157,1427,188,1830]" captionTargetId="figure@2.[151,1436,174,1842]" captionTargetPageId="2" captionText="FIGURES 1–6. Clidicus of Sri Lanka; dorsal habitus (1, 3–6) and protrochanter in lateral view (2). Clidicus minilankanus sp. n., holotype male (1) and paratype male (2; arrow showing modified protrochanter); C. loebli Besuchet, holotype female (3); C. monstrosus (Walker), lectotype female (4); C. mussardi Besuchet, holotype male (5); C. quadricollis Besuchet, holotype female (6)." figureDoi="http://doi.org/10.5281/zenodo.3602262" httpUri="https://zenodo.org/record/3602262/files/figure.png" pageId="6" pageNumber="93">Figs 1, 6</figureCitation>
|
||
). The male of
|
||
<taxonomicName id="4C4C24EA4B012317F3EDFD920926FD98" authorityName="Besuchet" authorityYear="1971" box="[151,319,549,575]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="quadricollis">
|
||
<emphasis id="B938837B4B012317F3EDFD920926FD98" box="[151,319,549,575]" italics="true" pageId="6" pageNumber="93">C. quadricollis</emphasis>
|
||
</taxonomicName>
|
||
remains unknown, but females of
|
||
<taxonomicName id="4C4C24EA4B012317F1BEFD920B62FD98" authorityName="Jałoszyński" authorityYear="2020" box="[708,891,549,575]" class="Insecta" family="Staphylinidae" genus="Clidicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="93" phylum="Arthropoda" rank="species" species="minilankanus">
|
||
<emphasis id="B938837B4B012317F1BEFD920B62FD98" box="[708,891,549,575]" italics="true" pageId="6" pageNumber="93">C. minilankanus</emphasis>
|
||
</taxonomicName>
|
||
have the same shape of the humeri as males and consequently the shape of the elytra in these two species is clearly different.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |