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<document id="1597BA5A02613372BF61749E5C6ADA46" ID-CLB-Dataset="6447" ID-DOI="10.11646/zootaxa.4969.1.7" ID-GBIF-Dataset="565d09b1-8934-4ecd-8e46-aebdbf3c801f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4745993" ID-ZooBank="997EC67F-7A07-4C40-93BC-56F8505CAE26" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620647120386" checkinUser="plazi" docAuthor="Cutrim, Marcelo, Neto, Alberto Moreira Da Silva, García Aldrete, Alfonso N. &amp; Rafael, José Albertino" docDate="2021" docId="039FF66CFFD47418FF09D71A683BFE03" docLanguage="en" docName="zootaxa.4969.1.7.pdf" docOrigin="Zootaxa 4969 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loneura willineri Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael 2021, n. sp." docType="treatment" docUuid="E0ECEE81-AD25-405F-97AA-B49FF26589CE" docUuidSource="ZooBank" docVersion="8" lastPageNumber="140" masterDocId="FFA68E14FFD5741DFF9ED63D6C7FFF8B" masterDocTitle="A new species of Loneura Navás and taxonomic update of L. boliviana Williner and L. meridionalis García Aldrete (Psocodea: Psocomorpha: Ptiloneuridae)" masterLastPageNumber="148" masterPageNumber="135" pageNumber="136" updateTime="1698938531557" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="920103130D750826708829DF1C566468">A new species of Loneura Navás and taxonomic update of L. boliviana Williner and L. meridionalis García Aldrete (Psocodea: Psocomorpha: Ptiloneuridae)</mods:title>
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<mods:namePart id="1AE4410DAE27C65112D00EBB29529BEA">Cutrim, Marcelo</mods:namePart>
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<mods:affiliation id="320A28D573E70F933E3E94BB7AB6E3F7">Instituto Nacional de Pesquisas da Amazônia-INPA, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas, BRASIL. marcelocutrim @ gmail. com; https: // orcid. org / 0000 - 0001 - 7147 - 5300</mods:affiliation>
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<mods:namePart id="4251F2A2920B6DA37292B5AF2B517750">Neto, Alberto Moreira Da Silva</mods:namePart>
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<mods:affiliation id="52BB4C0C3FA53F7128224BE55A9DBE46">Instituto Nacional de Pesquisas da Amazônia-INPA, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas, BRASIL. marcelocutrim @ gmail. com; https: // orcid. org / 0000 - 0001 - 7147 - 5300 &amp; bio. alberto @ gmail. com; https: // orcid. org / 0000 - 0002 - 4522 - 3756</mods:affiliation>
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<mods:namePart id="9219705C0A00F8900A34BF450AEFF6B9">García Aldrete, Alfonso N.</mods:namePart>
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<mods:affiliation id="7A298CD8C13AEE1923125F40054EC3C1">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, Ciudad de México, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
<mods:nameIdentifier id="3C9C3FA63A5BBCCD5E28D5EC587C3F7E" type="email">anga@ib.unam.mx</mods:nameIdentifier>
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<mods:namePart id="5E0606275DF3619F57A25F45BFB005C9">Rafael, José Albertino</mods:namePart>
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<mods:affiliation id="B6C4E78EE8B30F0EFC26BA2B48316CB9">Instituto Nacional de Pesquisas da Amazônia-INPA, Programa de Pós-Graduação em Entomologia, Manaus, Amazonas, BRASIL. marcelocutrim @ gmail. com; https: // orcid. org / 0000 - 0001 - 7147 - 5300 &amp; jarafael @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<treatment id="039FF66CFFD47418FF09D71A683BFE03" ID-DOI="http://doi.org/10.5281/zenodo.4745995" ID-GBIF-Taxon="182398201" ID-Zenodo-Dep="4745995" LSID="urn:lsid:zoobank.org:act:E0ECEE81-AD25-405F-97AA-B49FF26589CE" httpUri="http://treatment.plazi.org/id/039FF66CFFD47418FF09D71A683BFE03" lastPageId="5" lastPageNumber="140" pageId="1" pageNumber="136">
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<paragraph id="8B89477AFFD4741CFF09D71A6DC9FECA" blockId="1.[151,438,294,357]" box="[151,438,294,321]" pageId="1" pageNumber="136">
<heading id="D0C1F016FFD4741CFF09D71A6DC9FECA" bold="true" box="[151,438,294,321]" fontSize="11" level="1" pageId="1" pageNumber="136" reason="1">
<emphasis id="B9429B68FFD4741CFF09D71A6DC9FECA" bold="true" box="[151,438,294,321]" pageId="1" pageNumber="136">
<taxonomicName id="4C363CF9FFD4741CFF09D71A6D13FECA" authority="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael, 2021" authorityName="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael" authorityYear="2021" box="[151,364,294,321]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="willineri" status="sp. nov.">
<emphasis id="B9429B68FFD4741CFF09D71A6D13FECA" bold="true" box="[151,364,294,321]" italics="true" pageId="1" pageNumber="136">Loneura willineri</emphasis>
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<taxonomicNameLabel id="A2712613FFD4741CFEEDD71A6DC9FECA" box="[371,438,295,321]" pageId="1" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32C14F1FFD4741CFF09D7766D66FEEE" box="[151,281,331,357]" pageId="1" pageNumber="136" type="description">
<paragraph id="8B89477AFFD4741CFF09D7766D66FEEE" blockId="1.[151,438,294,357]" box="[151,281,331,357]" pageId="1" pageNumber="136">
(
<figureCitation id="130D5BFFFFD4741CFF01D7766D6EFEEE" box="[159,273,331,357]" captionStart-0="FIGURES 17" captionStart-1="FIGURES 814" captionStart-2="FIGURES 1521" captionStartId-0="2.[151,264,1887,1911]" captionStartId-1="3.[151,264,1949,1973]" captionStartId-2="4.[151,264,1860,1884]" captionTargetBox-0="[166,1421,219,1860]" captionTargetBox-1="[209,1389,197,1925]" captionTargetBox-2="[171,1419,540,1837]" captionTargetId-0="figure-20@2.[166,1421,179,1860]" captionTargetId-1="figure-17@3.[191,1396,181,1925]" captionTargetId-2="figure-247@4.[151,1436,525,1837]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." captionText-1="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." captionText-2="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.4745997" figureDoi-1="http://doi.org/10.5281/zenodo.4745999" figureDoi-2="http://doi.org/10.5281/zenodo.4746003" httpUri-0="https://zenodo.org/record/4745997/files/figure.png" httpUri-1="https://zenodo.org/record/4745999/files/figure.png" httpUri-2="https://zenodo.org/record/4746003/files/figure.png" pageId="1" pageNumber="136">Figs 121</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C14F1FFD4741CFF09D7AF694BFADE" pageId="1" pageNumber="136" type="diagnosis">
