171 lines
27 KiB
XML
171 lines
27 KiB
XML
<document id="2D1E829435765B9EA263C53C4026D7CF" ID-CLB-Dataset="138998" ID-DOI="10.11646/zootaxa.5182.5.1" ID-GBIF-Dataset="1b35fb1e-b051-429f-87fe-079aac9196ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="7059565" ID-ZooBank="E70EB74A-DC7E-482D-AE4D-D58FB3CD4F5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1662620157126" checkinUser="plazi" docAuthor="Anilkumar, Pooja Avinipully, Wesener, Thomas & Moritz, Leif" docDate="2022" docId="0A3287A5B908E81D6DC1FB46FC055A1D" docLanguage="en" docName="zootaxa.5182.5.1.pdf" docOrigin="Zootaxa 5182 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theratta Anilkumar & Wesener & Moritz 2022, n. gen." docType="treatment" docVersion="7" lastPageNumber="406" masterDocId="F60BFFDDB90DE8186D56FFAEFF965D7B" masterDocTitle="First record of the order Polyzoniida from the Indian subcontinent with an integrative description of a new genus (Diplopoda, Colobognatha, Siphonotidae)" masterLastPageNumber="428" masterPageNumber="401" pageNumber="406" updateTime="1699378897687" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F73BC0067AC25C86BF214D9A03380369">
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<mods:title id="0F955104339176631C1320052CA31699">First record of the order Polyzoniida from the Indian subcontinent with an integrative description of a new genus (Diplopoda, Colobognatha, Siphonotidae)</mods:title>
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<mods:name id="F7DA35B14018E391C550285751B3FA4B" type="personal">
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<mods:roleTerm id="3A70D8266EF9D0D304FD7697449E46A7">Author</mods:roleTerm>
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<mods:namePart id="FE23E54516B29B1A1EF79620FE0AA301">Anilkumar, Pooja Avinipully</mods:namePart>
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<mods:affiliation id="DE05C04B3D9CB5F18C100B8D3174F2F0">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Bonn, Germany.</mods:affiliation>
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<mods:name id="BC7DD04E9A2B6A7845B267A1D62172B0" type="personal">
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<mods:roleTerm id="6A4749E61DB9FEFF791E4113437BFFF4">Author</mods:roleTerm>
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<mods:namePart id="227804A3E94A055E1B548D57D718251C">Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier id="FDFDD100AF9B2B9324C5F72C2775BA84" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
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<mods:affiliation id="4F159D8C6D0B8FB35D0640D685B698D2">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Bonn, Germany. & T. Wesener @ leibniz-lib. de; https: // orcid. org / 0000 - 0002 - 2028 - 3541</mods:affiliation>
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<mods:nameIdentifier id="A3FC174AC5D8667A307B7F34287F1746" type="email">esener@leibniz-lib.de</mods:nameIdentifier>
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<mods:namePart id="AEB0103932E77BCC96DC0093D6102F22">Moritz, Leif</mods:namePart>
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<mods:nameIdentifier id="C5247CE2EFC21620E54818FD4A2F1FE7" type="ORCID">0000-0002-6028-5189</mods:nameIdentifier>
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<mods:affiliation id="8CA04960625E7AC625430AEEE2500419">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Bonn, Germany. & Institute of Evolutionary Biology and Animal Ecology, University of Bonn, Bonn, Germany. & moritz. leif @ gmail. com; https: // orcid. org / 0000 - 0002 - 6028 - 5189</mods:affiliation>
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<mods:nameIdentifier id="AC25A6EFBEAE75A781441D92F6EEAA6D" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:title id="FB0FAE3048FBD96BD71E0661BA438634">Zootaxa</mods:title>
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<mods:part id="531939F27E5DBFB804D4C1D5269CCBEF">
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<mods:date id="4166CC2D3FB4F63A6F7B3A363DC9DA40">2022</mods:date>
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<mods:number id="52E535243C79165C7210C7E3436759F0">2022-09-08</mods:number>
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<mods:number id="E8F084214ED85FC68D2A24B9FE694674">5182</mods:number>
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<treatment id="0A3287A5B908E81D6DC1FB46FC055A1D" ID-DOI="http://doi.org/10.5281/zenodo.7061880" ID-GBIF-Taxon="199973859" ID-Zenodo-Dep="7061880" LSID="urn:lsid:plazi:treatment:0A3287A5B908E81D6DC1FB46FC055A1D" httpUri="http://treatment.plazi.org/id/0A3287A5B908E81D6DC1FB46FC055A1D" lastPageNumber="406" pageId="5" pageNumber="406">
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<subSubSection id="CA816538B908E81D6DC1FB46FCA25879" box="[151,820,1256,1283]" pageId="5" pageNumber="406" type="nomenclature">
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<paragraph id="822436B3B908E81D6DC1FB46FCA25879" blockId="5.[151,820,1256,1283]" box="[151,820,1256,1283]" pageId="5" pageNumber="406">
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<heading id="D96C81DFB908E81D6DC1FB46FCA25879" bold="true" box="[151,820,1256,1283]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
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<emphasis id="B0EFEAA1B908E81D6DC1FB46FCA25879" bold="true" box="[151,820,1256,1283]" pageId="5" pageNumber="406">
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Genus
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<taxonomicName id="459B4D30B908E81D6DB8FB46FEC05878" ID-CoL="18012452-f6a0-48f9-acbf-2dfd643e4f4e" authority="Anilkumar, Wesener & Moritz" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[238,342,1256,1283]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B0EFEAA1B908E81D6DB8FB46FEC05878" bold="true" box="[238,342,1256,1283]" italics="true" pageId="5" pageNumber="406">Theratta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="ABDC57DAB908E81D6C08FB47FE265878" box="[350,432,1257,1283]" pageId="5" pageNumber="406" rank="genus">n. gen.</taxonomicNameLabel>
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Anilkumar, Wesener & Moritz
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CA816538B908E81D6DC1FA9FFBBA58E8" pageId="5" pageNumber="406" type="type_taxon">
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<paragraph id="822436B3B908E81D6DC1FA9FFC615831" blockId="5.[151,1015,1328,1355]" box="[151,1015,1328,1355]" pageId="5" pageNumber="406">
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<emphasis id="B0EFEAA1B908E81D6DC1FA9FFEAA5831" bold="true" box="[151,316,1328,1355]" pageId="5" pageNumber="406">
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<typeStatus id="5D208811B908E81D6DC1FA9FFF425830" box="[151,212,1329,1355]" pageId="5" pageNumber="406">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="459B4D30B908E81D6C15FA9EFDBC5831" ID-CoL="eaf450b3-4517-43c9-8bb3-cea8f566fbbf" authority="Anilkumar & Wesener & Moritz, 2022" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[323,554,1328,1354]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="mannavan" status="sp. nov.">
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<emphasis id="B0EFEAA1B908E81D6C15FA9EFDBC5831" box="[323,554,1328,1354]" italics="true" pageId="5" pageNumber="406">Theratta mannavan</emphasis>
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</taxonomicName>
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<emphasis id="B0EFEAA1B908E81D6F67FA9FFC615831" bold="true" box="[561,1015,1328,1355]" pageId="5" pageNumber="406">
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<taxonomicNameLabel id="ABDC57DAB908E81D6F67FA9FFDE25830" box="[561,628,1329,1355]" pageId="5" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
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Anilkumar, Wesener & Moritz
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</emphasis>
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</paragraph>
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<paragraph id="822436B3B908E81D6DC1FAD6FBBA58E8" blockId="5.[151,1068,1400,1427]" box="[151,1068,1400,1427]" pageId="5" pageNumber="406">
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<emphasis id="B0EFEAA1B908E81D6DC1FAD6FE2A58E9" bold="true" box="[151,444,1400,1426]" pageId="5" pageNumber="406">Other species included:</emphasis>
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<taxonomicName id="459B4D30B908E81D6C95FAD6FD2258E9" ID-CoL="f5f95154-c8f2-410b-bbe0-6cd6e148afbf" authority="Anilkumar & Wesener & Moritz, 2022" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[451,692,1400,1426]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="eravikulam" status="sp. nov.">
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<emphasis id="B0EFEAA1B908E81D6C95FAD6FD2258E9" box="[451,692,1400,1426]" italics="true" pageId="5" pageNumber="406">Theratta eravikulam</emphasis>
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</taxonomicName>
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<emphasis id="B0EFEAA1B908E81D6FEDFAD7FD6B58E8" bold="true" box="[699,765,1401,1427]" pageId="5" pageNumber="406">
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<taxonomicNameLabel id="ABDC57DAB908E81D6FEDFAD7FD6B58E8" box="[699,765,1401,1427]" pageId="5" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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and
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<taxonomicName id="459B4D30B908E81D6E60FAD6FC7458E9" ID-CoL="c2b3d43d-9f83-4fb8-a3e8-cb53d4b01603" authority="Anilkumar & Wesener & Moritz, 2022" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[822,994,1400,1426]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="shola" status="sp. nov.">
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<emphasis id="B0EFEAA1B908E81D6E60FAD6FC7458E9" box="[822,994,1400,1426]" italics="true" pageId="5" pageNumber="406">Theratta shola</emphasis>
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</taxonomicName>
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<emphasis id="B0EFEAA1B908E81D6EBFFAD7FBBA58E8" bold="true" box="[1001,1068,1401,1427]" pageId="5" pageNumber="406">
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<taxonomicNameLabel id="ABDC57DAB908E81D6EBFFAD7FBBA58E8" box="[1001,1068,1401,1427]" pageId="5" pageNumber="406" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="CA816538B908E81D6DC1FA6EFEE85A64" pageId="5" pageNumber="406" type="diagnosis">
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<paragraph id="822436B3B908E81D6DC1FA6EFEE85A64" blockId="5.[151,1437,1472,1895]" pageId="5" pageNumber="406">
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<emphasis id="B0EFEAA1B908E81D6DC1FA6EFE8458A1" bold="true" box="[151,274,1472,1498]" pageId="5" pageNumber="406">Diagnosis.</emphasis>
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Habitus, size and coloration of
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<taxonomicName id="459B4D30B908E81D6FDEFA6EFCD258A1" box="[648,836,1472,1498]" pageId="5" pageNumber="406">
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<emphasis id="B0EFEAA1B908E81D6FDEFA6EFD7E58A1" box="[648,744,1472,1498]" italics="true" pageId="5" pageNumber="406">Theratta</emphasis>
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species
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</taxonomicName>
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resemble those of the widespread introduced tramp species
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<taxonomicName id="459B4D30B908E81D6DA6FA4AFE585885" box="[240,462,1508,1534]" class="Diplopoda" family="Siphonotidae" genus="Rhinotus" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="purpureus">
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<emphasis id="B0EFEAA1B908E81D6DA6FA4AFE585885" box="[240,462,1508,1534]" italics="true" pageId="5" pageNumber="406">Rhinotus purpureus</emphasis>
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</taxonomicName>
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. Distinguished from the three other genera of the tribe
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<taxonomicName id="459B4D30B908E81D6914FA4AFB3A5885" authorityName="Hoffman" authorityYear="1977" box="[1090,1196,1508,1534]" class="Diplopoda" family="Siphonotidae" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="tribe" tribe="Rhinotini">Rhinotini</taxonomicName>
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by the bifurcation of the podomere II of anterior gonopod into a mesal process on the posterior side, with a furrow between the mesal process and the main part of the podomere II (
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<figureCitation id="1AA02A36B908E81D6FC2F982FC865B3C" box="[660,784,1580,1607]" captionStart="FIGURE 5" captionStartId="9.[151,250,1716,1740]" captionTargetBox="[201,1386,159,1692]" captionTargetId="figure-17@9.[201,1386,159,1692]" captionTargetPageId="9" captionText="FIGURE 5. Theratta mannavan n. sp., SEM. A, E, F: M holotype (ZFMK-MYR 9482B1). B: M paratype (ZFMK-MYR 9483A). C: F paratype (ZFMK-MYR 9482C). D: M paratype (ZFMK-MYR 9482B2). A: Telson, ventral view. B: Telson, lateral view. C: Female gonopore, ventral view. D: Gonopods, lateral view. E: Gonopods, posterior view. F: Gonopods, posterio-lateral view. Abbreviations: av = anal valve; cx = coxa; cxp = coxal pouch; fe = femur; go = gonopore; L = lamellae; le = leg; mp = mesal process; oz = ozopore; pbr = penultimate body ring; pf = prefemur; pl = pleurite; po = postfemur; st = sternite; ta = tarsus; te = telson; ti = tibia; ve = external valve of vulva; vi = internal valve of vulva. I–IV indicates the number of podomeres of gonopods. The black arrow on (E, F) indicates the remnant of podomere divisions of gonopods and the white arrow on (E, F) indicates the distal process present on the mesal process of the anterior gonopod. Scale bars: A = 200 µm; B, C = 100 µm; D–F = 20 µm." figureDoi="http://doi.org/10.5281/zenodo.7059581" httpUri="https://zenodo.org/record/7059581/files/figure.png" pageId="5" pageNumber="406">Figs 5D–F</figureCitation>
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); modification of the anterior side of anterior gonopod into a lamellar process (
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<figureCitation id="1AA02A36B908E81D6C25F9FEFE5A5B11" box="[371,460,1616,1642]" captionStart="FIGURE 6" captionStartId="10.[151,250,1784,1808]" captionTargetBox="[225,1362,159,1763]" captionTargetId="figure-27@10.[225,1362,159,1763]" captionTargetPageId="10" captionText="FIGURE 6. Theratta mannavan n. sp., SEM. A, D, F, J: M paratype (ZFMK-MYR 9483A). B, E: M holotype (ZFMK-MYR 9482B2). C: M paratype (ZFMK-MYR 11393). G, H, I: M paratype (ZFMK-MYR 9482B1). A: Gonopods, dorsal view. B, C: Anterior gonopod, anterior view. D: Anterior gonopod, posterior view. E: Anterior gonopod, mesal view. F: Body teeth. G: Teeth on the mesal process of the anterior gonopod. H: Feather like serrated setae. I: Posterior gonopod, posterior view. J: Posterior gonopod, anterior view. Abbreviations: L = lamellae; mp = mesal process; st = sternite. I–IV indicates the number of podomeres of gonopods. The white arrow on (I) indicates the remnant of podomere division on the posterior gonopod. Scale bars: A, C–E, I, J = 20 µm; B, H = 10 µm; F, G = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.7059583" httpUri="https://zenodo.org/record/7059583/files/figure.png" pageId="5" pageNumber="406">Fig. 6C</figureCitation>
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); absence of conical elevations on the sternite of the posterior gonopod (
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<figureCitation id="1AA02A36B908E81D6854F9FEFAF95B10" box="[1282,1391,1616,1643]" captionStart="FIGURE 6" captionStartId="10.[151,250,1784,1808]" captionTargetBox="[225,1362,159,1763]" captionTargetId="figure-27@10.[225,1362,159,1763]" captionTargetPageId="10" captionText="FIGURE 6. Theratta mannavan n. sp., SEM. A, D, F, J: M paratype (ZFMK-MYR 9483A). B, E: M holotype (ZFMK-MYR 9482B2). C: M paratype (ZFMK-MYR 11393). G, H, I: M paratype (ZFMK-MYR 9482B1). A: Gonopods, dorsal view. B, C: Anterior gonopod, anterior view. D: Anterior gonopod, posterior view. E: Anterior gonopod, mesal view. F: Body teeth. G: Teeth on the mesal process of the anterior gonopod. H: Feather like serrated setae. I: Posterior gonopod, posterior view. J: Posterior gonopod, anterior view. Abbreviations: L = lamellae; mp = mesal process; st = sternite. I–IV indicates the number of podomeres of gonopods. The white arrow on (I) indicates the remnant of podomere division on the posterior gonopod. Scale bars: A, C–E, I, J = 20 µm; B, H = 10 µm; F, G = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.7059583" httpUri="https://zenodo.org/record/7059583/files/figure.png" pageId="5" pageNumber="406">Figs 6I, J</figureCitation>
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), as found in the genera
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<taxonomicName id="459B4D30B908E81D6C21F9DAFE4D5BF5" authorityName="Cook" authorityYear="1896" box="[375,475,1652,1678]" class="Diplopoda" family="Siphonotidae" genus="Rhinotus" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
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<emphasis id="B0EFEAA1B908E81D6C21F9DAFE4D5BF5" box="[375,475,1652,1678]" italics="true" pageId="5" pageNumber="406">Rhinotus</emphasis>
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</taxonomicName>
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(see Wesener 2014) and
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<taxonomicName id="459B4D30B908E81D6FA2F9DAFC075BF5" authorityName="Attems" authorityYear="1931" box="[756,913,1652,1678]" class="Diplopoda" family="Siphonotidae" genus="Siphonoconus" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
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<emphasis id="B0EFEAA1B908E81D6FA2F9DAFC075BF5" box="[756,913,1652,1678]" italics="true" pageId="5" pageNumber="406">Siphonoconus</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="E60A4B42B908E81D6E9CF9DBFBC85BF4" author="Attems, C. G." box="[970,1118,1653,1679]" pageId="5" pageNumber="406" pagination="115 - 192" refId="ref13800" refString="Attems, C. G. (1931) Myriopoden von Java, Sumatra und Bali. In: Thienemann A. (Eds.), Archiv fur Hydrobiologie, E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, 8, 115 - 192." type="journal article" year="1931">Attems 1931</bibRefCitation>
|
||
). Eyes consisting of a single pair of ommatidia (
|
||
<figureCitation id="1AA02A36B908E81D6C25F936FE595BC8" box="[371,463,1688,1715]" captionStart="FIGURE 3" captionStartId="7.[151,250,1720,1744]" captionTargetBox="[249,1340,159,1697]" captionTargetId="figure-17@7.[248,1340,159,1697]" captionTargetPageId="7" captionText="FIGURE 3. Theratta mannavan n. sp., SEM. A, D, E: M paratype (ZFMK-MYR 9483A). B, C, F, G: M holotype (ZFMKMYR9482B1). H: M paratype (ZFMK-MYR 9482B2). A: Head, frontal view. B: Antenna. C: Sensilla basiconica on antennomeres 5 and 6. D: Apical disc with four sensory cones. E: Mouth parts, dorsal view. F: Mouth parts, ventral view. G: Anterior body, lateral view. H: Tergites, lateral view. Abbreviations: ac = apical cone; ag = anterior gonopod; an = antenna; br = body ring; co = collum; gn = gnathochilarium; le = leg; lb = labrum; ms = macrosetae; mz = metazonite; om = ommatidia; oz = ozopore; pg = posterior gonopod; pl = pleurite; pz = prozonite; sb = sensilla basiconica; sp = spiracle; st = sternite. I–VII indicates the number of antennomeres. The white arrow on (B) indicates the row of striae at the anterior margin of the antennomeres and on (H) indicates the lateral, ridge-like extension of the prozonite. Scale bars: A, G, H = 100 µm; B, D–F = 20 µm; C = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7059577" httpUri="https://zenodo.org/record/7059577/files/figure.png" pageId="5" pageNumber="406">Fig. 3A</figureCitation>
|
||
); head with a pair of macrosetae anterior to the ommatidia (
|
||
<figureCitation id="1AA02A36B908E81D692AF936FB4C5BC8" box="[1148,1242,1688,1715]" captionStart="FIGURE 2" captionStartId="6.[151,250,1927,1951]" captionTargetBox="[151,1436,645,1904]" captionTargetId="figure-232@6.[151,1436,633,1904]" captionTargetPageId="6" captionText="FIGURE 2. Theratta mannavan n. sp.. A, B: M paratype (ZFMK-MYR 9482A1). A: Habitus. B: Head, lateral view. C, D: Habitats, Mannavan Shola, Annamudi Shola National Park, Kerala, India. Abbreviations: an = antenna; co = collum; gp = gonopod; ms = macroseta; om = ommatidia." figureDoi="http://doi.org/10.5281/zenodo.7059573" httpUri="https://zenodo.org/record/7059573/files/figure.png" pageId="5" pageNumber="406">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="1AA02A36B908E81D69B0F937FA9A5BC8" box="[1254,1292,1689,1715]" captionStart="FIGURE 3" captionStartId="7.[151,250,1720,1744]" captionTargetBox="[249,1340,159,1697]" captionTargetId="figure-17@7.[248,1340,159,1697]" captionTargetPageId="7" captionText="FIGURE 3. Theratta mannavan n. sp., SEM. A, D, E: M paratype (ZFMK-MYR 9483A). B, C, F, G: M holotype (ZFMKMYR9482B1). H: M paratype (ZFMK-MYR 9482B2). A: Head, frontal view. B: Antenna. C: Sensilla basiconica on antennomeres 5 and 6. D: Apical disc with four sensory cones. E: Mouth parts, dorsal view. F: Mouth parts, ventral view. G: Anterior body, lateral view. H: Tergites, lateral view. Abbreviations: ac = apical cone; ag = anterior gonopod; an = antenna; br = body ring; co = collum; gn = gnathochilarium; le = leg; lb = labrum; ms = macrosetae; mz = metazonite; om = ommatidia; oz = ozopore; pg = posterior gonopod; pl = pleurite; pz = prozonite; sb = sensilla basiconica; sp = spiracle; st = sternite. I–VII indicates the number of antennomeres. The white arrow on (B) indicates the row of striae at the anterior margin of the antennomeres and on (H) indicates the lateral, ridge-like extension of the prozonite. Scale bars: A, G, H = 100 µm; B, D–F = 20 µm; C = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7059577" httpUri="https://zenodo.org/record/7059577/files/figure.png" pageId="5" pageNumber="406">3A</figureCitation>
|
||
); tarsal claw with a thin paronychium extending beyond the claw, curved and bulged distally (
|
||
<figureCitation id="1AA02A36B908E81D6976F912FBE05BAC" box="[1056,1142,1724,1751]" captionStart="FIGURE 4" captionStartId="8.[151,250,1644,1668]" captionTargetBox="[151,1436,160,1621]" captionTargetId="figure-27@8.[151,1436,159,1621]" captionTargetPageId="8" captionText="FIGURE 4. Theratta mannavan n. sp., SEM. A–D: M paratype (ZFMK-MYR 9483A). E, F: M holotype (ZFMK-MYR 9482B1). A: Tergites, dorsal view. B: First three leg pairs with gonopore, ventral view. C: Gonopore, posterior view. D: Detail of gonopore, ventral view. E: Leg. F: Paronychium. Abbreviations: cl = claw; cx = coxa; cxp = coxal pouch; fe = femur; le = leg; mz = metazonite; oz = ozopore; pe = penes; pf = prefemur; pn = paronychium; po = postfemur; pz = prozonite; sp = spiracle; st = sternite; ta = tarsus; ti = tibia; to = trochanter; vdo = vasa deferentia opening. The white arrow on (A) indicates the row of striae at the anterior margin of the metazonite. Scale bars: A = 200 µm; B, C, E = 20 µm; D, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7059579" httpUri="https://zenodo.org/record/7059579/files/figure.png" pageId="5" pageNumber="406">Fig. 4F</figureCitation>
|
||
); tergites with groove-like striae; anterior margin of metazonites with a distinct row of striae (
|
||
<figureCitation id="1AA02A36B908E81D6EDCF94EFC735B80" box="[906,997,1760,1787]" captionStart="FIGURE 4" captionStartId="8.[151,250,1644,1668]" captionTargetBox="[151,1436,160,1621]" captionTargetId="figure-27@8.[151,1436,159,1621]" captionTargetPageId="8" captionText="FIGURE 4. Theratta mannavan n. sp., SEM. A–D: M paratype (ZFMK-MYR 9483A). E, F: M holotype (ZFMK-MYR 9482B1). A: Tergites, dorsal view. B: First three leg pairs with gonopore, ventral view. C: Gonopore, posterior view. D: Detail of gonopore, ventral view. E: Leg. F: Paronychium. Abbreviations: cl = claw; cx = coxa; cxp = coxal pouch; fe = femur; le = leg; mz = metazonite; oz = ozopore; pe = penes; pf = prefemur; pn = paronychium; po = postfemur; pz = prozonite; sp = spiracle; st = sternite; ta = tarsus; ti = tibia; to = trochanter; vdo = vasa deferentia opening. The white arrow on (A) indicates the row of striae at the anterior margin of the metazonite. Scale bars: A = 200 µm; B, C, E = 20 µm; D, F = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7059579" httpUri="https://zenodo.org/record/7059579/files/figure.png" pageId="5" pageNumber="406">Fig. 4A</figureCitation>
|
||
); lateral margin of the pleurite covered with setae (
|
||
<figureCitation id="1AA02A36B908E81D6C41F8AAFEE75A64" box="[279,369,1796,1823]" captionStart="FIGURE 3" captionStartId="7.[151,250,1720,1744]" captionTargetBox="[249,1340,159,1697]" captionTargetId="figure-17@7.[248,1340,159,1697]" captionTargetPageId="7" captionText="FIGURE 3. Theratta mannavan n. sp., SEM. A, D, E: M paratype (ZFMK-MYR 9483A). B, C, F, G: M holotype (ZFMKMYR9482B1). H: M paratype (ZFMK-MYR 9482B2). A: Head, frontal view. B: Antenna. C: Sensilla basiconica on antennomeres 5 and 6. D: Apical disc with four sensory cones. E: Mouth parts, dorsal view. F: Mouth parts, ventral view. G: Anterior body, lateral view. H: Tergites, lateral view. Abbreviations: ac = apical cone; ag = anterior gonopod; an = antenna; br = body ring; co = collum; gn = gnathochilarium; le = leg; lb = labrum; ms = macrosetae; mz = metazonite; om = ommatidia; oz = ozopore; pg = posterior gonopod; pl = pleurite; pz = prozonite; sb = sensilla basiconica; sp = spiracle; st = sternite. I–VII indicates the number of antennomeres. The white arrow on (B) indicates the row of striae at the anterior margin of the antennomeres and on (H) indicates the lateral, ridge-like extension of the prozonite. Scale bars: A, G, H = 100 µm; B, D–F = 20 µm; C = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.7059577" httpUri="https://zenodo.org/record/7059577/files/figure.png" pageId="5" pageNumber="406">Fig. 3H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CA816538B908E81D6D91F887FC055A1D" pageId="5" pageNumber="406" type="etymology">
|
||
<paragraph id="822436B3B908E81D6D91F887FC055A1D" blockId="5.[151,1437,1472,1895]" pageId="5" pageNumber="406">
|
||
<emphasis id="B0EFEAA1B908E81D6D91F887FED85A38" bold="true" box="[199,334,1833,1859]" pageId="5" pageNumber="406">Etymology.</emphasis>
|
||
Genus epithet,
|
||
<taxonomicName id="459B4D30B908E81D6F58F886FDF85A39" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[526,622,1832,1858]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B0EFEAA1B908E81D6F58F886FDF85A39" box="[526,622,1832,1858]" italics="true" pageId="5" pageNumber="406">Theratta</emphasis>
|
||
</taxonomicName>
|
||
, noun, feminine. Millipedes are called ‘
|
||
<taxonomicName id="459B4D30B908E81D6904F886FB245A39" authorityName="Anilkumar & Wesener & Moritz" authorityYear="2022" box="[1106,1202,1832,1858]" class="Diplopoda" family="Siphonotidae" genus="Theratta" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B0EFEAA1B908E81D6904F886FB245A39" box="[1106,1202,1832,1858]" italics="true" pageId="5" pageNumber="406">Theratta</emphasis>
|
||
</taxonomicName>
|
||
’ in Malayalam, the language spoken in the Indian province (
|
||
<collectingRegion id="405FF851B908E81D6F09F8E2FD3B5A1D" box="[607,685,1868,1894]" country="India" name="Kerala" pageId="5" pageNumber="406">Kerala</collectingRegion>
|
||
) of the
|
||
<typeStatus id="5D208811B908E81D6E55F8E3FCA55A1C" box="[771,819,1869,1895]" pageId="5" pageNumber="406">type</typeStatus>
|
||
species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |