342 lines
39 KiB
XML
342 lines
39 KiB
XML
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<mods:title id="092F58A7C3D8069BFFAAB2B96EF36F0C">Taxonomic notes concerning the genus Catharsius Hope, 1837 (Scarabaeidae Scarabaeinae)</mods:title>
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<mods:namePart id="08DAEC83648B48B4216C9928F3B3C468">Takano, Hitoshi</mods:namePart>
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<mods:affiliation id="E339011EE276DA687222AC2F505789E8">African Natural History Research Trust, Street Court, Kingsland, Leominster HR 6 9 QA, UK. & Department of Zoology, Oxford University, South Parks Road, Oxford OX 1 3 PS, UK & Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:title id="F6CCCFA61093B41DA37A8C989DD65AD0">Zootaxa</mods:title>
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<mods:part id="FA9285FB6511F6493A8292AAE836AB2D">
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<mods:date id="346BFD2F34994BA0462C45EE130CD1B0">2021</mods:date>
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<mods:number id="3801E4DD2176AC5E0CA348F46B5620F7">2021-10-14</mods:number>
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<treatment id="865687A5FF94FFFD27974EEF2527FD69" ID-DOI="http://doi.org/10.5281/zenodo.5568716" ID-GBIF-Taxon="189077018" ID-Zenodo-Dep="5568716" LSID="urn:lsid:plazi:treatment:865687A5FF94FFFD27974EEF2527FD69" httpUri="http://treatment.plazi.org/id/865687A5FF94FFFD27974EEF2527FD69" lastPageId="4" lastPageNumber="284" pageId="3" pageNumber="283">
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<subSubSection id="46E56538FF94FFFA27974EEF237CFE6C" box="[151,617,474,501]" pageId="3" pageNumber="283" type="nomenclature">
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<taxonomicName id="C9FF4D30FF94FFFA27974EEF206EFE6D" ID-CoL="8RXXJ" authority="Takano, 2021" authorityName="Takano" authorityYear="2021" box="[151,379,474,501]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="283" phylum="Arthropoda" rank="species" species="haroldi" status="sp. nov.">
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<emphasis id="3C8BEAA1FF94FFFA27974EEF206EFE6D" bold="true" box="[151,379,474,501]" italics="true" pageId="3" pageNumber="283">Catharsius haroldi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="27B857DAFF94FFFA26824EEE20F5FE6D" box="[386,480,475,501]" pageId="3" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="96C42A36FF94FFFA26EF4EEF2374FE6C" box="[495,609,474,500]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 1–9. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="3" pageNumber="283">Figs. 6–9</figureCitation>
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)
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</emphasis>
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<taxonomicName id="C9FF4D30FF94FFFA27974D2A2354FDAF" ID-CoL="8R7Y8" authority="sensu Ferreira, 1960" authorityName="sensu Ferreira" authorityYear="1960" box="[151,577,543,567]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="283" phylum="Arthropoda" rank="species" species="harpagus">
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<emphasis id="3C8BEAA1FF94FFFA27974D2A2078FDAF" box="[151,365,543,567]" italics="true" pageId="3" pageNumber="283">Catharsius harpagus</emphasis>
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sensu
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<bibRefCitation id="6A6E4B42FF94FFFA26B14D2A2354FDAF" author="Ferreira, M. C." box="[433,577,543,567]" pageId="3" pageNumber="283" pagination="155 - 270" refId="ref4027" refString="Ferreira, M. C. (1960) Revisao das especies Etiopicas do subgenera Catharsius s. str. Revista Entomologia Mocambique, 3 (2), 155 - 270." type="journal article" year="1960">Ferreira, 1960</bibRefCitation>
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</taxonomicName>
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(nec Harold): 225; 1964a: 7; 1967: 234; 1972: 270;
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<treatmentCitation id="8F5E10A2FF94FFFA23554D2A2472FDAF" ID-DOI="http://doi.org/10.5281/zenodo.6079997" ID-Zenodo-Dep="6079997" author="Genier, F. & Josso, J. - F." box="[1109,1383,543,567]" httpUri="http://treatment.plazi.org/id/03B487F3FFC8FFBD9396FBE5FB40F91D" page="335" pageId="3" pageNumber="283" year="2016">
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<bibRefCitation id="6A6E4B42FF94FFFA23554D2A2472FDAF" author="Genier, F. & Josso, J. - F." box="[1109,1383,543,567]" pageId="3" pageNumber="283" pagination="333 - 342" refId="ref4365" refString="Genier, F. & Josso, J. - F. (2016) Notes on Afrotropical Catharsius species described by Edgar von Harold (Coleoptera: Scarabaeidae, Scarabaeinae). Zootaxa, 4072 (3), 333 - 342. https: // doi. org / 10.11646 / zootaxa. 4072.3.3" type="journal article" year="2016">Génier & Josso, 2016: 335</bibRefCitation>
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.
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<emphasis id="3C8BEAA1FF94FFFA27974D502058FDE7" bold="true" box="[151,333,613,639]" pageId="3" pageNumber="283">Type material.</emphasis>
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</paragraph>
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<typeStatus id="D1448811FF94FFFA26544D5020DDFDE7" box="[340,456,613,639]" pageId="3" pageNumber="283" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="18F9FD3AFF94FFFA26D54D5320FEFD1A" box="[469,491,614,642]" pageId="3" pageNumber="283" type="male">♂</specimenCount>
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(
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<collectionCode id="68EEAE76FF94FFFA26FB4D502348FDE7" box="[507,605,613,639]" pageId="3" pageNumber="283">BMNH</collectionCode>
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):
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</emphasis>
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</heading>
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“
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<collectingCountry id="76E87623FF94FFFA27D04DBC2081FD3B" box="[208,404,649,675]" name="Mozambique" pageId="3" pageNumber="283">MOZAMBIQUE</collectingCountry>
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,
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</quantity>
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, Transect 4, /
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<collectingRegion id="CC3BF851FF94FFFA256D4DBF23D1FD3C" box="[621,708,650,676]" country="Mozambique" name="Maputo" pageId="3" pageNumber="283">Maputo</collectingRegion>
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<collectingMunicipality id="EE24ACC9FF94FFFA25CC4DBC2291FD3C" box="[716,900,649,676]" pageId="3" pageNumber="283">Special Reserve</collectingMunicipality>
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, / (
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<location id="0B206068FF94FFFA24A94DBC25D5FD3B" LSID="urn:lsid:plazi:treatment:865687A5FF94FFFD27974EEF2527FD69:0B206068FF94FFFA24A94DBC25D5FD3B" box="[937,1216,649,675]" country="Mozambique" latitude="-26.48564" longLatPrecision="1" longitude="32.752472" municipality="Special Reserve" name="Hygrophilous Grassland" pageId="3" pageNumber="283" stateProvince="Maputo">Hygrophilous Grassland</location>
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,
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/ from
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) /
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,
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<collectingDate id="6A05E99BFF94FFFA25A14D98220AFD5F" box="[673,799,685,712]" pageId="3" pageNumber="283" value="2016-12-05">5.XII.2016</collectingDate>
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.
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<location id="0B206068FF94FFFA242B4D9B25D9FD5F" LSID="urn:lsid:plazi:treatment:865687A5FF94FFFD27974EEF2527FD69:0B206068FF94FFFA242B4D9B25D9FD5F" box="[811,1228,685,712]" country="Mozambique" latitude="-26.48564" longLatPrecision="1" longitude="32.752472" municipality="Special Reserve" name="Human Dung Pitfall / Aristophanous" pageId="3" pageNumber="283" stateProvince="Maputo">Human Dung Pitfall / Aristophanous</location>
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, M.,
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<collectorName id="A30A5365FF94FFFA22074D982482FD50" box="[1287,1431,685,712]" pageId="3" pageNumber="283">Cristóvão, J.</collectorName>
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, /
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<collectorName id="A30A5365FF94FFFA27A74DE4200BFD73" box="[167,286,721,747]" pageId="3" pageNumber="283">Laszlo, G.</collectorName>
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,
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<collectorName id="A30A5365FF94FFFA262C4DE4208FFD74" box="[300,410,721,748]" pageId="3" pageNumber="283">Miles, W.</collectorName>
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leg. /
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<collectionCode id="68EEAE76FF94FFFA26E44DE4232FFD73" box="[484,570,721,747]" pageId="3" pageNumber="283">BMNH</collectionCode>
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(E) 2016–262 /
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<location id="0B206068FF94FFFA25EC4DE42245FD73" LSID="urn:lsid:plazi:treatment:865687A5FF94FFFD27974EEF2527FD69:0B206068FF94FFFA25EC4DE42245FD73" box="[748,848,721,747]" country="Mozambique" latitude="-26.48564" longLatPrecision="1" longitude="32.752472" municipality="Special Reserve" name="Trip Ref" pageId="3" pageNumber="283" stateProvince="Maputo">Trip Ref</location>
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: MZ–001 (
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<specimenCode id="5E599EC8FF94FFFA24CE4DE42546FD73" box="[974,1107,721,747]" pageId="3" pageNumber="283">ANHRT 22</specimenCode>
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) //
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<specimenCode id="5E599EC8FF94FFFA237C4DE42471FD73" box="[1148,1380,721,747]" pageId="3" pageNumber="283">NHMUK010367768</specimenCode>
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[QR code]”.
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<materialsCitation id="BE973CEEFF94FFFA27C74C2C2316FCAE" ID-GBIF-Occurrence="3387267303" box="[199,515,793,822]" pageId="3" pageNumber="283" typeStatus="paratype">
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<typeStatus id="D1448811FF94FFFA27C74C2C2051FCAB" box="[199,324,793,819]" pageId="3" pageNumber="283" type="paratype">Paratypes</typeStatus>
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(
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):
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<paragraph id="0E4036B3FF94FFFA27C74C082565FBA8" blockId="3.[151,1437,613,2008]" pageId="3" pageNumber="283">
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<collectingCountry id="76E87623FF94FFFA27C74C082086FCCF" box="[199,403,829,855]" name="Mozambique" pageId="3" pageNumber="283">MOZAMBIQUE</collectingCountry>
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</emphasis>
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:
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<collectingRegion id="CC3BF851FF94FFFA269F4C082300FCCF" box="[415,533,829,855]" country="Mozambique" name="Maputo" pageId="3" pageNumber="283">MAPUTO</collectingRegion>
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: Maputo Special Reserve,
|
||
<quantity id="C9079B56FF94FFFA24364C08224FFCCF" box="[822,858,829,855]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="283" unit="m" value="2.0">2m</quantity>
|
||
(
|
||
<geoCoordinate id="6BCB5074FF94FFFA24664C0822D0FCCF" box="[870,965,829,855]" degrees="26" direction="south" minutes="29" orientation="latitude" pageId="3" pageNumber="283" precision="925" value="-26.483334">26°29’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BCB5074FF94FFFA24CD4C08253EFCC0" box="[973,1067,829,856]" degrees="32" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="283" precision="925" value="32.75">32°45’E</geoCoordinate>
|
||
),
|
||
<date id="7A411073FF94FFFA233F4C0825DDFCC0" box="[1087,1224,829,856]" pageId="3" pageNumber="283" value="2017-06-04" valueMax="2017-06-05" valueMin="2017-06-04">4–5.vi.2017</date>
|
||
, M.Aristophanous, G. László, W. Miles & A. Vetina (
|
||
<specimenCount id="18F9FD3AFF94FFFA25164C54232CFCE3" box="[534,569,865,891]" pageId="3" pageNumber="283" type="male">1♂</specimenCount>
|
||
<specimenCount id="18F9FD3AFF94FFFA25404C54236DFCE3" box="[576,632,865,891]" pageId="3" pageNumber="283" type="female">3♀♀</specimenCount>
|
||
<collectionCode id="68EEAE76FF94FFFA257F4C5723CDFCE4" box="[639,728,866,892]" pageId="3" pageNumber="283">BMNH</collectionCode>
|
||
);
|
||
<materialsCitation id="BE973CEEFF94FFFA25EC4C57256CFC38" ID-GBIF-Occurrence="3387267301" pageId="3" pageNumber="283">
|
||
Maputo Special Reserve, Transect
|
||
<quantity id="C9079B56FF94FFFA23734C5425A6FCE3" box="[1139,1203,865,891]" metricMagnitude="0" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="283" unit="m" value="4.2">4, 2m</quantity>
|
||
(
|
||
<geoCoordinate id="6BCB5074FF94FFFA23C34C542437FCE3" box="[1219,1314,865,891]" degrees="26" direction="south" minutes="29" orientation="latitude" pageId="3" pageNumber="283" precision="925" value="-26.483334">26°29’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BCB5074FF94FFFA222E4C54249AFCE4" box="[1326,1423,865,892]" degrees="32" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="283" precision="925" value="32.75">32°45’E</geoCoordinate>
|
||
),
|
||
<date id="7A411073FF94FFFA27974CB0201EFC38" box="[151,267,901,928]" pageId="3" pageNumber="283" value="2016-12-05">5.xii.2016</date>