<paragraph id="8B89477AFFD4741CFF09D7AF6844FD92" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF09D7AF6D6DFE27" bold="true" box="[151,274,402,428]" pageId="1" pageNumber="136">Diagnosis.</emphasis>
Belonging in species group IB of
<bibRefCitation id="EFA73A8BFFD4741CFD0CD7AE6FB6FE26" author="Garcia Aldrete, A. N. &amp; Gonzalez Obando, R. &amp; Sarria Sarria, F. A." box="[658,969,403,429]" pageId="1" pageNumber="136" pagination="55 - 62" refId="ref5766" refString="Garcia Aldrete, A. N., Gonzalez Obando, R. &amp; Sarria Sarria, F. A. (2011) Three new species of Loneura (Psocodea: ' Psocoptera': Ptiloneuridae) from Gorgona Island, Cauca, Colombia, with a new infrageneric classification. Zootaxa, 3050 (1), 55 - 62. https: // doi. org / 10.11646 / zootaxa. 3050.1.3" type="journal article" year="2011">
García Aldrete
<emphasis id="B9429B68FFD4741CFCA0D7AE6F10FE26" box="[830,879,403,429]" italics="true" pageId="1" pageNumber="136">et al</emphasis>
. (2011)
</bibRefCitation>
, this species is related to
<taxonomicName id="4C363CF9FFD4741CFB7BD7AE6912FE26" authorityName="Williner" authorityYear="1949" box="[1253,1389,403,429]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="B9429B68FFD4741CFB7BD7AE6912FE26" box="[1253,1389,403,429]" italics="true" pageId="1" pageNumber="136">L. boliviana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C363CF9FFD4741CFF09D78A6D3DFE5A" box="[151,322,439,465]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="1" pageNumber="136" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B9429B68FFD4741CFF09D78A6D3DFE5A" box="[151,322,439,465]" italics="true" pageId="1" pageNumber="136">L. meridionalis</emphasis>
</taxonomicName>
by the hyaline wings and hypandrium arrangement but differs from it in having the phallosome with external parameres long and slender, in having the anterior pair of endophallic sclerites basally straight, besides having an anterior central projection with a group of spines along the outer margin.
</paragraph>
<paragraph id="8B89477AFFD4741CFF59D41E691EFD46" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D41E6D21FDB7" bold="true" box="[199,350,546,573]" pageId="1" pageNumber="136">Male. Color.</emphasis>
Compound eyes black, ocelli hyaline, with ochre centripetal crescents; head pattern (
<figureCitation id="130D5BFFFFD4741CFAD8D41E69F4FDB6" box="[1350,1419,547,573]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 1</figureCitation>
). Scape brown, pedicel pale brown, f1f3 pale yellow. Femora pale with a brown belt at proximal ends; tibiae pale yellow with distal ends brown; tarsomeres 1 pale brown with distal ends brown, tarsomeres 23 brown. Forewings (
<figureCitation id="130D5BFFFFD4741CFF3ED4B26C9DFD22" box="[160,226,655,681]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 2</figureCitation>
) almost hyaline, veins brown, pterostigma with proximal and distal pale brown bands, a brown spot at RRs crossvein and at confluence of CuP1A. Hindwings (
<figureCitation id="130D5BFFFFD4741CFD75D48E6F51FD46" box="[747,814,691,717]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
) hyaline, veins brown, a brown spot at wing base.
</paragraph>
<paragraph id="8B89477AFFD4741CFF59D4EA694BFADE" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D4EA6D1EFD7A" bold="true" box="[199,353,727,753]" pageId="1" pageNumber="136">Morphology.</emphasis>
Compound eyes without interommatidial setae. Outer cusp of lacinial tips (
<figureCitation id="130D5BFFFFD4741CFB5BD4EA6975FD7A" box="[1221,1290,727,753]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 4</figureCitation>
) broad, with five denticles. Right forewing (
<figureCitation id="130D5BFFFFD4741CFE69D4C66E44FC9E" box="[503,571,763,789]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 2</figureCitation>
) pterostigma elongate, narrow anteriorly, wider in the middle; areola postica tall, broadly triangular, apically round; Rs long, slightly concave proximally, R
<subScript id="17B2453FFFD4741CFBB0D50C6838FCB4" attach="left" box="[1070,1095,817,831]" fontSize="6" pageId="1" pageNumber="136">2+3</subScript>
almost straight, R
<subScript id="17B2453FFFD4741CFA85D50C694AFCB4" attach="left" box="[1307,1333,817,831]" fontSize="6" pageId="1" pageNumber="136">4+5</subScript>
sinuous; M stem slightly concave proximally, then almost straight, with six primary branches, M
<subScript id="17B2453FFFD4741CFB1FD56868F6FCE8" attach="left" box="[1153,1161,853,867]" fontSize="6" pageId="1" pageNumber="136">6</subScript>
forked, resulting in M
<subScript id="17B2453FFFD4741CFA13D56869E3FCE8" attach="left" box="[1421,1436,853,867]" fontSize="6" pageId="1" pageNumber="136">6a</subScript>
and M
<subScript id="17B2453FFFD4741CFF7ED5446C8FFC0C" attach="left" box="[224,240,889,903]" fontSize="6" pageId="1" pageNumber="136">6b</subScript>
. Right hindwing (
<figureCitation id="130D5BFFFFD4741CFE21D55A6E7DFC0A" box="[447,514,871,897]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 3</figureCitation>
) M with four primary branches, M
<subScript id="17B2453FFFD4741CFC10D5446FE9FC0C" attach="left" box="[910,918,889,903]" fontSize="6" pageId="1" pageNumber="136">4</subScript>
forked, resulting in M
<subScript id="17B2453FFFD4741CFB0BD54468DAFC0C" attach="left" box="[1173,1189,889,903]" fontSize="6" pageId="1" pageNumber="136">4a</subScript>
and M
<subScript id="17B2453FFFD4741CFB6BD544697AFC0C" attach="left" box="[1269,1285,889,903]" fontSize="6" pageId="1" pageNumber="136">4b</subScript>
(
<figureCitation id="130D5BFFFFD4741CFA8BD55A69E7FC0A" box="[1301,1432,870,897]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="1" pageNumber="136">Figs 1521</figureCitation>
; see also variations of the wings of female specimens). Hypandrium (
<figureCitation id="130D5BFFFFD4741CFC3CD5B66F99FC2E" box="[930,998,906,933]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 5</figureCitation>
) of three sclerites, central sclerite “U” shaped, convex anteriorly, with two lateral posterior projections, rounded distally, curved outward; irregular side sclerites narrow distally, setae as illustrated. Phallosome (
<figureCitation id="130D5BFFFFD4741CFCB8D5EE6F14FC65" box="[806,875,979,1006]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 6</figureCitation>
) with side struts basally not fused, V-shaped, not fused posteriorly to external parameres, these long and slender, proximally slender and widening distally ending in a curved inward projection with a field of pores; two pairs of endophallic sclerites; an anterior pair U-shaped, anteriorly straight and serrated at the base, with two projections, a central inner shorter, very thin distally, posteriorly with a sclerotized area and a spine group along the outer margin, and a lateral external longer and curved outward, with a sclerotized area on both inner and external margins, posteriorly with a short and slightly acuminate projection, laterally with a radular posterior margin and a lobed end; two partially sclerotized membranes curved outward between the base of the anterior pair of endophallic sclerites; a posterior pair elongate, irregular, proximally concave with a very thin curved, apical end scythe-shaped and distally turned outward with short acuminated end projection. Epiproct (
<figureCitation id="130D5BFFFFD4741CFE98D32A6D36FABA" box="[262,329,1303,1329]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 7</figureCitation>
) trapezoidal, posteriorly slightly rounded, with two lateral large setae as illustrated. Right paraproct (
<figureCitation id="130D5BFFFFD4741CFF3ED3066C9CFADE" box="[160,227,1339,1365]" captionStart="FIGURES 17" captionStartId="2.[151,264,1887,1911]" captionTargetBox="[166,1421,219,1860]" captionTargetId="figure-20@2.[166,1421,179,1860]" captionTargetPageId="2" captionText="FIGURES 17. Loneura willineri n. sp. Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="1" pageNumber="136">Fig. 7</figureCitation>
) broad-based, triangular, sensory fields with 34 trichobothria on basal rosettes, setae as illustrated.
</paragraph>
</subSubSection>
<subSubSection id="C32C14F1FFD47419FF59D3626824FE7E" lastPageId="4" lastPageNumber="139" pageId="1" pageNumber="136" type="description">
<paragraph id="8B89477AFFD4741CFF59D3626F60FA16" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D3626D01FAF2" bold="true" box="[199,382,1375,1401]" pageId="1" pageNumber="136">Measurements.</emphasis>
FW: 5785, HW: 3914, F: 1478, T: 2489, t1: 1020, t2: 53, t3: 136, f1: 1296, f2: 1303, Mx4: 334, IO: 650, D: 409, d: 255, PO: 0.62. IO/d: 2.54, IO/D: 1.58.
</paragraph>
<paragraph id="8B89477AFFD4741CFF59D39A6EDFFA4A" blockId="1.[151,1437,402,2013]" box="[199,672,1446,1473]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D39A6D0AFA4B" bold="true" box="[199,373,1446,1473]" pageId="1" pageNumber="136">Female. Color.</emphasis>
Essentially as in the male.
</paragraph>
<paragraph id="8B89477AFFD4741CFF59D3F6695CF81E" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D3F66D1EFA6E" bold="true" box="[199,353,1483,1509]" pageId="1" pageNumber="136">Morphology.</emphasis>
Compound eyes (
<figureCitation id="130D5BFFFFD4741CFDA6D3F66E01FA6E" box="[568,638,1483,1509]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 8</figureCitation>
) without interommatidial setae. Outer cusp of lacinial tips (
<figureCitation id="130D5BFFFFD4741CFADED3F669ECFA6E" box="[1344,1427,1483,1509]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 11</figureCitation>
) broad, with five denticles. Right forewing (
<figureCitation id="130D5BFFFFD4741CFDE6D3D26EC4F982" box="[632,699,1519,1545]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 9</figureCitation>
) pterostigma elongate, narrow basally, wider in the middle; areola postica tall, trapezoid, apically rounded; Rs long, slightly concave basally, R
<subScript id="17B2453FFFD4741CFC7DD0186F83F9B8" attach="left" box="[995,1020,1573,1587]" fontSize="6" pageId="1" pageNumber="136">2+3</subScript>
almost straight, R
<subScript id="17B2453FFFD4741CFB58D018689FF9B8" attach="left" box="[1222,1248,1573,1587]" fontSize="6" pageId="1" pageNumber="136">4+5</subScript>
sinuous; M stem slightly concave proximally, then almost straight, with seven primary branches, M
<subScript id="17B2453FFFD4741CFBB8D0746851F9DC" attach="left" box="[1062,1070,1609,1623]" fontSize="6" pageId="1" pageNumber="136">7</subScript>
forked, resulting in M
<subScript id="17B2453FFFD4741CFAB9D0746949F9DC" attach="left" box="[1319,1334,1609,1623]" fontSize="6" pageId="1" pageNumber="136">7a</subScript>
and M
<subScript id="17B2453FFFD4741CFA1AD07469EBF9DC" attach="left" box="[1412,1428,1609,1623]" fontSize="6" pageId="1" pageNumber="136">7b</subScript>
; Right hindwing (
<figureCitation id="130D5BFFFFD4741CFECBD0666DD8F9FE" box="[341,423,1627,1653]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 10</figureCitation>
) M with three primary branches (
<figureCitation id="130D5BFFFFD4741CFC81D0666FDEF9FE" box="[799,929,1626,1653]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="1" pageNumber="136">Figs 1521</figureCitation>
; see also variations of the wing female specimens). Subgenital plate (
<figureCitation id="130D5BFFFFD4741CFE34D0426D84F912" box="[426,507,1663,1689]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 12</figureCitation>
) broad, straight basally, subtriangular, with sides converging to a pointed apex, with lateral pigmented areas, setae directed to the inner and anterior area as illustrated. Ninth sternum (
<figureCitation id="130D5BFFFFD4741CFB7ED09E694FF936" box="[1248,1328,1699,1725]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 13</figureCitation>
) strongly sclerotized, subtrapezoidal, anteriorly with a deep central cleft, posteriorly straight at base, with two latero-posterior elliptic areas and other almost square strongly pigmented at its center. Gonapophyses (
<figureCitation id="130D5BFFFFD4741CFBC1D0D668CDF88E" box="[1119,1202,1771,1797]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 13</figureCitation>
): v1 long, with outer margin heavily sclerotized, slightly wider in the middle and narrowing at the end, with acuminate apex; v
<subScript id="17B2453FFFD4741CFAA0D11C6928F8A4" attach="left" box="[1342,1367,1825,1839]" fontSize="6" pageId="1" pageNumber="136">2+3</subScript>
stout, with a large heel proximally, wide at the middle and narrowing at the end with three large setae on outer lobes and long acuminate apex. Epiproct (
<figureCitation id="130D5BFFFFD4741CFD9DD16A6E29F8FA" box="[515,598,1879,1905]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 14</figureCitation>
) wide-based, rounded and with setae anteriorly as illustrated. Right Paraproct (
<figureCitation id="130D5BFFFFD4741CFF7ED1466D4EF81E" box="[224,305,1915,1941]" captionStart="FIGURES 814" captionStartId="3.[151,264,1949,1973]" captionTargetBox="[209,1389,197,1925]" captionTargetId="figure-17@3.[191,1396,181,1925]" captionTargetPageId="3" captionText="FIGURES 814. Loneura willineri n. sp. Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="1" pageNumber="136">Fig. 14</figureCitation>
) subtrapezoidal, sensory fields with 27 trichobothria on basal rosettes, setae as illustrated.
</paragraph>
<paragraph id="8B89477AFFD4741CFF59D1A26F67F856" blockId="1.[151,1437,402,2013]" pageId="1" pageNumber="136">
<emphasis id="B9429B68FFD4741CFF59D1A26D01F832" bold="true" box="[199,382,1951,1977]" pageId="1" pageNumber="136">Measurements.</emphasis>
FW: 6060, HW: 4018, F: 1208, T: 1503, t1: 706, t2: 84, t3: 138, f1: 1167, f2: 1269, Mx4: 332, IO: 670, D: 382, d: 270, PO: 0.70, IO/D: 1.75 IO/d: 2.48.
</paragraph>
<caption id="DF4917F2FFD7741FFF09D1626F8AF811" ID-DOI="http://doi.org/10.5281/zenodo.4745997" ID-Zenodo-Dep="4745997" httpUri="https://zenodo.org/record/4745997/files/figure.png" pageId="2" pageNumber="137" startId="2.[151,264,1887,1911]" targetBox="[166,1421,219,1860]" targetPageId="2">
<paragraph id="8B89477AFFD7741FFF09D1626F8AF811" blockId="2.[151,1436,1887,1947]" pageId="2" pageNumber="137">
<emphasis id="B9429B68FFD7741FFF09D1626D44F8FC" bold="true" box="[151,315,1887,1911]" pageId="2" pageNumber="137">FIGURES 17.</emphasis>
<taxonomicName id="4C363CF9FFD7741FFEDCD15D6D8AF8FC" authority="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael, 2021" authorityName="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael" authorityYear="2021" box="[322,501,1887,1911]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="137" phylum="Arthropoda" rank="species" species="willineri" status="sp. nov.">
<emphasis id="B9429B68FFD7741FFEDCD15D6D8AF8FC" box="[322,501,1887,1911]" italics="true" pageId="2" pageNumber="137">Loneura willineri</emphasis>
</taxonomicName>
<emphasis id="B9429B68FFD7741FFE65D1626E4AF8FC" bold="true" box="[507,565,1887,1911]" pageId="2" pageNumber="137">
<taxonomicNameLabel id="A2712613FFD7741FFE65D1626E4AF8FC" box="[507,565,1887,1911]" pageId="2" pageNumber="137" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male. 1. Front view of head. 2. Right forewing. 3. Right hindwing. 4. Lacinial tip. 5. Hypandrium. 6. Phallosome. 7. Clunium, epiproct and right paraproct. Scales in mm.
</paragraph>
</caption>
<caption id="DF4917F2FFD6741EFF09D1A0694AF852" ID-DOI="http://doi.org/10.5281/zenodo.4745999" ID-Zenodo-Dep="4745999" httpUri="https://zenodo.org/record/4745999/files/figure.png" pageId="3" pageNumber="138" startId="3.[151,264,1949,1973]" targetBox="[209,1389,197,1925]" targetPageId="3">
<paragraph id="8B89477AFFD6741EFF09D1A0694AF852" blockId="3.[151,1437,1949,2010]" pageId="3" pageNumber="138">
<emphasis id="B9429B68FFD6741EFF09D1A06D39F83D" bold="true" box="[151,326,1949,1974]" pageId="3" pageNumber="138">FIGURES 814.</emphasis>
<taxonomicName id="4C363CF9FFD6741EFED5D1A36D83F83D" authority="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael, 2021" authorityName="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael" authorityYear="2021" box="[331,508,1950,1974]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="138" phylum="Arthropoda" rank="species" species="willineri" status="sp. nov.">
<emphasis id="B9429B68FFD6741EFED5D1A36D83F83D" box="[331,508,1950,1974]" italics="true" pageId="3" pageNumber="138">Loneura willineri</emphasis>
</taxonomicName>
<emphasis id="B9429B68FFD6741EFD9FD1A36E46F83D" bold="true" box="[513,569,1950,1974]" pageId="3" pageNumber="138">
<taxonomicNameLabel id="A2712613FFD6741EFD9FD1A36E46F83D" box="[513,569,1950,1974]" pageId="3" pageNumber="138" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Female. 8. Front view of head. 9. Right forewing. 10. Right hindwing. 11. Lacinial tip. 12. Subgenital plate. 13. Ninth sternum and gonapophyses. 14. Clunium, epiproct and right paraproct. Scales in mm.
</paragraph>
</caption>
<paragraph id="8B89477AFFD17419FF59D6AA6824FE7E" blockId="4.[151,1437,150,507]" pageId="4" pageNumber="139">
<emphasis id="B9429B68FFD17419FF59D6AA6E0FFF3B" bold="true" box="[199,624,150,177]" pageId="4" pageNumber="139">Fore &amp; hindwing venation variation</emphasis>
. Five female specimens from
<collectingCountry id="F32107EAFFD17419FC22D6AA6871FF3A" box="[956,1038,151,177]" name="Bolivia" pageId="4" pageNumber="139">Bolivia</collectingCountry>
(4) and
<collectingCountry id="F32107EAFFD17419FBF6D6AA68E4FF3A" box="[1128,1179,151,177]" name="Peru" pageId="4" pageNumber="139">Peru</collectingCountry>
(1) showed the following variations: (A) with no vein variation; female (B) left forewing M with seven primary branches with M
<subScript id="17B2453FFFD17419FAA3D6F0693AFF50" attach="left" box="[1341,1349,205,219]" fontSize="6" pageId="4" pageNumber="139">7</subScript>
forked, resulting in M
<subScript id="17B2453FFFD17419FEA9D6CC6D39FF74" attach="left" box="[311,326,241,255]" fontSize="6" pageId="4" pageNumber="139">7a</subScript>
and M
<subScript id="17B2453FFFD17419FE09D6CC6DD8FF74" attach="left" box="[407,423,241,255]" fontSize="6" pageId="4" pageNumber="139">7b</subScript>
, from which an incomplete vein derives (
<figureCitation id="130D5BFFFFD17419FCE1D6E36FB4FF72" box="[895,971,222,249]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig.15</figureCitation>
); female (C), left forewing M with seven primary branches with M
<subScript id="17B2453FFFD17419FE28D7286DC1FEA8" attach="left" box="[438,446,277,291]" fontSize="6" pageId="4" pageNumber="139">7</subScript>
three branched, resulting in M
<subScript id="17B2453FFFD17419FC82D7286F4BFEA8" attach="left" box="[796,820,277,291]" fontSize="6" pageId="4" pageNumber="139">7a1</subScript>
, M
<subScript id="17B2453FFFD17419FCC2D7286F0CFEA8" attach="left" box="[860,883,277,291]" fontSize="6" pageId="4" pageNumber="139">7a2</subScript>
and M
<subScript id="17B2453FFFD17419FC5AD7286FABFEA8" attach="left" box="[964,980,277,291]" fontSize="6" pageId="4" pageNumber="139">7b</subScript>
(
<figureCitation id="130D5BFFFFD17419FC7BD73E6846FE95" box="[997,1081,259,286]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 16</figureCitation>
); female (D), right forewing M with six primary branches, with M
<subScript id="17B2453FFFD17419FD89D7046E60FECC" attach="left" box="[535,543,313,327]" fontSize="6" pageId="4" pageNumber="139">2</subScript>
and M
<subScript id="17B2453FFFD17419FDF3D7046E0AFECC" attach="left" box="[621,629,313,327]" fontSize="6" pageId="4" pageNumber="139">6</subScript>
forked, resulting in M
<subScript id="17B2453FFFD17419FCEED7046F00FECC" attach="left" box="[880,895,313,327]" fontSize="6" pageId="4" pageNumber="139">2a</subScript>
and M
<subScript id="17B2453FFFD17419FC53D7046FA2FECC" attach="left" box="[973,989,313,327]" fontSize="6" pageId="4" pageNumber="139">2b</subScript>
, M
<subScript id="17B2453FFFD17419FB9DD704686DFECC" attach="left" box="[1027,1042,313,327]" fontSize="6" pageId="4" pageNumber="139">6a</subScript>
and M
<subScript id="17B2453FFFD17419FBFFD704680EFECC" attach="left" box="[1121,1137,313,327]" fontSize="6" pageId="4" pageNumber="139">6b</subScript>
from which an incomplete vein derives (
<figureCitation id="130D5BFFFFD17419FEB3D7766D01FEEE" box="[301,382,331,357]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 17</figureCitation>
), left hindwing M with three primary branches and then a M
<subScript id="17B2453FFFD17419FB85D760685CFEE0" attach="left" box="[1051,1059,349,363]" fontSize="6" pageId="4" pageNumber="139">4</subScript>
bifurcation originating directly in the M vein, resulting in M
<subScript id="17B2453FFFD17419FE23D7BC6DB2FE04" attach="left" box="[445,461,385,399]" fontSize="6" pageId="4" pageNumber="139">4a</subScript>
and M
<subScript id="17B2453FFFD17419FD82D7BC6E53FE04" attach="left" box="[540,556,385,399]" fontSize="6" pageId="4" pageNumber="139">4b</subScript>
(
<figureCitation id="130D5BFFFFD17419FDA2D7526EF0FE02" box="[572,655,367,393]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 18</figureCitation>
); female (E), right forewing M with seven primary branches, with M
<subScript id="17B2453FFFD17419FA0AD7BC69E3FE04" attach="left" box="[1428,1436,385,399]" fontSize="6" pageId="4" pageNumber="139">6</subScript>
and M
<subScript id="17B2453FFFD17419FF41D7986C98FE38" attach="left" box="[223,231,421,435]" fontSize="6" pageId="4" pageNumber="139">7</subScript>
forked, resulting in M
<subScript id="17B2453FFFD17419FE7CD7986D8EFE38" attach="left" box="[482,497,421,435]" fontSize="6" pageId="4" pageNumber="139">6a</subScript>
and M
<subScript id="17B2453FFFD17419FDDED7986E30FE38" attach="left" box="[576,591,421,435]" fontSize="6" pageId="4" pageNumber="139">6b</subScript>
, M
<subScript id="17B2453FFFD17419FDE8D7986EFAFE38" attach="left" box="[630,645,421,435]" fontSize="6" pageId="4" pageNumber="139">7a</subScript>
and M
<subScript id="17B2453FFFD17419FD4AD7986E9BFE38" attach="left" box="[724,740,421,435]" fontSize="6" pageId="4" pageNumber="139">7b</subScript>
and an incomplete vein drifting from the areola postica (Cu
<subScript id="17B2453FFFD17419FAE1D79869ECFE38" attach="left" box="[1407,1427,421,435]" fontSize="6" pageId="4" pageNumber="139">A1</subScript>
) (
<figureCitation id="130D5BFFFFD17419FF3ED78A6C8DFE5A" box="[160,242,439,465]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 19</figureCitation>
), left forewing M with seven primary branches, with M
<subScript id="17B2453FFFD17419FCC0D7F46F19FE5C" attach="left" box="[862,870,457,471]" fontSize="6" pageId="4" pageNumber="139">7</subScript>
three branched arc shaped in equal parts, resulting in M
<subScript id="17B2453FFFD17419FF53D7D06CA3FE70" attach="left" box="[205,220,493,507]" fontSize="6" pageId="4" pageNumber="139">7a</subScript>
, M
<subScript id="17B2453FFFD17419FE9DD7D06D6CFE70" attach="left" box="[259,275,493,507]" fontSize="6" pageId="4" pageNumber="139">7b</subScript>
and M
<subScript id="17B2453FFFD17419FEFDD7D06D0DFE70" attach="left" box="[355,370,493,507]" fontSize="6" pageId="4" pageNumber="139">7c</subScript>
, (
<figureCitation id="130D5BFFFFD17419FE17D7E66DA3FE7E" box="[393,476,475,501]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 20</figureCitation>
), right hindwing M with five primary branches (
<figureCitation id="130D5BFFFFD17419FC65D7E66831FE7E" box="[1019,1102,475,501]" captionStart="FIGURES 1521" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[171,1419,540,1837]" captionTargetId="figure-247@4.[151,1436,525,1837]" captionTargetPageId="4" captionText="FIGURES 1521. Variation in venation of Loneura willineri n. sp. specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4917F2FFD17419FF09D1796F12F82F" ID-DOI="http://doi.org/10.5281/zenodo.4746003" ID-Zenodo-Dep="4746003" httpUri="https://zenodo.org/record/4746003/files/figure.png" pageId="4" pageNumber="139" startId="4.[151,264,1860,1884]" targetBox="[171,1419,540,1837]" targetPageId="4">
<paragraph id="8B89477AFFD17419FF09D1796F12F82F" blockId="4.[151,1437,1860,1957]" pageId="4" pageNumber="139">
<emphasis id="B9429B68FFD17419FF09D1796D2BF8D7" bold="true" box="[151,340,1860,1884]" pageId="4" pageNumber="139">FIGURES 1521.</emphasis>
Variation in venation of
<taxonomicName id="4C363CF9FFD17419FDD3D1786E80F8D6" authority="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael, 2021" authorityName="Cutrim &amp; Neto &amp; García Aldrete &amp; Rafael" authorityYear="2021" box="[589,767,1861,1885]" class="Insecta" family="Ptiloneuridae" genus="Loneura" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="139" phylum="Arthropoda" rank="species" species="willineri" status="sp. nov.">
<emphasis id="B9429B68FFD17419FDD3D1786E80F8D6" box="[589,767,1861,1885]" italics="true" pageId="4" pageNumber="139">Loneura willineri</emphasis>
</taxonomicName>
<emphasis id="B9429B68FFD17419FC9AD1786F42F8D6" bold="true" box="[772,829,1861,1885]" pageId="4" pageNumber="139">
<taxonomicNameLabel id="A2712613FFD17419FC9AD1786F42F8D6" box="[772,829,1861,1885]" pageId="4" pageNumber="139" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
specimens. 15. Left forewing of female (B). 16. Left forewing of female (C). 17. Right forewing of female (D). 18. Left hindwing of female (D). 19. Right forewing of female (E). 20. Left forewing of female (E). 21. Left hindwing of female (E). Scales in mm.
</paragraph>
</caption>
<subSubSection id="C32C14F1FFD07418FF59D6AB683BFE03" pageId="5" pageNumber="140" type="materials_examined">
<paragraph id="8B89477AFFD07418FF59D6AB683BFE03" blockId="5.[151,1436,150,393]" pageId="5" pageNumber="140">
<emphasis id="B9429B68FFD07418FF59D6AB6DCEFF3B" bold="true" box="[199,433,150,176]" pageId="5" pageNumber="140">Material examined.</emphasis>
<materialsCitation id="3B5E4D27FFD07418FE26D6AB6EF4FF5E" ID-GBIF-Occurrence="3118913302" collectingDate="2005-04-24" collectionCode="UNAM" collectorName="On &amp; A. N. Garcia-Aldrete" country="Bolivia" location="Samaipata" pageId="5" pageNumber="140" specimenCount="1" stateProvince="Santa Cruz" typeStatus="holotype">
<typeStatus id="548DF9D8FFD07418FE26D6AB6E5FFF3B" box="[440,544,150,176]" pageId="5" pageNumber="140" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED27DFBFFFD07418FDF3D6AB6EB6FF3B" box="[621,713,150,176]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="5" pageNumber="140">UNAM</collectionCode>
).
<collectingCountry id="F32107EAFFD07418FD45D6AB6F4FFF3B" box="[731,816,150,176]" name="Bolivia" pageId="5" pageNumber="140">Bolivia</collectingCountry>
,
<collectingRegion id="49F28998FFD07418FCA2D6AB6FC4FF3B" box="[828,955,150,176]" country="Bolivia" name="Santa Cruz" pageId="5" pageNumber="140">Santa Cruz</collectingRegion>
,
<quantity id="4CCEEA9FFFD07418FC58D6AB686AFF3B" box="[966,1045,150,177]" metricMagnitude="3" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="140" unit="km" value="5.8">5.8 km</quantity>
E
<location id="8EE911A1FFD07418FBAAD6AB68D3FF3B" LSID="urn:lsid:plazi:treatment:039FF66CFFD47418FF09D71A683BFE03:8EE911A1FFD07418FBAAD6AB68D3FF3B" box="[1076,1196,150,176]" country="Bolivia" name="Samaipata" pageId="5" pageNumber="140" stateProvince="Santa Cruz">Samaipata</location>
,
<date id="FF8861BAFFD07418FB29D6AB6948FF3A" box="[1207,1335,150,177]" pageId="5" pageNumber="140" value="2005-04-24">
<collectingDate id="EFCC9852FFD07418FB29D6AB6948FF3A" box="[1207,1335,150,177]" pageId="5" pageNumber="140" value="2005-04-24">24.IV.2005</collectingDate>
</date>
.
<collectorName id="26C322ACFFD07418FADFD6AB691CFF3B" box="[1345,1379,150,176]" pageId="5" pageNumber="140">On</collectorName>
rock faces at creek side. (
<collectorName id="26C322ACFFD07418FE14D6876EFAFF5E" box="[394,645,186,213]" pageId="5" pageNumber="140">A. N. García-Aldrete</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B5E4D27FFD07418FD04D6866D16FF73" ID-GBIF-Occurrence="3118913305" collectingDate="1980-12-15" collectingDateMax="2005-05-04" collectingDateMin="1980-12-15" collectionCode="UNAM" collectorName="A. N. Garcia Aldrete" country="Bolivia" elevation="1396" latitude="-16.254116" location="Machu Pichu" longLatPrecision="1" longitude="-67.77942" municipality="On" pageId="5" pageNumber="140" specimenCode="ELM5007" specimenCount="4" specimenCount-female="4" stateProvince="Cusco" typeStatus="paratype">
<typeStatus id="548DF9D8FFD07418FD04D6866E83FF5E" box="[666,764,187,213]" pageId="5" pageNumber="140" type="paratype">Paratype</typeStatus>
female,
<emphasis id="B9429B68FFD07418FCFBD6876FA6FF5F" bold="true" box="[869,985,186,212]" pageId="5" pageNumber="140">
<collectorName id="26C322ACFFD07418FCFBD6876FA6FF5F" box="[869,985,186,212]" pageId="5" pageNumber="140">Female A</collectorName>
</emphasis>
, same data as the holotype.
<specimenCount id="9D308CF3FFD07418FAB2D68769E3FF5F" box="[1324,1436,186,212]" pageId="5" pageNumber="140" type="female">3 females</specimenCount>
(
<collectionCode id="ED27DFBFFFD07418FF01D6E36C84FF73" box="[159,251,222,248]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="5" pageNumber="140">UNAM</collectionCode>
).
<collectingCountry id="F32107EAFFD07418FE90D6E36D1AFF73" box="[270,357,222,248]" name="Bolivia" pageId="5" pageNumber="140">Bolivia</collectingCountry>
:
</materialsCitation>
<materialsCitation id="3B5E4D27FFD07418FEEED6E36E0EFE97" ID-GBIF-Occurrence="3118913304" collectingDate="1980-12-15" collectingDateMax="2005-05-04" collectingDateMin="1980-12-15" collectionCode="UNAM" collectorName="A. N. Garcia Aldrete" country="Bolivia" elevation="1396" latitude="-16.254116" location="Machu Pichu" longLatPrecision="1" longitude="-67.77942" municipality="On" pageId="5" pageNumber="140" specimenCode="ELM5007" specimenCount="4" specimenCount-female="4" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="B9429B68FFD07418FEEED6E36D9DFF73" bold="true" box="[368,482,222,248]" pageId="5" pageNumber="140">
<collectorName id="26C322ACFFD07418FEEED6E36D9DFF73" box="[368,482,222,248]" pageId="5" pageNumber="140">Female B</collectorName>
</emphasis>
:
<collectorName id="26C322ACFFD07418FE6FD6E26E3EFF72" box="[497,577,223,249]" pageId="5" pageNumber="140">La Paz</collectorName>
,
<collectorName id="26C322ACFFD07418FDD0D6E26EAAFF72" box="[590,725,223,249]" pageId="5" pageNumber="140">Nor Yungas</collectorName>
,
<locationDeviation id="9A3B5E58FFD07418FD7FD6E36FC0FF73" box="[737,959,222,248]" location="Coroico" pageId="5" pageNumber="140">
<quantity id="4CCEEA9FFFD07418FD7FD6E36F56FF73" box="[737,809,222,248]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="140" unit="km" value="12.0">12 km</quantity>
SW Coroico
</locationDeviation>
,
<date id="FF8861BAFFD07418FC55D6E36844FF72" box="[971,1083,222,249]" pageId="5" pageNumber="140" value="2004-04-05">
<collectingDate id="EFCC9852FFD07418FC55D6E36844FF72" box="[971,1083,222,249]" pageId="5" pageNumber="140" value="2004-04-05">5.IV.2004</collectingDate>
</date>
, 1694m,
<geoCoordinate id="EE0221BDFFD07418FB3AD6E3694EFF73" box="[1188,1329,222,248]" degrees="16" direction="south" minutes="14.082" orientation="latitude" pageId="5" pageNumber="140" precision="1" value="-16.2347">16º14.082S</geoCoordinate>
:
<geoCoordinate id="EE0221BDFFD07418FAAAD6E36CAEFE97" pageId="5" pageNumber="140">
67º4087 7
<collectorName id="26C322ACFFD07418FF2FD73F6CAEFE97" box="[177,209,258,284]" pageId="5" pageNumber="140">W</collectorName>
</geoCoordinate>
. On rock wall.
<collectorName id="26C322ACFFD07418FEE0D73F6E12FE97" box="[382,621,258,285]" pageId="5" pageNumber="140">A. N. García Aldrete</collectorName>
;
</materialsCitation>
<materialsCitation id="3B5E4D27FFD07418FDE7D73F683AFECB" ID-GBIF-Occurrence="3118913301" collectingDate="1980-12-15" collectingDateMax="2005-05-04" collectingDateMin="1980-12-15" collectionCode="UNAM" collectorName="A. N. Garcia Aldrete" country="Bolivia" elevation="1396" latitude="-16.254116" location="Machu Pichu" longLatPrecision="1" longitude="-67.77942" municipality="On" pageId="5" pageNumber="140" specimenCode="ELM5007" specimenCount="4" specimenCount-female="4" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="B9429B68FFD07418FDE7D73F6E9BFE97" bold="true" box="[633,740,258,284]" pageId="5" pageNumber="140">female C</emphasis>
:
<collectorName id="26C322ACFFD07418FD6AD73E6F3BFE96" box="[756,836,259,285]" pageId="5" pageNumber="140">La Paz</collectorName>
,
<collectorName id="26C322ACFFD07418FCCFD73E6FA7FE96" box="[849,984,259,285]" pageId="5" pageNumber="140">Nor Yungas</collectorName>
,
<locationDeviation id="9A3B5E58FFD07418FC7BD73F68D5FE97" box="[997,1194,258,285]" location="Yolosa" pageId="5" pageNumber="140">
<quantity id="4CCEEA9FFFD07418FC7BD73F6860FE97" box="[997,1055,258,284]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="140" unit="km" value="1.0">1 km</quantity>
NW Yolosa
</locationDeviation>
, towards
<collectorName id="26C322ACFFD07418FA89D73F69E7FE97" box="[1303,1432,258,284]" pageId="5" pageNumber="140">Chuspipata</collectorName>
