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<document id="6DAB86B0A64BF7DE4473E7ABD20C22FD" ID-DOI="10.26107/RBZ-2024-0017" ID-ISSN="2345-7600" ID-ZooBank="6CE5E7E8-2138-40F8-A503-4DE964EFAF94" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1723050159934" checkinUser="felipe" docAuthor="Páll-Gergely, Barna, , András Hunyadi, Sutcharit, &amp; Chirasak" docDate="2024" docId="87386C2A13746260FEDE15A85021FE47" docLanguage="en" docName="RafflesBZool.72.203-213.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Diaphera parini Páll-Gergely, , Sutcharit &amp; Chirasak, 2024, new species" docType="treatment" docVersion="1" lastPageNumber="207" masterDocId="7B01145213706264FFFA15145429FFF2" masterDocTitle="New and little-known Diapheridae of Cambodia and Thailand (Gastropoda: Stylommatophora: Streptaxoidea)" masterLastPageNumber="213" masterPageNumber="203" pageNumber="207" updateTime="1723053732350" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BCD60D6DA499D6529770133EB20CF5F4">New and little-known Diapheridae of Cambodia and Thailand (Gastropoda: Stylommatophora: Streptaxoidea)</mods:title>
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<mods:namePart id="AE4024AEC5352644E42B825BA129D395">Páll-Gergely, Barna</mods:namePart>
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<mods:namePart id="BAE36786F09B939D38C92BD962D8D195">, András Hunyadi</mods:namePart>
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<mods:namePart id="693E40FEF4368F510A5D4DD843578EE6">Sutcharit,</mods:namePart>
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<mods:roleTerm id="6000DFEEE76EAF30CCD544C9034156C6">Author</mods:roleTerm>
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<mods:namePart id="E05AF2392EF54545B7444C2F0AE3162F">Chirasak</mods:namePart>
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<mods:typeOfResource id="3EF5D6476CB339E33E2E5244C8691211">text</mods:typeOfResource>
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<mods:title id="018E9762E61E0C986EC16768EE4F62D2">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="30E2E7EB1120D0FC05A7878721B5EA9B">2024</mods:date>
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<mods:number id="19C6ECBCEB45779A6F228A2843EABD93">2024-07-26</mods:number>
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<mods:identifier id="93F3C96519845078F61FD760CE74945D" type="DOI">10.26107/RBZ-2024-0017</mods:identifier>
<mods:identifier id="3A52B8F3E5B24F11B803BA79BF5BFDDD" type="ISSN">2345-7600</mods:identifier>
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<treatment id="87386C2A13746260FEDE15A85021FE47" LSID="urn:lsid:plazi:treatment:87386C2A13746260FEDE15A85021FE47" httpUri="http://treatment.plazi.org/id/87386C2A13746260FEDE15A85021FE47" lastPageNumber="207" pageId="4" pageNumber="207">
<subSubSection id="478B8EB713746260FEDE15A85645FF27" box="[292,620,188,213]" pageId="4" pageNumber="207" type="nomenclature">
<paragraph id="0F2EDD3C13746260FEDE15A85645FF27" blockId="4.[292,620,188,245]" box="[292,620,188,213]" pageId="4" pageNumber="207">
<heading id="54666A5013746260FEDE15A85645FF27" bold="true" box="[292,620,188,213]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="207" reason="2">
<emphasis id="3DE5012E13746260FEDE15A85645FF27" bold="true" box="[292,620,188,213]" pageId="4" pageNumber="207">
<taxonomicName id="C891A6BF13746260FEDE15A855FEFF27" authorityName="Páll-Gergely &amp; &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[292,471,188,213]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="parini" status="sp. nov.">
<emphasis id="3DE5012E13746260FEDE15A855FEFF27" bold="true" box="[292,471,188,213]" italics="true" pageId="4" pageNumber="207">Diaphera parini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="26D6BC5513746260FE1C15A95645FF27" box="[486,620,189,213]" pageId="4" pageNumber="207" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FE7915C95625FF07" box="[387,524,221,245]" pageId="4" pageNumber="207" type="description">
<paragraph id="0F2EDD3C13746260FE7915C95625FF07" blockId="4.[292,620,188,245]" box="[387,524,221,245]" pageId="4" pageNumber="207">
(
<figureCitation id="97AAC1B913746260FE7115C955F1FF07" box="[395,472,221,245]" captionStart="Fig" captionStartId="3.[113,148,1964,1986]" captionTargetBox="[113,1451,260,1948]" captionTargetId="figure-10@3.[113,1451,260,1948]" captionTargetPageId="3" captionText="Fig. 2. Diaphera parini, new species, from Rayong Province, Thailand. A, holotype CUMZ 14213 with three enlarged apertures. B, paratype coll. HA from the type locality. Abbreviations: pal: parieto-angular lamella, c: columellar lamella, lpt: lower palatal tooth, upt: upper palatal tooth." pageId="4" pageNumber="207">
Figs.