|
||
, M. Aristophanous, J. Cristóvão, G. László & W. Miles (
|
||
<specimenCount id="18F9FD3AFF94FFFA24934CB022DEFC38" box="[915,971,901,928]" pageId="3" pageNumber="283" type="male">5♂♂</specimenCount>
|
||
<specimenCount id="18F9FD3AFF94FFFA24D44CB0251EFC07" box="[980,1035,901,927]" pageId="3" pageNumber="283" type="female">6♀♀</specimenCount>
|
||
<collectionCode id="68EEAE76FF94FFFA23144CB32578FC38" box="[1044,1133,902,928]" pageId="3" pageNumber="283">BMNH</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="BE973CEEFF94FFFA23814CB321E9FC70" ID-GBIF-Occurrence="3387267302" pageId="3" pageNumber="283">
|
||
Maputo Special Reserve, Transect
|
||
<quantity id="C9079B56FF94FFFA27FE4C9C202AFC5B" box="[254,319,937,964]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="283" unit="m" value="5.2">5, 2m</quantity>
|
||
(
|
||
<geoCoordinate id="6BCB5074FF94FFFA264F4C9C20BBFC5B" box="[335,430,937,963]" degrees="26" direction="south" minutes="29" orientation="latitude" pageId="3" pageNumber="283" precision="925" value="-26.483334">26°29’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BCB5074FF94FFFA26BA4C9C230EFC5C" box="[442,539,937,964]" degrees="32" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="283" precision="925" value="32.75">32°45’E</geoCoordinate>
|
||
),
|
||
<date id="7A411073FF94FFFA252F4C9C23B6FC5C" box="[559,675,937,964]" pageId="3" pageNumber="283" value="2016-12-05">5.xii.2016</date>
|
||
, M. Aristophanous, J. Cristóvão, G. László & W. Miles (
|
||
<specimenCount id="18F9FD3AFF94FFFA22244C9C2448FC5B" box="[1316,1373,937,963]" pageId="3" pageNumber="283" type="male">2♂♂</specimenCount>
|
||
<specimenCount id="18F9FD3AFF94FFFA22644C9C2489FC5C" box="[1380,1436,937,964]" pageId="3" pageNumber="283" type="female">5♀♀</specimenCount>
|
||
<collectionCode id="68EEAE76FF94FFFA27974CFB21E5FC70" box="[151,240,974,1000]" pageId="3" pageNumber="283">BMNH</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="BE973CEEFF94FFFA26034CFB2387FB94" ID-GBIF-Occurrence="3387267307" pageId="3" pageNumber="283">
|
||
Maputo Special Reserve, Transect
|
||
<quantity id="C9079B56FF94FFFA25894CF823DFFC7F" box="[649,714,973,999]" metricMagnitude="0" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="283" unit="m" value="6.2">6, 2m</quantity>
|
||
(
|
||
<geoCoordinate id="6BCB5074FF94FFFA25D94CF8222DFC7F" box="[729,824,973,999]" degrees="26" direction="south" minutes="29" orientation="latitude" pageId="3" pageNumber="283" precision="925" value="-26.483334">26°29’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BCB5074FF94FFFA24444CF822B4FC70" box="[836,929,973,1000]" degrees="32" direction="east" minutes="45" orientation="longitude" pageId="3" pageNumber="283" precision="925" value="32.75">32°45’E</geoCoordinate>
|
||
),
|
||
<date id="7A411073FF94FFFA24B94CF82538FC70" box="[953,1069,973,1000]" pageId="3" pageNumber="283" value="2016-12-05">5.xii.2016</date>
|
||
, M. Aristophanous, J. Cristóvão, G. László & W. Miles (
|
||
<specimenCount id="18F9FD3AFF94FFFA26A94CC420F7FB93" box="[425,482,1009,1035]" pageId="3" pageNumber="283" type="male">6♂♂</specimenCount>
|
||
<specimenCount id="18F9FD3AFF94FFFA26EC4CC42336FB93" box="[492,547,1009,1035]" pageId="3" pageNumber="283" type="female">4♀♀</specimenCount>
|
||
<collectionCode id="68EEAE76FF94FFFA252D4CC72393FB94" box="[557,646,1010,1036]" pageId="3" pageNumber="283">BMNH</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="BE973CEEFF94FFFA259C4CC72578FBA8" ID-GBIF-Occurrence="3387267304" pageId="3" pageNumber="283">