,
<date id="FF8861BAFFD07418FF09D71B6C81FECA" box="[151,254,294,321]" pageId="5" pageNumber="140" value="2005-05-04">
<collectingDate id="EFCC9852FFD07418FF09D71B6C81FECA" box="[151,254,294,321]" pageId="5" pageNumber="140" value="2005-05-04">4.V.2005</collectingDate>
</date>
,
<geoCoordinate id="EE0221BDFFD07418FE94D71B6DEFFECA" box="[266,400,294,321]" degrees="16" direction="south" minutes="15.247" orientation="latitude" pageId="5" pageNumber="140" precision="1" value="-16.254116">16º15.247S</geoCoordinate>
:
<geoCoordinate id="EE0221BDFFD07418FE0DD71B6E57FECA" box="[403,552,294,321]" degrees="67" direction="west" minutes="46.765" orientation="longitude" pageId="5" pageNumber="140" precision="1" value="-67.77942">67º46.765W</geoCoordinate>
,
<quantity id="4CCEEA9FFFD07418FDADD71B6EF5FECA" box="[563,650,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.396" pageId="5" pageNumber="140" unit="m" value="1396.0">
<elevation id="001BA049FFD07418FDADD71B6EF5FECA" box="[563,650,294,321]" metricMagnitude="3" metricUnit="m" metricValue="1.396" pageId="5" pageNumber="140" unit="m" value="1396.0">1396 m</elevation>
</quantity>
, on rock in slope,
<collectorName id="26C322ACFFD07418FCCBD71B683DFECB" box="[853,1090,294,321]" pageId="5" pageNumber="140">A. N. García Aldrete</collectorName>
;
</materialsCitation>
<materialsCitation id="3B5E4D27FFD07418FBD2D71B68A5FEEF" ID-GBIF-Occurrence="3118913306" collectingDate="1980-12-15" collectingDateMax="2005-05-04" collectingDateMin="1980-12-15" collectionCode="UNAM" collectorName="A. N. Garcia Aldrete" country="Bolivia" elevation="1396" latitude="-16.254116" location="Machu Pichu" longLatPrecision="1" longitude="-67.77942" municipality="On" pageId="5" pageNumber="140" specimenCode="ELM5007" specimenCount="4" specimenCount-female="4" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="B9429B68FFD07418FBD2D71B68C9FECB" bold="true" box="[1100,1206,294,320]" pageId="5" pageNumber="140">female D</emphasis>
: Santa Cruz,
<locationDeviation id="9A3B5E58FFD07418FAD1D71B6D57FEEF" location="Samaipata" pageId="5" pageNumber="140">
<quantity id="4CCEEA9FFFD07418FAD1D71B69E3FECB" box="[1359,1436,294,321]" metricMagnitude="3" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="140" unit="km" value="5.8">5.8 km</quantity>
E Samaipata
</locationDeviation>
,
<date id="FF8861BAFFD07418FEADD7776DCCFEEE" box="[307,435,330,357]" pageId="5" pageNumber="140" value="2005-04-24">
<collectingDate id="EFCC9852FFD07418FEADD7776DCCFEEE" box="[307,435,330,357]" pageId="5" pageNumber="140" value="2005-04-24">24.IV.2005</collectingDate>
</date>
.
<collectingMunicipality id="6BEDDD00FFD07418FE20D7776D9FFEEF" box="[446,480,330,356]" pageId="5" pageNumber="140">On</collectingMunicipality>
rock faces, at creek side.
<collectorName id="26C322ACFFD07418FC9AD7776F85FEEF" box="[772,1018,330,357]" pageId="5" pageNumber="140">A. N. García-Aldrete.</collectorName>
<specimenCount id="9D308CF3FFD07418FB9ED777681DFEEF" box="[1024,1122,330,356]" pageId="5" pageNumber="140" type="female">1 female</specimenCount>
(
<collectionCode id="ED27DFBFFFD07418FBECD77768B1FEEF" box="[1138,1230,330,356]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="5" pageNumber="140">UNAM</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B5E4D27FFD07418FB7FD777683FFE03" ID-GBIF-Occurrence="3118913303" collectingDate="1980-12-15" collectingDateMax="2005-05-04" collectingDateMin="1980-12-15" collectionCode="UNAM" collectorName=" E. L. Mockford" country="Bolivia" elevation="1396" latitude="-16.254116" location="Machu Pichu" longLatPrecision="1" longitude="-67.77942" municipality="On" pageId="5" pageNumber="140" specimenCode="ELM5007" specimenCount="4" specimenCount-female="4" stateProvince="Cusco" typeStatus="paratype">
<emphasis id="B9429B68FFD07418FB7FD777692CFEEF" bold="true" box="[1249,1363,330,356]" pageId="5" pageNumber="140">
<collectorName id="26C322ACFFD07418FB7FD777692CFEEF" box="[1249,1363,330,356]" pageId="5" pageNumber="140">Female E</collectorName>
</emphasis>
:
<collectingCountry id="F32107EAFFD07418FAFCD77669E7FEEE" box="[1378,1432,331,357]" name="Peru" pageId="5" pageNumber="140">Peru</collectingCountry>
,
<specimenCode id="DB90EF01FFD07418FF09D7536D72FE02" box="[151,269,366,393]" collectionCode="ELM" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="5" pageNumber="140">ELM5007</specimenCode>
,
<collectingRegion id="49F28998FFD07418FE86D7536D1EFE03" box="[280,353,366,392]" country="Peru" name="Cusco" pageId="5" pageNumber="140">Cusco</collectingRegion>
,
<location id="8EE911A1FFD07418FEF2D7536E7BFE03" LSID="urn:lsid:plazi:treatment:039FF66CFFD47418FF09D71A683BFE03:8EE911A1FFD07418FEF2D7536E7BFE03" box="[364,516,366,392]" country="Bolivia" latitude="-16.254116" longLatPrecision="1" longitude="-67.77942" municipality="On" name="Machu Pichu" pageId="5" pageNumber="140" stateProvince="Cusco">Machu Pichu</location>
,
<date id="FF8861BAFFD07418FD91D7536EE5FE02" box="[527,666,366,393]" pageId="5" pageNumber="140" value="1980-12-15">
<collectingDate id="EFCC9852FFD07418FD91D7536EE5FE02" box="[527,666,366,393]" pageId="5" pageNumber="140" value="1980-12-15">15.XII.1980</collectingDate>
</date>
, in recesses on ruins.
<collectorName id="26C322ACFFD07418FC10D752683FFE03" box="[910,1088,366,393]" pageId="5" pageNumber="140">E. L. Mockford</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>