<geoCoordinate id="6AA5BBFB13746260FE3315C955F1FF07" box="[457,472,221,245]" degrees="2, 6" direction="east" orientation="longitude" pageId="4" pageNumber="207" precision="5555" value="2.6">2</geoCoordinate>
</figureCitation>
,
<figureCitation id="97AAC1B913746260FE1C15C9562CFF07" box="[486,517,221,245]" captionStart="Fig" captionStartId="8.[136,171,1556,1578]" captionTargetBox="[136,1474,189,1540]" captionTargetId="figure-8@8.[136,1474,189,1540]" captionTargetPageId="8" captionText="Fig. 6. A, distribution map and BE, synoptic view of four Diaphera examined in this study. B, Diaphera saurini Benthem Jutting, 1962, C, Diaphera prima Panha, 2010, D, Diaphera pongrati, new species, and E, Diaphera parini, new species." pageId="4" pageNumber="207">6E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FF721409572EFDE7" pageId="4" pageNumber="207" type="materials_examined">
<paragraph id="0F2EDD3C13746260FF721409551BFEC7" blockId="4.[135,776,285,533]" box="[136,306,285,309]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FF721409551BFEC7" bold="true" box="[136,306,285,309]" pageId="4" pageNumber="207">Type material.</emphasis>
</paragraph>
<paragraph id="0F2EDD3C13746260FF72142955C7FE27" blockId="4.[135,776,285,533]" pageId="4" pageNumber="207">
<materialsCitation id="BFF9D76113746260FF72142955C7FE27" collectingDate="2023-03-09" collectionCode="CUMZ" collectorName="A. Hunyadi" country="Thailand" elevation="70" latitude="13.037884" location="Wat Tham Khao Bot" longLatPrecision="1" longitude="101.63533" municipality="Khao Chamao District" pageId="4" pageNumber="207" specimenCode="CUMZ 14213" specimenCount="1" stateProvince="Rayong" typeStatus="holotype">
<emphasis id="3DE5012E13746260FF72142954DEFEA7" bold="true" box="[136,247,317,341]" pageId="4" pageNumber="207">
<typeStatus id="D02A639E13746260FF72142954DBFEA7" box="[136,242,317,341]" pageId="4" pageNumber="207" type="holotype">Holotype</typeStatus>
.
</emphasis>
1 shell (
<figureCitation id="97AAC1B913746260FEAF14295585FEA7" box="[341,428,317,341]" captionStart="Fig" captionStartId="3.[113,148,1964,1986]" captionTargetBox="[113,1451,260,1948]" captionTargetId="figure-10@3.[113,1451,260,1948]" captionTargetPageId="3" captionText="Fig. 2. Diaphera parini, new species, from Rayong Province, Thailand. A, holotype CUMZ 14213 with three enlarged apertures. B, paratype coll. HA from the type locality. Abbreviations: pal: parieto-angular lamella, c: columellar lamella, lpt: lower palatal tooth, upt: upper palatal tooth." pageId="4" pageNumber="207">Fig. 2A</figureCitation>
; SH =
<quantity id="C86970D913746260FE0314295667FEA6" box="[505,590,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="207" unit="mm" value="3.2">3.2 mm</quantity>
, SW =
<quantity id="C86970D913746260FD5A1429572AFEA6" box="[672,771,317,341]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="4" pageNumber="207" unit="mm" value="1.21">1.21 mm</quantity>
, AH:
<quantity id="C86970D913746260FF3814495500FE86" box="[194,297,349,373]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="207" unit="mm" value="0.81">0.81 mm</quantity>
, AW:
<quantity id="C86970D913746260FE8C144955E6FE87" box="[374,463,349,373]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="207" unit="mm" value="0.8">0.8 mm</quantity>
) (
<specimenCode id="5F37754713746260FE1D144956A3FE87" box="[487,650,349,373]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="4" pageNumber="207">CUMZ 14213</specimenCode>
),
<collectingCountry id="77869DAC13746260FD5A1449572AFE87" box="[672,771,349,373]" name="Thailand" pageId="4" pageNumber="207">Thailand</collectingCountry>
,
<collectingRegion id="CD5513DE13746260FF721469556CFE67" box="[136,325,381,405]" country="Thailand" name="Rayong" pageId="4" pageNumber="207">Rayong Province</collectingRegion>
,
<collectingMunicipality id="EF4A474613746260FEAA1469566FFE67" box="[336,582,381,405]" pageId="4" pageNumber="207">Khao Chamao District</collectingMunicipality>
, rock wall behind
<location id="0A4E8BE713746260FF7314895543FE46" LSID="urn:lsid:plazi:treatment:87386C2A13746260FEDE15A85021FE47:0A4E8BE713746260FF7314895543FE46" box="[137,362,413,437]" country="Thailand" latitude="13.037884" longLatPrecision="1" longitude="101.63533" municipality="Khao Chamao District" name="Wat Tham Khao Bot" pageId="4" pageNumber="207" stateProvince="Rayong">Wat Tham Khao Bot</location>
,
<elevation id="84BC3A0F13746260FE8E148955F9FE47" box="[372,464,413,437]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="207" unit="m" value="70.0">
<quantity id="C86970D913746260FE8E14895583FE46" box="[372,426,413,437]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="207" unit="m" value="70.0">70 m</quantity>
asl
</elevation>
,
<geoCoordinate id="6AA5BBFB13746260FE21148F564CFE46" box="[475,613,411,436]" degrees="13" direction="north" minutes="02.273" orientation="latitude" pageId="4" pageNumber="207" precision="1" value="13.037884">13°02.273N</geoCoordinate>
,
<geoCoordinate id="6AA5BBFB13746260FD95148F572DFE46" box="[623,772,411,436]" degrees="101" direction="east" minutes="38.120" orientation="longitude" pageId="4" pageNumber="207" precision="1" value="101.63533">101°38.120E</geoCoordinate>
, coll.
<collectorName id="A264B8EA13746260FF4514A95568FE27" box="[191,321,445,469]" pageId="4" pageNumber="207">A. Hunyadi</collectorName>
,
<date id="7B2FFBFC13746260FEB714A955C3FE27" box="[333,490,445,469]" pageId="4" pageNumber="207" value="2023-03-09">
<collectingDate id="6B6B021413746260FEB714A955C3FE27" box="[333,490,445,469]" pageId="4" pageNumber="207" value="2023-03-09">9 March 2023</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="0F2EDD3C13746260FF7214C9572EFDE7" blockId="4.[135,776,285,533]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FF7214C95528FE07" bold="true" box="[136,257,477,501]" pageId="4" pageNumber="207">
<typeStatus id="D02A639E13746260FF7214C954D5FE07" box="[136,252,477,501]" pageId="4" pageNumber="207" type="paratype">Paratypes</typeStatus>
.