|
||
Maputo Special Reserve, West Gate,
|
||
<quantity id="C9079B56FF94FFFA23494CC4256EFB93" box="[1097,1147,1009,1035]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="283" unit="m" value="22.0">22m</quantity>
|
||
(
|
||
<geoCoordinate id="6BCB5074FF94FFFA238D4CC425F8FB93" box="[1165,1261,1009,1035]" degrees="26" direction="south" minutes="30" orientation="latitude" pageId="3" pageNumber="283" precision="925" value="-26.5">26°30’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="6BCB5074FF94FFFA23FB4CC42449FB93" box="[1275,1372,1009,1035]" degrees="32" direction="east" minutes="43" orientation="longitude" pageId="3" pageNumber="283" precision="925" value="32.716667">32°43’E</geoCoordinate>
|
||
),
|
||
<date id="7A411073FF94FFFA22724CC42004FBB7" pageId="3" pageNumber="283" value="2016-11-21" valueMax="2016-11-30" valueMin="2016-11-21">21– 30.xi.2016</date>
|
||
, M. Aristophanous, J. Cristóvão, G. László & W. Miles (
|
||
<specimenCount id="18F9FD3AFF94FFFA248D4B2022D3FBB7" box="[909,966,1045,1071]" pageId="3" pageNumber="283" type="male">2♂♂</specimenCount>
|
||
<specimenCount id="18F9FD3AFF94FFFA24CD4B202510FBB7" box="[973,1029,1045,1071]" pageId="3" pageNumber="283" type="female">2♀♀</specimenCount>
|
||
<collectionCode id="68EEAE76FF94FFFA230C4B232570FBA8" box="[1036,1125,1046,1072]" pageId="3" pageNumber="283">BMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46E56538FF94FFFD27C74B0C2057FE49" lastPageId="4" lastPageNumber="284" pageId="3" pageNumber="283" type="description">
|
||
<paragraph id="0E4036B3FF94FFFA27C74B0C21E7F92F" blockId="3.[151,1437,613,2008]" pageId="3" pageNumber="283">
|
||
<emphasis id="3C8BEAA1FF94FFFA27C74B0C231BFBCB" bold="true" box="[199,526,1081,1107]" pageId="3" pageNumber="283">
|
||
Description. Male
|
||
<typeStatus id="D1448811FF94FFFA26A74B0C231BFBCB" box="[423,526,1081,1107]" pageId="3" pageNumber="283" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="96C42A36FF94FFFA25204B0C239BFBCC" box="[544,654,1081,1108]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 1–9. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="3" pageNumber="283">Figs. 6–7</figureCitation>
|
||
).
|
||
<emphasis id="3C8BEAA1FF94FFFA25A44B0C23CBFBCB" bold="true" box="[676,734,1081,1107]" pageId="3" pageNumber="283">Size:</emphasis>
|
||
Length
|
||
<quantity id="C9079B56FF94FFFA24404B0C2286FBCC" box="[832,915,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="283" unit="mm" value="25.0">25 mm</quantity>
|
||
, width
|
||
<quantity id="C9079B56FF94FFFA24E94B0C2528FBCC" box="[1001,1085,1081,1108]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="283" unit="mm" value="17.0">17 mm</quantity>
|
||
.
|
||
<emphasis id="3C8BEAA1FF94FFFA234A4B0C258BFBCB" bold="true" box="[1098,1182,1081,1107]" pageId="3" pageNumber="283">Colour</emphasis>
|
||
: Black with a reddish undertone; pubescence light brown.
|
||
<emphasis id="3C8BEAA1FF94FFFA25344B682368FBEF" bold="true" box="[564,637,1117,1143]" pageId="3" pageNumber="283">Head:</emphasis>
|
||
Anterior clypeal edge evenly arcuate with continuous clypeal margin. Clypeal surface rugose anteriorly, becoming more granulose posteriorly; genal edge rounded and acutely angled, surface with granules disappearing and becoming glabrous internally. Clypeogenal suture well-defined to base of cephalic horn. Cephalic horn long and straight, wide in basal third, edges of basal portion converging slightly; apicolateral angles rounded and obtuse. Cephalic suture laterally well-defined from base to apex.
|
||
<emphasis id="3C8BEAA1FF94FFFA221A4BD82489FA9F" bold="true" box="[1306,1436,1261,1287]" pageId="3" pageNumber="283">Pronotum:</emphasis>
|
||
Anterior margin continuous, obliquely orientated anteriorly on each side of eyes. Lateral edges broadly arcuate. Pronotal disc with short but sharply pointed horns anteriorly, gently diverging and curved upwards, placed evenly between midline and lateral margin; apices of horns in line with edge of clypeogenal suture. Pronotal disc with continuous margin posteriorly; surface of disc covered in fine granules except for a glabrous oviform region on each side of midline; lateral declivity with fovea; surface completely granulate. Surface of anterior declivity glabrous; in frontal view, glabrous area extends from horns to anterior margin.
|
||
<emphasis id="3C8BEAA1FF94FFFA24AA4AF022EAFA47" bold="true" box="[938,1023,1477,1503]" pageId="3" pageNumber="283">Elytra:</emphasis>
|
||
Carinate basally; elytral striae welldefined, with weak punctures encroaching slightly on intervals; surface of interstriae glabrous except for
|
||
<quantity id="C9079B56FF94FFFA22594ADC2488F99C" box="[1369,1437,1513,1540]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="283" unit="mm" value="2.0">2 mm</quantity>
|
||
portion basally of fine alutaceous microsculpture; carina on ninth interval very short.
|
||
<emphasis id="3C8BEAA1FF94FFFA235B493825CDF9BF" bold="true" box="[1115,1240,1549,1575]" pageId="3" pageNumber="283">Pygidium:</emphasis>
|
||
Transverse, fully margined, glabrous centrally, becoming more alutaceous towards edges; fine oviform punctures throughout.
|
||
<emphasis id="3C8BEAA1FF94FFFA225B49042489F9D3" bold="true" box="[1371,1436,1585,1611]" pageId="3" pageNumber="283">Legs:</emphasis>
|
||
Protibia tridentate; spur curved gently apically; three longitudinal striae ventrally. Metatibial spur long, truncate apically.
|
||
<emphasis id="3C8BEAA1FF94FFFA27FE494C206EF90B" bold="true" box="[254,379,1657,1683]" pageId="3" pageNumber="283">Aedeagus:</emphasis>
|
||
Parameres almost as long as phallobase, laterally arcuate, with shallow concavity anteriorly (
|
||
<figureCitation id="96C42A36FF94FFFA27A049A821F1F92F" box="[160,228,1693,1719]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 1–9. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="3" pageNumber="283">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0E4036B3FF94FFFD27C749F42082FEFD" blockId="3.[151,1437,613,2008]" lastBlockId="4.[151,1437,151,753]" lastPageId="4" lastPageNumber="284" pageId="3" pageNumber="283">
|
||
<emphasis id="3C8BEAA1FF94FFFA27C749F42083F943" bold="true" box="[199,406,1729,1755]" pageId="3" pageNumber="283">
|
||
Female
|
||
<typeStatus id="D1448811FF94FFFA262B49F42083F943" box="[299,406,1729,1755]" pageId="3" pageNumber="283" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="96C42A36FF94FFFA26AB49F420E0F943" box="[427,501,1729,1755]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1842,1866]" captionTargetBox="[232,1332,187,1804]" captionTargetId="figure-15@2.[219,1368,179,1816]" captionTargetPageId="2" captionText="FIGURES 1–9. Catharsius Hope, 1837 species. 1, C. harpagus Harold, 1877 (lectotype ♂), habitus, dorsal view; 2, idem, associated labels; 3, C. laticeps sensu Péringuey (nec Boheman), reproduction of figures 23 and 23a from plate XXXIII of Péringuey (1901); 4, C. parafastidiosus Ferreira, 1971 (holotype ♂), habitus, dorsal view; 5, idem, associated labels; 6, C. haroldi sp. nov. (holotype ♂), habitus, dorsal view; 7, idem, lateral view; 8, idem, aedeagus, lateral view; 9, C. haroldi sp. nov. (paratype ♀), habitus, dorsal view. Scale equals 5 mm unless otherwise specified." figureDoi="http://doi.org/10.5281/zenodo.5568730" httpUri="https://zenodo.org/record/5568730/files/figure.png" pageId="3" pageNumber="283">Fig. 9</figureCitation>
|
||
).
|
||
<emphasis id="3C8BEAA1FF94FFFA251049F4234CF943" bold="true" box="[528,601,1729,1755]" pageId="3" pageNumber="283">Head:</emphasis>
|
||
Anterior clypeal edge evenly arcuate with continuous clypeal margin. Clypeal surface transversely rugose anteriorly, becoming more granulose posteriorly; genal edge rounded and acutely angled, surface with granules disappearing and becoming glabrous internally. Clypeogenal suture welldefined to base of clypeal carina. Clypeal carina raised, broadly triangular in frontal view and weakly tridentate, central point slightly bifurcate.
|
||
<emphasis id="3C8BEAA1FF94FFFA26FA48642369F8F3" bold="true" box="[506,636,1873,1899]" pageId="3" pageNumber="283">Pronotum:</emphasis>
|
||
Anterior margin continuous, obliquely oriented anteriorly on each side of eyes. Lateral edges broadly arcuate. Pronotal disc with short carina anteriorly, arcuate and medially emarginate; carina no longer than distance between eyes. Surface of disc completely granulose except for a small glabrous region on either side of midline; glabrous area rounded internally, tapering to a point laterally. Lateral declivity with fovea; surface completely granulate. Surface of anterior declivity completely granulate.
|
||
<emphasis id="3C8BEAA1FF93FFFD23804FA225C0FF29" bold="true" box="[1152,1237,151,177]" pageId="4" pageNumber="284">Elytra:</emphasis>
|
||
Carinate basally; elytral striae well-defined, with weak punctures encroaching slightly on intervals; surface of interstriae glabrous except for
|
||
<quantity id="C9079B56FF93FFFD260D4FEA2058FF61" box="[269,333,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="284" unit="mm" value="2.0">2 mm</quantity>
|
||
portion anteriorly of fine alutaceous microsculpture; carina on ninth interval very short.
|
||
<emphasis id="3C8BEAA1FF93FFFD22204FEB2489FF60" bold="true" box="[1312,1436,222,248]" pageId="4" pageNumber="284">Pygidium:</emphasis>
|
||
Transverse, fully margined, glabrous centrally, becoming more alutaceous towards edges; fine oviform punctures throughout.
|
||
<emphasis id="3C8BEAA1FF93FFFD261F4E122042FED9" bold="true" box="[287,343,295,321]" pageId="4" pageNumber="284">Legs</emphasis>
|
||
: Protibia tridentate; spur curved gently apically; three longitudinal striae ventrally. Metatibial spur long, truncate apically.
|
||
</paragraph>
|
||
<paragraph id="0E4036B3FF93FFFD27C74E5B2057FE49" blockId="4.[151,1437,151,753]" pageId="4" pageNumber="284">
|
||
<emphasis id="3C8BEAA1FF93FFFD27C74E5B202AFE10" bold="true" box="[199,319,366,392]" pageId="4" pageNumber="284">Variation.</emphasis>
|
||
Length: male
|
||
<quantity id="C9079B56FF93FFFD26E54E5B2377FE11" box="[485,610,366,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="4" pageNumber="284" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 mm</quantity>
|
||
, female
|
||
<quantity id="C9079B56FF93FFFD25C34E5A2255FE11" box="[707,832,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="4" pageNumber="284" unit="mm" value="23.5" valueMax="26.0" valueMin="21.0">21–26 mm</quantity>
|
||
. In smaller males the anterior declivity is completely granulose; the pronotal horns are reduced to tubercles and the cephalic horn is reduced to a broad equilateral triangle in frontal view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46E56538FF93FFFD27C74EEE25CCFD81" pageId="4" pageNumber="284" type="etymology">
|
||
<paragraph id="0E4036B3FF93FFFD27C74EEE25CCFD81" blockId="4.[151,1437,151,753]" pageId="4" pageNumber="284">
|
||
<emphasis id="3C8BEAA1FF93FFFD27C74EEE208BFE6D" bold="true" box="[199,414,475,501]" pageId="4" pageNumber="284">
|
||
<emphasis id="3C8BEAA1FF93FFFD27C74EEE2083FE6D" bold="true" box="[199,406,475,501]" italics="true" pageId="4" pageNumber="284">Derivatio nominis</emphasis>
|
||
.
|
||
</emphasis>
|
||
This species is named after the coleopterist Edgar von Harold (1830–1886), who described
|
||
<taxonomicName id="C9FF4D30FF93FFFD27974ECA2096FD81" authorityName="sensu Ferreira" authorityYear="1960" box="[151,387,511,537]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="284" phylum="Arthropoda" rank="species" species="harpagus">
|
||
<emphasis id="3C8BEAA1FF93FFFD27974ECA2096FD81" box="[151,387,511,537]" italics="true" pageId="4" pageNumber="284">Catharsius harpagus</emphasis>
|
||
</taxonomicName>
|
||
, the species with which subsequent authors have confused this new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46E56538FF93FFFD27C74D17239AFD55" pageId="4" pageNumber="284" type="diagnosis">
|
||
<paragraph id="0E4036B3FF93FFFD27C74D17239AFD55" blockId="4.[151,1437,151,753]" pageId="4" pageNumber="284">
|
||
<emphasis id="3C8BEAA1FF93FFFD27C74D172057FDA4" bold="true" box="[199,322,546,572]" pageId="4" pageNumber="284">Diagnosis.</emphasis>
|
||
This species, although very distinctive, might be confused with the smaller
|
||
<taxonomicName id="C9FF4D30FF93FFFD238D4D1621CCFDF9" authority="(De Geer, 1778)" baseAuthorityName="De Geer" baseAuthorityYear="1778" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="284" phylum="Arthropoda" rank="species" species="tricornutus">
|
||
<emphasis id="3C8BEAA1FF93FFFD238D4D16243CFDA4" box="[1165,1321,547,573]" italics="true" pageId="4" pageNumber="284">C. tricornutus</emphasis>
|
||
(De Geer, 1778)
|
||
</taxonomicName>
|
||
with which it is found sympatrically. In the males of the latter, the more conical pronotal horns are longer, placed nearer the midline and diverge strongly. In the females, the clypeal carina of
|
||
<taxonomicName id="C9FF4D30FF93FFFD23774D5E240DFD1C" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[1143,1304,619,645]" class="Insecta" family="Scarabaeidae" genus="Catharsius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="284" phylum="Arthropoda" rank="species" species="tricornutus">
|
||
<emphasis id="3C8BEAA1FF93FFFD23774D5E240DFD1C" box="[1143,1304,619,645]" italics="true" pageId="4" pageNumber="284">C. tricornutus</emphasis>
|
||
</taxonomicName>
|
||
is strongly bifurcate medially giving it a quadridentate appearance in frontal view, and the pronotal carina is straight, weakly produced and shorter than the clypeal carina.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="46E56538FF93FFFD27C74DE32527FD69" box="[199,1074,726,753]" pageId="4" pageNumber="284" type="distribution">
|
||
<paragraph id="0E4036B3FF93FFFD27C74DE32527FD69" blockId="4.[151,1437,151,753]" box="[199,1074,726,753]" pageId="4" pageNumber="284">
|
||
<emphasis id="3C8BEAA1FF93FFFD27C74DE32074FD68" bold="true" box="[199,353,726,752]" pageId="4" pageNumber="284">Distribution.</emphasis>
|
||
<collectingCountry id="76E87623FF93FFFD26674DE220EEFD69" box="[359,507,727,753]" name="Mozambique" pageId="4" pageNumber="284">Mozambique</collectingCountry>
|
||
and
|
||
<collectingCountry id="76E87623FF93FFFD25324DE223D5FD69" box="[562,704,727,753]" name="South Africa" pageId="4" pageNumber="284">South Africa</collectingCountry>
|
||
(
|
||
<emphasis id="3C8BEAA1FF93FFFD25D04DE223EDFD68" box="[720,760,727,752]" italics="true" pageId="4" pageNumber="284">e.g.</emphasis>
|
||
,
|
||
<bibRefCitation id="6A6E4B42FF93FFFD24064DE22533FD69" author="Van Rensburg, B. J. & McGeoch, M. A. & Chown, S. L. & Van Jaarsveld, A. S." box="[774,1062,727,753]" pageId="4" pageNumber="284" pagination="145 - 153" refId="ref5196" refString="Van Rensburg, B. J., McGeoch, M. A., Chown, S. L. & Van Jaarsveld, A. S. (1999) Conservation of heterogeneity among dung beetles in the Maputaland Centre of Endemism, South Africa. Biological Conservation, 88 (2), 145 - 153. https: // doi. org / 10.1016 / S 0006 - 3207 (98) 00109 - 8" type="journal article" year="1999">
|
||
Van Rensburg
|
||
<emphasis id="3C8BEAA1FF93FFFD24AB4DE222F6FD69" box="[939,995,727,753]" italics="true" pageId="4" pageNumber="284">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |