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<document id="E7B183AD2ABA8DE359C0C08DD2A0857B" ID-DOI="10.5281/zenodo.158101" ID-GBIF-Dataset="1010f4fc-4ae7-49bf-a6d5-3582c56d1bec" ID-ISSN="1175­5326" ID-Zenodo-Dep="158101" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461038695694" checkinUser="plazi" docAuthor="Pogue, Michael G." docDate="2004" docId="03920A30FF8CFFCC8806FE86FC7B833E" docLanguage="en" docName="zt00788.pdf" docOrigin="Zootaxa 788" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Schinia immaculata Pogue, new species" docType="treatment" docVersion="8" lastPageNumber="4" masterDocId="FFAB7248FF8DFFCF890EFF9AFFDB8148" masterDocTitle="A new species of Schinia Hübner from riparian habitats in the Grand Canyon (Lepidoptera: Noctuidae: Heliothinae)" masterLastPageNumber="4" masterPageNumber="1" pageNumber="2" updateTime="1698575063737" updateUser="plazi">
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<mods:title id="CF6EBA9B2FFF75C7A7D60CE9729CC242">A new species of Schinia Hübner from riparian habitats in the Grand Canyon (Lepidoptera: Noctuidae: Heliothinae)</mods:title>
</mods:titleInfo>
<mods:name id="4D021816CD1AE3DF2A41F46FB66FDEE7" type="personal">
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<mods:namePart id="969AE047B55B71F250A166CB25CEAC9E">Pogue, Michael G.</mods:namePart>
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<mods:typeOfResource id="DC70E7E633DBC9CFA378FBA55722155F">text</mods:typeOfResource>
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<mods:date id="F50BAEFFAC374A5ED670113900BFFF9B">2004</mods:date>
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<treatment id="03920A30FF8CFFCC8806FE86FC7B833E" ID-DOI="http://doi.org/10.5281/zenodo.6271985" ID-GBIF-Taxon="119609422" ID-Zenodo-Dep="6271985" LSID="urn:lsid:plazi:treatment:03920A30FF8CFFCC8806FE86FC7B833E" httpUri="http://treatment.plazi.org/id/03920A30FF8CFFCC8806FE86FC7B833E" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
<subSubSection id="C321E8ADFF8CFFCE8806FE86FE7D8011" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8B84BB26FF8CFFCE8806FE86FD36807D" blockId="1.[264,749,283,345]" box="[264,749,283,310]" pageId="1" pageNumber="2">
<heading id="D0CC0C4AFF8CFFCE8806FE86FD36807D" bold="true" box="[264,749,283,310]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="B94F6734FF8CFFCE8806FE86FD36807D" bold="true" box="[264,749,283,310]" pageId="1" pageNumber="2">
<taxonomicName id="4C3BC0A5FF8CFFCE8806FE86FD8A807D" ID-CoL="948S6" authority="Pogue" authorityName="Pogue" box="[264,593,283,310]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="immaculata" status="sp. nov.">
<emphasis id="B94F6734FF8CFFCE8806FE86FE20807E" bold="true" box="[264,507,284,310]" italics="true" pageId="1" pageNumber="2">Schinia immaculata</emphasis>
Pogue
</taxonomicName>
,
<taxonomicNameLabel id="A27CDA4FFF8CFFCE8B50FE81FD36807D" box="[606,749,283,309]" pageId="1" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B84BB26FF8CFFCE8806FEA5FE7D8011" blockId="1.[264,749,283,345]" box="[264,422,319,345]" pageId="1" pageNumber="2">
(
<figureCitation id="1300A7A3FF8CFFCE881FFEA5FE468011" box="[273,413,319,345]" captionStart="FIGURES 1 6" captionStartId="2.[264,382,1413,1437]" captionTargetBox="[300,1299,285,1389]" captionTargetId="figure@2.[270,1317,280,1409]" captionTargetPageId="2" captionText="FIGURES 1 6. Adults and male genitalia. 1, Schinia immaculata, male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2, S biundulata, female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3, S. immaculata, male genitalia; 4, S. immaculata, aedoeagus; 5, S. biundulata, male genitalia; 6, S. biundulata, aedoeagus." httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="1" pageNumber="2">Figs. 1, 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8CFFCE8806FEE3FB568310" pageId="1" pageNumber="2" type="diagnosis">
<paragraph id="8B84BB26FF8CFFCE8806FEE3FB568310" blockId="1.[264,1325,377,1903]" pageId="1" pageNumber="2">
<emphasis id="B94F6734FF8CFFCE8806FEE3FE5380DB" bold="true" box="[264,392,377,403]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
<taxonomicName id="4C3BC0A5FF8CFFCE889CFEE3FD5B80DA" box="[402,640,377,402]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8CFFCE889CFEE3FD5B80DA" box="[402,640,377,402]" italics="true" pageId="1" pageNumber="2">Schinia immaculata</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C3BC0A5FF8CFFCE8A07FEE3FC2380DB" authority="Smith" authorityName="Smith" box="[777,1016,377,403]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B94F6734FF8CFFCE8A07FEE3FC7380DA" box="[777,936,377,402]" italics="true" pageId="1" pageNumber="2">S. biundulata</emphasis>
Smith
</taxonomicName>
in size, forewing ground color, and dark hindwing, but
<taxonomicName id="4C3BC0A5FF8CFFCE8B7BFE3BFCCF80F2" box="[629,788,417,442]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B94F6734FF8CFFCE8B7BFE3BFCCF80F2" box="[629,788,417,442]" italics="true" pageId="1" pageNumber="2">S. biundulata</emphasis>
</taxonomicName>
has well defined antemedial and postmedial lines, which are absent in
<taxonomicName id="4C3BC0A5FF8CFFCE8B34FE53FD3A80AA" box="[570,737,457,482]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8CFFCE8B34FE53FD3A80AA" box="[570,737,457,482]" italics="true" pageId="1" pageNumber="2">S. immaculata</emphasis>
</taxonomicName>
. In male genitalia the shape of the valve is different, with
<taxonomicName id="4C3BC0A5FF8CFFCE8876FE6AFDF98341" box="[376,546,496,521]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8CFFCE8876FE6AFDF98341" box="[376,546,496,521]" italics="true" pageId="1" pageNumber="2">S. immaculata</emphasis>
</taxonomicName>
having a round apex (
<figureCitation id="1300A7A3FF8CFFCE8A22FE6AFCA88342" box="[812,883,496,522]" captionStart="FIGURES 1 6" captionStartId="2.[264,382,1413,1437]" captionTargetBox="[300,1299,285,1389]" captionTargetId="figure@2.[270,1317,280,1409]" captionTargetPageId="2" captionText="FIGURES 1 6. Adults and male genitalia. 1, Schinia immaculata, male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2, S biundulata, female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3, S. immaculata, male genitalia; 4, S. immaculata, aedoeagus; 5, S. biundulata, male genitalia; 6, S. biundulata, aedoeagus." httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
) versus a more pointed apex (
<figureCitation id="1300A7A3FF8CFFCE8DD5FE6AFAF98342" box="[1243,1314,496,522]" captionStart="FIGURES 1 6" captionStartId="2.[264,382,1413,1437]" captionTargetBox="[300,1299,285,1389]" captionTargetId="figure@2.[270,1317,280,1409]" captionTargetPageId="2" captionText="FIGURES 1 6. Adults and male genitalia. 1, Schinia immaculata, male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2, S biundulata, female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3, S. immaculata, male genitalia; 4, S. immaculata, aedoeagus; 5, S. biundulata, male genitalia; 6, S. biundulata, aedoeagus." httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="1" pageNumber="2">Fig. 5</figureCitation>
) in
<taxonomicName id="4C3BC0A5FF8CFFCE8827FD8DFE128378" box="[297,457,535,560]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B94F6734FF8CFFCE8827FD8DFE128378" box="[297,457,535,560]" italics="true" pageId="1" pageNumber="2">S. biundulata</emphasis>
</taxonomicName>
; and the uncus is relatively longer, approximately 0.38 X length of the valve in
<taxonomicName id="4C3BC0A5FF8CFFCE8863FDA5FDCF8310" box="[365,532,575,600]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8CFFCE8863FDA5FDCF8310" box="[365,532,575,600]" italics="true" pageId="1" pageNumber="2">S. immaculata</emphasis>
</taxonomicName>
, compared to 0.32 X length of valve in
<emphasis id="B94F6734FF8CFFCE8AE7FDA5FB568310" box="[1001,1165,575,600]" italics="true" pageId="1" pageNumber="2">
<taxonomicName id="4C3BC0A5FF8CFFCE8AE7FDA5FB528310" box="[1001,1161,575,600]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="biundulata">S. biundulata</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8CFFCE8836FDFCFDC785B0" pageId="1" pageNumber="2" type="description">
<paragraph id="8B84BB26FF8CFFCE8836FDFCFADD8598" blockId="1.[264,1325,377,1903]" pageId="1" pageNumber="2">
<emphasis id="B94F6734FF8CFFCE8836FDFCFE1483C8" bold="true" box="[312,463,614,640]" pageId="1" pageNumber="2">Description.</emphasis>
<emphasis id="B94F6734FF8CFFCE88D2FDFCFDFE8337" box="[476,549,614,639]" italics="true" pageId="1" pageNumber="2">Head:</emphasis>
Frons and vertex cream; labial palpus long, extending beyond frons, cream; eyes large and globular.
<emphasis id="B94F6734FF8CFFCE8BEFFD14FC9B83EF" box="[737,832,654,679]" italics="true" pageId="1" pageNumber="2">Thorax:</emphasis>
Scales hair­like overlaying wide scales, cream; fore femur cream with long, white, ventral fringe of hair­like scales; tibia white, largest spine on inner side elongate, round, 1 shorter slender spine dorsal to it, outer side with 23 slender spines that become progressively shorter proximally; tarsi white; middle and hind legs white; underside white.
<emphasis id="B94F6734FF8CFFCE8BC3FCB6FC93820D" box="[717,840,812,837]" italics="true" pageId="1" pageNumber="2">Forewing:</emphasis>
Length 10.5311.70, mean 11.21 +
<quantity id="4CC316C3FF8CFFCE8DF9FCB1FEED8225" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="2" unit="mm" value="1.21">1.21 mm</quantity>
(n=4). Ground color cream with a faint tinge of rufous on the scales; pattern absent; fringe white.
<emphasis id="B94F6734FF8CFFCE88A1FCE1FDF782DC" box="[431,556,891,916]" italics="true" pageId="1" pageNumber="2">Hindwing:</emphasis>
Ground color gray; pattern absent; base of fringe cream, rest white.
<emphasis id="B94F6734FF8CFFCE8852FC38FE0E82F3" box="[348,469,930,955]" italics="true" pageId="1" pageNumber="2">Abdomen:</emphasis>
Dorsum white, distal border of sternites cream, ventrum white; scent pockets and hair pencils on second sternite absent.
<emphasis id="B94F6734FF8CFFCE8A6AFC53FC0882AA" box="[868,979,969,994]" italics="true" pageId="1" pageNumber="2">Genitalia</emphasis>
(
<figureCitation id="1300A7A3FF8CFFCE8AEAFC53FB8382AB" box="[996,1112,969,995]" captionStart="FIGURES 1 6" captionStartId="2.[264,382,1413,1437]" captionTargetBox="[300,1299,285,1389]" captionTargetId="figure@2.[270,1317,280,1409]" captionTargetPageId="2" captionText="FIGURES 1 6. Adults and male genitalia. 1, Schinia immaculata, male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2, S biundulata, female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3, S. immaculata, male genitalia; 4, S. immaculata, aedoeagus; 5, S. biundulata, male genitalia; 6, S. biundulata, aedoeagus." httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="1" pageNumber="2">Figs. 34</figureCitation>
): Uncus elongate, approximately 0.38 X length of valve, apex blunt with small hook; valve moderate (length approximately 7.1 X width), costal margin slightly bent at 2/3 length, posterior margin gently curved, apex round; ampulla small (0.04 X length of valve); corona with stout spines; sacculus with ventral margin produced; juxta rectangular, width greater than height, dorsal margin broadly V­shaped; saccus V­shaped; aedoeagus slightly curved; vesica emerging ventrally, basal diverticulum present, 3 1/2 coils; cornuti finely scobinate.
</paragraph>
<paragraph id="8B84BB26FF8CFFCE8836FB44FDC785B0" blockId="1.[264,1325,377,1903]" box="[312,540,1246,1272]" pageId="1" pageNumber="2">
<emphasis id="B94F6734FF8CFFCE8836FB44FE4185BF" box="[312,410,1246,1271]" italics="true" pageId="1" pageNumber="2">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8CFFCE8836FA9FFCC884F5" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8B84BB26FF8CFFCE8836FA9FFCC884F5" blockId="1.[264,1325,377,1903]" pageId="1" pageNumber="2">
<emphasis id="B94F6734FF8CFFCE8836FA9FFE368457" bold="true" box="[312,493,1285,1311]" pageId="1" pageNumber="2">
<typeStatus id="54800584FF8CFFCE8836FA9FFEAE8457" box="[312,373,1285,1311]" pageId="1" pageNumber="2">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54800584FF8CFFCE88FAFA9FFD488457" box="[500,659,1285,1311]" pageId="1" pageNumber="2" type="holotype">HOLOTYPE</typeStatus>
: ɗ, Arizona, Coconino Co., Colorado River, Grand Canyon, river mile 23.0 L, shore,
<geoCoordinate id="EE0FDDE1FF8CFFCE8B68FAB6FD2B840E" box="[614,752,1324,1350]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="5" value="36.6003">36.6003° N</geoCoordinate>
,
<geoCoordinate id="EE0FDDE1FF8CFFCE8BF2FAB6FC4C840E" box="[764,919,1324,1350]" direction="west" orientation="longitude" pageId="1" pageNumber="2" precision="5" value="-111.7668">111.7668° W</geoCoordinate>
,
<date id="FF859DE6FF8CFFCE8AADFAB6FBE1840E" box="[931,1082,1324,1350]" pageId="1" pageNumber="2" value="2003-05-02">May 2, 2003</date>
, R. J. Delph, genitalia slide
<collectionCode id="ED2A23E3FF8CFFCE8865FACEFE1B8426" LSID="urn:lsid:biocol.org:col:34871" box="[363,448,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2">USNM</collectionCode>
47867,
<collectionCode id="ED2A23E3FF8CFFCE8B11FACEFDAF8426" LSID="urn:lsid:biocol.org:col:34871" box="[543,628,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2">USNM</collectionCode>
<collectionCode id="ED2A23E3FF8CFFCE8B75FACEFD6F8426" LSID="urn:lsid:biocol.org:col:13589" box="[635,692,1364,1390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="1" pageNumber="2">ENT</collectionCode>
0 0 219964. The
<typeStatus id="54800584FF8CFFCE8A7AFACEFC018426" box="[884,986,1364,1390]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
is provisionally deposited at the
<collectionCode id="ED2A23E3FF8CFFCE883AFAE1FE5784DD" LSID="urn:lsid:biocol.org:col:34871" box="[308,396,1403,1429]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="2">USNM</collectionCode>
, Washington, DC, pending mutual resolution and agreement with the National Park Service regarding specimen deposition.
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8CFFCE8836FA50FDEF8627" pageId="1" pageNumber="2" type="distribution">
<paragraph id="8B84BB26FF8CFFCE8836FA50FDEF8627" blockId="1.[264,1325,377,1903]" pageId="1" pageNumber="2">
<emphasis id="B94F6734FF8CFFCE8836FA50FDA384AC" bold="true" box="[312,632,1482,1508]" pageId="1" pageNumber="2">Distribution and Habitat.</emphasis>
Specimens were taken from 3 riparian zones of the Colorado River in the Grand Canyon in Coconino Co., Arizona: shoreline, old high water, and new high water. The shoreline zone is situated one meter above the existing daily highwater shoreline, which varies over time depending on water releases from Glen Canyon Dam. The new high water zone is situated just above shoreline and was characterized by vegetation dominated by tamarisk (
<taxonomicName id="4C3BC0A5FF8CFFCE8BA1F90AFCD687E1" box="[687,781,1680,1705]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Violales" pageId="1" pageNumber="2" phylum="Magnoliophyta" rank="genus">
<emphasis id="B94F6734FF8CFFCE8BA1F90AFCD687E1" box="[687,781,1680,1705]" italics="true" pageId="1" pageNumber="2">Tamarix</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C3BC0A5FF8CFFCE8A45F90AFC2987E2" box="[843,1010,1680,1706]" class="Magnoliopsida" family="Tamaricaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Violales" pageId="1" pageNumber="2" phylum="Magnoliophyta" rank="family">Tamaricaceae</taxonomicName>
). The old high water zone was the highest elevation hydrologic zone and was characterized by mesquite (
<taxonomicName id="4C3BC0A5FF8CFFCE8DCDF922FAF18799" box="[1219,1322,1720,1745]" class="Magnoliopsida" family="Fabaceae" genus="Prosopis" kingdom="Animalia" order="Fabales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis id="B94F6734FF8CFFCE8DCDF922FAF18799" box="[1219,1322,1720,1745]" italics="true" pageId="1" pageNumber="2">Prosopis</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C3BC0A5FF8CFFCE8834F945FE7787B1" box="[314,428,1759,1785]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), acacia (
<emphasis id="B94F6734FF8CFFCE8B13F945FDB587B0" box="[541,622,1759,1784]" italics="true" pageId="1" pageNumber="2">Acacia</emphasis>
sp.,
<taxonomicName id="4C3BC0A5FF8CFFCE8BA4F945FCC787B1" box="[682,796,1759,1785]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), and desert shrubs. A total of 34 sites were selected within these zones to monitor arthropods. This information was summarized from the Grand Canyon Arthropod Inventory &amp; Monitoring website: http://bugs.bio.nau.edu/ grand_canyon/index.htm.
</paragraph>
</subSubSection>
<caption id="DF44EBAEFF8FFFCD8806FA1FFB43874C" httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="2" pageNumber="3" targetBox="[300,1299,285,1389]" targetPageId="2">
<paragraph id="8B84BB26FF8FFFCD8806FA1FFB43874C" blockId="2.[264,1323,1413,1541]" pageId="2" pageNumber="3">
<emphasis id="B94F6734FF8FFFCD8806FA1FFE6D84D5" bold="true" box="[264,438,1413,1437]" pageId="2" pageNumber="3">FIGURES 16.</emphasis>
Adults and male genitalia. 1,
<emphasis id="B94F6734FF8FFFCD8A0AFA1CFC3984D5" box="[772,994,1414,1437]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C3BC0A5FF8FFFCD8A0AFA1CFC0584D5" box="[772,990,1414,1437]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="immaculata">Schinia immaculata</taxonomicName>
,
</emphasis>
male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2,
<emphasis id="B94F6734FF8FFFCD8D49FA32FB0E84F7" box="[1095,1237,1448,1471]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4C3BC0A5FF8FFFCD8D49FA32FB0A84F7" box="[1095,1233,1448,1471]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="biundulata">S biundulata</taxonomicName>
,
</emphasis>
female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3,
<taxonomicName id="4C3BC0A5FF8FFFCD8A93FA51FBE184AA" box="[925,1082,1483,1506]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8FFFCD8A93FA51FBE184AA" box="[925,1082,1483,1506]" italics="true" pageId="2" pageNumber="3">S. immaculata</emphasis>
</taxonomicName>
, male genitalia; 4,
<taxonomicName id="4C3BC0A5FF8FFFCD8C18FA51FE5D874D" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8FFFCD8C18FA51FE5D874D" italics="true" pageId="2" pageNumber="3">S. immaculata</emphasis>
</taxonomicName>
, aedoeagus; 5,
<taxonomicName id="4C3BC0A5FF8FFFCD8B25FA74FD61874D" box="[555,698,1518,1541]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B94F6734FF8FFFCD8B25FA74FD61874D" box="[555,698,1518,1541]" italics="true" pageId="2" pageNumber="3">S. biundulata</emphasis>
</taxonomicName>
, male genitalia; 6,
<taxonomicName id="4C3BC0A5FF8FFFCD8A8BFA74FBC8874D" box="[901,1043,1518,1541]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="biundulata">
<emphasis id="B94F6734FF8FFFCD8A8BFA74FBC8874D" box="[901,1043,1518,1541]" italics="true" pageId="2" pageNumber="3">S. biundulata</emphasis>
</taxonomicName>
, aedoeagus.
</paragraph>
</caption>
<subSubSection id="C321E8ADFF8FFFCD8836F9FAFE4A8622" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8B84BB26FF8FFFCD8836F9FAFE4A8622" blockId="2.[264,1323,1632,1898]" pageId="2" pageNumber="3">
<emphasis id="B94F6734FF8FFFCD8836F9FAFDEE8732" bold="true" box="[312,565,1632,1658]" pageId="2" pageNumber="3">Material Examined.</emphasis>
<typeStatus id="54800584FF8FFFCD8B4EF9FAFD398732" box="[576,738,1632,1658]" pageId="2" pageNumber="3" type="paratype">PARATYPES</typeStatus>
(3°):
<emphasis id="B94F6734FF8FFFCD8A23F9FAFBFD8732" bold="true" box="[813,1062,1632,1658]" pageId="2" pageNumber="3">
<collectingCountry id="F32CFBB6FF8FFFCD8A23F9FAFC5A8732" box="[813,897,1632,1658]" name="United States of America" pageId="2" pageNumber="3">U.S.A.</collectingCountry>
: ARIZONA:
</emphasis>
COCONINO CO. 1° same data as
<typeStatus id="54800584FF8FFFCD88ADF912FDD187EA" box="[419,522,1672,1698]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
except:
<collectionCode id="ED2A23E3FF8FFFCD8B62F912FD1A87EA" LSID="urn:lsid:biocol.org:col:34871" box="[620,705,1672,1698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
<collectionCode id="ED2A23E3FF8FFFCD8BC6F912FCDA87EA" LSID="urn:lsid:biocol.org:col:13589" box="[712,769,1672,1698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="2" pageNumber="3">ENT</collectionCode>
0 0
<date id="FF859DE6FF8FFFCD8A2AF912FCA587EA" box="[804,894,1672,1698]" pageId="2" pageNumber="3" value="20-01-21">210120</date>
(
<collectionCode id="ED2A23E3FF8FFFCD8A83F912FC0887EA" LSID="urn:lsid:biocol.org:col:13688" box="[909,979,1672,1698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13688" name="Nanjing Agricultural University, Department of Plant Science" pageId="2" pageNumber="3">NAU</collectionCode>
); river mile 166.5 L, old high water,
<geoCoordinate id="EE0FDDE1FF8FFFCD8854F92AFE008782" box="[346,475,1712,1738]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="5" value="36.2542">36.2542 N</geoCoordinate>
,
<geoCoordinate id="EE0FDDE1FF8FFFCD88E4F92AFDA58782" box="[490,638,1712,1738]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="5" value="-112.8996">112.8996 W</geoCoordinate>
,
<date id="FF859DE6FF8FFFCD8B82F92AFCF68782" box="[652,813,1712,1738]" pageId="2" pageNumber="3" value="2003-04-14">14 Apr. 2003</date>
(1°), R. J. Delph,
<collectionCode id="ED2A23E3FF8FFFCD8D1DF92AFBB38782" LSID="urn:lsid:biocol.org:col:34871" box="[1043,1128,1712,1738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
<collectionCode id="ED2A23E3FF8FFFCD8D7CF92AFB708782" LSID="urn:lsid:biocol.org:col:13589" box="[1138,1195,1712,1738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="2" pageNumber="3">ENT</collectionCode>
0 0 219965 (
<collectionCode id="ED2A23E3FF8FFFCD881EF942FEB087BA" LSID="urn:lsid:biocol.org:col:34871" box="[272,363,1752,1778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
); river mile 202 R, new high water,
<geoCoordinate id="EE0FDDE1FF8FFFCD8A1FF942FC4B87BA" box="[785,912,1752,1778]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="5" value="36.0526">36.0526 N</geoCoordinate>
,
<geoCoordinate id="EE0FDDE1FF8FFFCD8A92F942FBF787BA" box="[924,1068,1752,1778]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="5" value="-113.3489">113.3489 W</geoCoordinate>
,
<date id="FF859DE6FF8FFFCD8D36F942FB0F87BA" box="[1080,1236,1752,1778]" pageId="2" pageNumber="3" value="2001-05-15">15 May 2001</date>
(1°), J. Rundall,
<collectionCode id="ED2A23E3FF8FFFCD887BF89AFE118652" LSID="urn:lsid:biocol.org:col:34871" box="[373,458,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
<collectionCode id="ED2A23E3FF8FFFCD88DCF89AFDD08652" LSID="urn:lsid:biocol.org:col:13589" box="[466,523,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="2" pageNumber="3">ENT</collectionCode>
0 0
<date id="FF859DE6FF8FFFCD8B3EF89AFD528652" box="[560,649,1792,1818]" pageId="2" pageNumber="3" value="19-01-21">210119</date>
(
<collectionCode id="ED2A23E3FF8FFFCD8B94F89AFD3B8652" LSID="urn:lsid:biocol.org:col:13688" box="[666,736,1792,1818]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13688" name="Nanjing Agricultural University, Department of Plant Science" pageId="2" pageNumber="3">NAU</collectionCode>
).
<typeStatus id="54800584FF8FFFCD8BFBF89AFCBC8652" box="[757,871,1792,1818]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
deposited in the National Museum of Natural History, Washington, DC (
<collectionCode id="ED2A23E3FF8FFFCD8BA0F8B2FCDD860A" LSID="urn:lsid:biocol.org:col:34871" box="[686,774,1832,1858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
) and Northern Arizona University, Flagstaff, AZ (
<collectionCode id="ED2A23E3FF8FFFCD8830F8CAFE5F8622" LSID="urn:lsid:biocol.org:col:13688" box="[318,388,1872,1898]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13688" name="Nanjing Agricultural University, Department of Plant Science" pageId="2" pageNumber="3">NAU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8EFFCC8836FE81FDFA8016" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8B84BB26FF8EFFCC8836FE81FDFA8016" blockId="3.[264,1324,283,630]" pageId="3" pageNumber="4">
<emphasis id="B94F6734FF8EFFCC8836FE81FE13807D" bold="true" box="[312,456,283,309]" pageId="3" pageNumber="4">Etymology:</emphasis>
The specific epithet,
<taxonomicName id="4C3BC0A5FF8EFFCC8BC4FE86FC8F807D" box="[714,852,284,309]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8EFFCC8BC4FE86FC8F807D" box="[714,852,284,309]" italics="true" pageId="3" pageNumber="4">immaculata</emphasis>
</taxonomicName>
, refers to the lack of any pattern on the forewing and hindwing.
</paragraph>
</subSubSection>
<subSubSection id="C321E8ADFF8EFFCC8836FEF1FC7B833E" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8B84BB26FF8EFFCC8836FEF1FC7B833E" blockId="3.[264,1324,283,630]" pageId="3" pageNumber="4">
<emphasis id="B94F6734FF8EFFCC8836FEF1FE1880CD" bold="true" box="[312,451,363,389]" pageId="3" pageNumber="4">Discussion.</emphasis>
Unlike most species of
<taxonomicName id="4C3BC0A5FF8EFFCC8BEEFEF6FCE380CD" box="[736,824,364,389]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B94F6734FF8EFFCC8BEEFEF6FCE380CD" box="[736,824,364,389]" italics="true" pageId="3" pageNumber="4">Schinia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3BC0A5FF8EFFCC8A49FEF6FC3480CD" box="[839,1007,364,389]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8EFFCC8A49FEF6FC3480CD" box="[839,1007,364,389]" italics="true" pageId="3" pageNumber="4">S. immaculata</emphasis>
</taxonomicName>
has no forewing markings in all but one
<typeStatus id="54800584FF8EFFCC88B8FE0EFDC580E6" box="[438,542,404,430]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
, which has a barely perceptible trace of both an antemedial and postmedial line (
<figureCitation id="1300A7A3FF8EFFCC88DAFE26FDC6809E" box="[468,541,444,470]" captionStart="FIGURES 1 6" captionStartId="2.[264,382,1413,1437]" captionTargetBox="[300,1299,285,1389]" captionTargetId="figure@2.[270,1317,280,1409]" captionTargetPageId="2" captionText="FIGURES 1 6. Adults and male genitalia. 1, Schinia immaculata, male, Arizona, Coconino Co. Colorado River, Grand Canyon, river mile 166.5 L, USNMENT 00229965; 2, S biundulata, female, Nevada, Humboldt Co. Sulphur, USNMENT 00220807; 3, S. immaculata, male genitalia; 4, S. immaculata, aedoeagus; 5, S. biundulata, male genitalia; 6, S. biundulata, aedoeagus." httpUri="https://zenodo.org/record/169633/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
). The form of the foretibia, male genitalia, and the coils of the aedoeagus are typical of
<taxonomicName id="4C3BC0A5FF8EFFCC8B37FE7EFD4A80B5" box="[569,657,484,509]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B94F6734FF8EFFCC8B37FE7EFD4A80B5" box="[569,657,484,509]" italics="true" pageId="3" pageNumber="4">Schinia</emphasis>
</taxonomicName>
. The species was collected in all riparian zones of the study area.
<taxonomicName id="4C3BC0A5FF8EFFCC8881FD96FDA2836D" box="[399,633,524,549]" class="Insecta" family="Noctuidae" genus="Schinia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="immaculata">
<emphasis id="B94F6734FF8EFFCC8881FD96FDA2836D" box="[399,633,524,549]" italics="true" pageId="3" pageNumber="4">Schinia immaculata</emphasis>
</taxonomicName>
is a spring flyer, being active from mid­April to mid­May. Other large river systems in the southwestern
<collectingCountry id="F32CFBB6FF8EFFCC8A11FDAEFC608306" box="[799,955,564,590]" name="United States of America" pageId="3" pageNumber="4">United States</collectingCountry>
should be investigated to try to learn more about the distribution of this unusual species.
</paragraph>
</subSubSection>
</treatment>
</document>