</emphasis>
1 shell (
<figureCitation id="97AAC1B913746260FEA314C95583FE07" box="[345,426,477,501]" captionStart="Fig" captionStartId="3.[113,148,1964,1986]" captionTargetBox="[113,1451,260,1948]" captionTargetId="figure-10@3.[113,1451,260,1948]" captionTargetPageId="3" captionText="Fig. 2. Diaphera parini, new species, from Rayong Province, Thailand. A, holotype CUMZ 14213 with three enlarged apertures. B, paratype coll. HA from the type locality. Abbreviations: pal: parieto-angular lamella, c: columellar lamella, lpt: lower palatal tooth, upt: upper palatal tooth." pageId="4" pageNumber="207">Fig. 2B</figureCitation>
) (coll. HA), same information as
<typeStatus id="D02A639E13746260FF7D14E954C1FDE7" box="[135,232,509,533]" pageId="4" pageNumber="207" type="holotype">holotype</typeStatus>
; 2 shells (coll. HA), same information as
<typeStatus id="D02A639E13746260FD5814E9572AFDE7" box="[674,771,509,533]" pageId="4" pageNumber="207" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FF721729572DFCA7" pageId="4" pageNumber="207" type="diagnosis">
<paragraph id="0F2EDD3C13746260FF721729572DFCA7" blockId="4.[135,777,573,853]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FF721729552AFDA7" bold="true" box="[136,259,573,597]" pageId="4" pageNumber="207">Diagnosis.</emphasis>
The smallest
<taxonomicName id="C891A6BF13746260FE541729565EFDA7" box="[430,631,573,597]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FE5417295633FDA7" box="[430,538,573,597]" italics="true" pageId="4" pageNumber="207">Diaphera</emphasis>
species
</taxonomicName>
in
<collectingCountry id="77869DAC13746260FD591729572FFDA7" box="[675,774,573,597]" name="Thailand" pageId="4" pageNumber="207">Thailand</collectingCountry>
and
<collectingCountry id="77869DAC13746260FF461749551AFD87" box="[188,307,605,629]" name="Cambodia" pageId="4" pageNumber="207">Cambodia</collectingCountry>
, with an ovoid shell and five apertural barriers (a very strong parieto-angular lamella, blunt upper palatal tooth, a more pointed lower palatal tooth, slightly elongated columellar lamella, and a low, blunt basal tooth). The sympatric
<taxonomicName id="C891A6BF13746260FED417CA55B8FD07" authorityName="Panha" authorityYear="2010" box="[302,401,734,757]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="prima">
<emphasis id="3DE5012E13746260FED417CA55B8FD07" box="[302,401,734,757]" italics="true" pageId="4" pageNumber="207">D. prima</emphasis>
</taxonomicName>
is much larger (
<figureCitation id="97AAC1B913746260FDBF17C95693FD07" box="[581,698,733,757]" captionStart="Fig" captionStartId="8.[136,171,1556,1578]" captionTargetBox="[136,1474,189,1540]" captionTargetId="figure-8@8.[136,1474,189,1540]" captionTargetPageId="8" captionText="Fig. 6. A, distribution map and BE, synoptic view of four Diaphera examined in this study. B, Diaphera saurini Benthem Jutting, 1962, C, Diaphera prima Panha, 2010, D, Diaphera pongrati, new species, and E, Diaphera parini, new species." pageId="4" pageNumber="207">Fig. 6C, E</figureCitation>
), more spindle-shaped, has a different sculpture (strong riblets along the suture but smooth mid-whorls), and a longer detached portion (
<taxonomicName id="C891A6BF13746260FF1C1629550CFCA7" baseAuthorityName="Mollendorff" baseAuthorityYear="1887" box="[230,293,829,853]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="tuba">“tuba</taxonomicName>
”). See also under
<taxonomicName id="C891A6BF13746260FE15162A5642FCA7" authorityName="Páll-Gergely &amp; &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[495,619,830,853]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="pongrati" status="new species">
<emphasis id="3DE5012E13746260FE15162A5642FCA7" box="[495,619,830,853]" italics="true" pageId="4" pageNumber="207">D. pongrati</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="26D6BC5513746260FD80162956D6FCA7" box="[634,767,829,853]" pageId="4" pageNumber="207" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FF72166950C3FF07" pageId="4" pageNumber="207" type="description">
<paragraph id="0F2EDD3C13746260FF721669559EF7E7" blockId="4.[135,778,893,2069]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FF721669553FFC67" bold="true" box="[136,278,893,917]" pageId="4" pageNumber="207">Description.</emphasis>
Shell white, opaque to slightly translucent. Spire-whorls elongate ovoid with a deep suture; apex blunt; last and penultimate whorls of comparable width. Whorls 6.5; protoconch-teleoconch boundary not clearly visible; all whorls strongly convex (bulging); first juvenile peristome developed at ca. 2 whorls; last juvenile peristome present ca. 1.52 whorls behind adult peristome. Protoconch glossy, smooth. Teleoconch moderately glossy, with very fine, low, inconspicuous ribs; last ca. half whorl, but mostly the detached part (
<taxonomicName id="C891A6BF13746260FEA5118955B7FB47" baseAuthorityName="Mollendorff" baseAuthorityYear="1887" box="[351,414,1181,1205]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="tuba">“tuba</taxonomicName>
”) with somewhat stronger ribs, area around the umbilicus also with stronger ribs, not reaching the middle of the body whorl; last ca. 0.25 whorls detached, detached part short, thick, aperture not protruding beyond spire-whorl plane. Detached portion (
<taxonomicName id="C891A6BF13746260FD9410095682FAC7" baseAuthorityName="Mollendorff" baseAuthorityYear="1887" box="[622,683,1309,1333]" class="Gastropoda" family="Diapheridae" genus="Diaphera" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="207" phylum="Mollusca" rank="species" species="tuba">“tuba</taxonomicName>
”) slopes downward at an angle of about 40°. Upper suture continues to back side of sinulus without forming a keel, it bears a shallow groove on inner side, dividing the upper part of the detached portion in the middle; outer (lateral) side of detached portion without groove, but with a slight depression corresponding with the lower palatal tooth. Lower suture continues towards the peristome, forming a very blunt periumbilical keel, with a short, shallow depression on the umbilical side corresponding with the columellar lamella. Aperture axis slightly oblique to shell axis; aperture ovoid, nearly rectangular, with a narrow, strongly isolated sinulus. Sinulus opens slightly upwards. Parieto-angular lamella very strongly developed, extremely high, long, strongly oblique; outer tip slightly protruding; the entire lamella seems to be turning slightly towards columellar side. Upper palatal tooth blunt, but strong, lower palatal tooth situated much deeper and lower in position, more pointed than upper palatal tooth. Columellar lamella slightly elongated, situated deep inside the aperture, its position corresponds with the end of the groove on the umbilical side of the tuba. Basal tooth low, blunt, only slightly indicated. Peristome thickened, strongly expanded (except for the sinulus area), its margin slightly reflected. Umbilicus closed.
</paragraph>
<paragraph id="0F2EDD3C13746260FCB915A950C3FF07" blockId="4.[835,1474,189,246]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FCB915A95070FF27" bold="true" box="[835,1113,189,213]" pageId="4" pageNumber="207">Measurements (in mm).</emphasis>
SH = 3.13.4, SW = 1.211.27, AH: 0.740.81, AW: 0.750.8 (n = 3).
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FCB91409515BFEA7" pageId="4" pageNumber="207" type="etymology">
<paragraph id="0F2EDD3C13746260FCB91409515BFEA7" blockId="4.[835,1474,285,341]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FCB9140957ECFEC7" bold="true" box="[835,965,285,309]" pageId="4" pageNumber="207">Etymology.</emphasis>
This new species is named after and dedicated to our friend Parin Jirapatrasilp, Thai malacologist.
</paragraph>
</subSubSection>
<subSubSection id="478B8EB713746260FCB914695021FE47" pageId="4" pageNumber="207" type="distribution">
<paragraph id="0F2EDD3C13746260FCB914695021FE47" blockId="4.[835,1474,381,437]" pageId="4" pageNumber="207">
<emphasis id="3DE5012E13746260FCB9146957F3FE67" bold="true" box="[835,986,381,405]" pageId="4" pageNumber="207">Distribution.</emphasis>
This species is known only from the
<typeStatus id="D02A639E13746260FA69146A51EBFE64" box="[1427,1474,382,406]" pageId="4" pageNumber="207">type</typeStatus>
locality (
<figureCitation id="97AAC1B913746260FC5E148957D5FE47" box="[932,1020,413,437]" captionStart="Fig" captionStartId="8.[136,171,1556,1578]" captionTargetBox="[136,1474,189,1540]" captionTargetId="figure-8@8.[136,1474,189,1540]" captionTargetPageId="8" captionText="Fig. 6. A, distribution map and BE, synoptic view of four Diaphera examined in this study. B, Diaphera saurini Benthem Jutting, 1962, C, Diaphera prima Panha, 2010, D, Diaphera pongrati, new species, and E, Diaphera parini, new species." pageId="4" pageNumber="207">Fig. 6A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>