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<document id="5287E012FEB034CD57B7B804CD1AB8E0" ID-CLB-Dataset="53154" ID-DOI="10.11646/zootaxa.5124.2.5" ID-GBIF-Dataset="e2bc8855-27f6-49b8-af3d-42c4939bc9d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="6405090" ID-ZooBank="B2745AA1-8B1D-47C2-B648-A12386BC2574" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648798060305" checkinUser="plazi" docAuthor="Mendes, Luanna L., Evenhuis, Neal L., Limeira-De-Oliveira, Francisco &amp; Lamas, Carlos J. E." docDate="2022" docId="038BF530FFF88434E68AFEC06B65F8E4" docLanguage="en" docName="zootaxa.5124.2.5.pdf" docOrigin="Zootaxa 5124 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glabellula catiapatiuae Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="207" masterDocId="FFB28D48FFFA8436E61DFE576F5DFFCD" masterDocTitle="First record of Glabellula Bezzi (Diptera, Mythicomyiidae: Glabellulinae) in South America, with description of three new species from Brazil" masterLastPageNumber="215" masterPageNumber="205" pageNumber="207" updateTime="1699319211259" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4B27396F1A3481ECB8135AFBF423F503">First record of Glabellula Bezzi (Diptera, Mythicomyiidae: Glabellulinae) in South America, with description of three new species from Brazil</mods:title>
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<mods:namePart id="2FD32A67A5D43BCA09837D13C55B1538">Mendes, Luanna L.</mods:namePart>
<mods:nameIdentifier id="09EF29E56081E7A64B6F4A77A21C1DEC" type="ORCID">0000-0002-5115-9914</mods:nameIdentifier>
<mods:affiliation id="7650F43D29C4E989C75433129DC9CC4B">Museu de Zoologia da Universidade de São Paulo, São Paulo, São Paulo, Avenida Nazaré, 481, Ipiranga, 04263 - 000, São Paulo, SP, Brazil. luannalmendes @ hotmail. com; https: // orcid. org / 0000 - 0002 - 5115 - 9914</mods:affiliation>
<mods:nameIdentifier id="EB8912CB798466910D7D6771409E9071" type="email">luannalmendes@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="398EF24E6FE7753D392C8B98EB921BF9">Evenhuis, Neal L.</mods:namePart>
<mods:nameIdentifier id="87915576188D1EA00C769E956FFE496F" type="ORCID">0000-0002-1314-755X</mods:nameIdentifier>
<mods:affiliation id="738F6C0AD1F0CEF66068B935A0B2B463">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA. neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
<mods:nameIdentifier id="4813D2853C68FB4A02511D2F911574FA" type="email">neale@bishopmuseum.org</mods:nameIdentifier>
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<mods:namePart id="180F8B2890F63AB90ABEDE80247AD00F">Limeira-De-Oliveira, Francisco</mods:namePart>
<mods:nameIdentifier id="A907433562240D056B7050A31B93CA20" type="ORCID">0000-0002-4980-5454</mods:nameIdentifier>
<mods:affiliation id="9A7588FA3CFB4330C631B407E206FF4A">Universidade Estadual do Maranhão, Laboratório de Estudos dos Invertebrados, 65604 - 380, Caxias, Maranhão, Brazil. limeiraf @ gmail. com; https: // orcid. org / 0000 - 0002 - 4980 - 5454</mods:affiliation>
<mods:nameIdentifier id="24845F0F3034EB5AC7ABF57447A1D120" type="email">limeiraf@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0A1D5799CA005E933CB737909F8FD319">Lamas, Carlos J. E.</mods:namePart>
<mods:nameIdentifier id="B9520F7B01C65717A8D58E5DEB8BEC1B" type="ORCID">0000-0002-7750-590X</mods:nameIdentifier>
<mods:affiliation id="6A05DFF282E64F7D43B08D3FA3924632">Museu de Zoologia da Universidade de São Paulo, São Paulo, São Paulo, Avenida Nazaré, 481, Ipiranga, 04263 - 000, São Paulo, SP, Brazil. einicker @ usp. br; https: // orcid. org / 0000 - 0002 - 7750 - 590 X</mods:affiliation>
<mods:nameIdentifier id="A520E7EFD78D98FD7F998B500055E664" type="email">einicker@usp.br</mods:nameIdentifier>
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<mods:title id="3CD8256C1B41ED58FEA7AB5B42938638">Zootaxa</mods:title>
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<mods:date id="DD805FE95B753891DBCE69E0C1E771F1">2022</mods:date>
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<treatment id="038BF530FFF88434E68AFEC06B65F8E4" ID-DOI="http://doi.org/10.5281/zenodo.6405079" ID-GBIF-Taxon="194442516" ID-Zenodo-Dep="6405079" LSID="urn:lsid:plazi:treatment:038BF530FFF88434E68AFEC06B65F8E4" httpUri="http://treatment.plazi.org/id/038BF530FFF88434E68AFEC06B65F8E4" lastPageNumber="207" pageId="2" pageNumber="207">
<subSubSection id="C33817ADFFF88434E68AFEC06D4AFF7C" box="[151,535,151,178]" pageId="2" pageNumber="207" type="nomenclature">
<paragraph id="8B9D4426FFF88434E68AFEC06D4AFF7C" blockId="2.[151,535,151,214]" box="[151,535,151,178]" pageId="2" pageNumber="207">
<heading id="D0D5F34AFFF88434E68AFEC06D4AFF7C" bold="true" box="[151,535,151,178]" fontSize="11" level="1" pageId="2" pageNumber="207" reason="1">
<emphasis id="B9569834FFF88434E68AFEC06D4AFF7C" bold="true" box="[151,535,151,178]" pageId="2" pageNumber="207">
<taxonomicName id="4C223FA5FFF88434E68AFEC06EEFFF7C" authority="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas, 2022" authorityName="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas" authorityYear="2022" box="[151,434,151,178]" class="Insecta" family="Bombyliidae" genus="Glabellula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="catiapatiuae" status="sp. nov.">
<emphasis id="B9569834FFF88434E68AFEC06EEFFF7C" bold="true" box="[151,434,151,178]" italics="true" pageId="2" pageNumber="207">Glabellula catiapatiuae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265254FFFF88434E7A4FEC06D4AFF7C" box="[441,535,151,177]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33817ADFFF88434E68AFEEC6E6AFF18" box="[151,311,187,214]" pageId="2" pageNumber="207" type="description">
<paragraph id="8B9D4426FFF88434E68AFEEC6E6AFF18" blockId="2.[151,535,151,214]" box="[151,311,187,214]" pageId="2" pageNumber="207">
(
<figureCitation id="131958A3FFF88434E682FEEC6E58FF1B" box="[159,261,187,214]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Figs 15</figureCitation>
,
<figureCitation id="131958A3FFF88434E70FFEEC6E72FF18" box="[274,303,187,213]" captionStart="FIGURES 1719" captionStartId="8.[151,264,637,661]" captionTargetBox="[180,1412,204,598]" captionTargetId="figure-20@8.[151,1436,181,613]" captionTargetPageId="8" captionText="FIGURES 1719. Sternites 7 and 8 of the new species of Glabellula, ventral view. 17, G. catiapatiuae sp. nov.; 18, G. rafaelae sp. nov.; 19, G. sarahae sp. nov. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405098" httpUri="https://zenodo.org/record/6405098/files/figure.png" pageId="2" pageNumber="207">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33817ADFFF88434E68AFF546C2AFE8C" pageId="2" pageNumber="207" type="diagnosis">
<paragraph id="8B9D4426FFF88434E68AFF546C2AFE8C" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E68AFF546E4FFED0" bold="true" box="[151,274,259,285]" pageId="2" pageNumber="207">Diagnosis.</emphasis>
Cell
<emphasis id="B9569834FFF88434E751FF536E29FED0" box="[332,372,260,285]" italics="true" pageId="2" pageNumber="207">cua</emphasis>
at wing margin closed; mesonotum, scutellum, tergites and femora entirely dark brown, without yellow areas; female sternite 7 with lateral digitiform projections.
</paragraph>
</subSubSection>
<subSubSection id="C33817ADFFF88434E6DAFF1C6B2BFE44" pageId="2" pageNumber="207" type="etymology">
<paragraph id="8B9D4426FFF88434E6DAFF1C6B2BFE44" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAFF1C6E13FEA8" bold="true" box="[199,334,331,357]" pageId="2" pageNumber="207">Etymology.</emphasis>
The specific epithet is a patronymic tribute to the late Dr. Catia Antunes de Mello-Patiu (MNRJ) for her great contribution to the knowledge of the diversity of
<taxonomicName id="4C223FA5FFF88434E552FF386CFEFE44" box="[847,931,367,393]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
in South America.
</paragraph>
</subSubSection>
<subSubSection id="C33817ADFFF88434E6DAFFC46DE4F9E0" pageId="2" pageNumber="207" type="description">
<paragraph id="8B9D4426FFF88434E6DAFFC46E88FA0F" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAFFC46D79FE60" bold="true" box="[199,548,403,429]" pageId="2" pageNumber="207">
Description.
<typeStatus id="5499FA84FFF88434E77CFFC46E93FE60" box="[353,462,403,429]" pageId="2" pageNumber="207" type="holotype">Holotype</typeStatus>
female
</emphasis>
(
<figureCitation id="131958A3FFF88434E428FFC46D2AFE60" box="[565,631,403,429]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 1</figureCitation>
). Body length:
<quantity id="4CDAE9C3FFF88434E536FFC46CCEFE63" box="[811,915,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.54" pageId="2" pageNumber="207" unit="mm" value="1.54">1.54 mm</quantity>
; wing length:
<quantity id="4CDAE9C3FFF88434E228FFC46BC0FE63" box="[1077,1181,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="207" unit="mm" value="1.25">1.25 mm</quantity>
.
<emphasis id="B9569834FFF88434E2B5FFC46BB5FE60" bold="true" box="[1192,1256,403,429]" pageId="2" pageNumber="207">Head</emphasis>
. Dark brown to black; eyes separated at vertex by approximately 1.25X distance between posterior ocelli; ocellar triangle shiny, ocelli yellow to dark yellow; frons dark brown, yellow in ventral and lateral margin, shiny, slightly depressed medially, with small yellow setae laterally; face yellow, with brown mark medially, slightly projecting beyond eye in lateral view; proboscis light brown to brown, smaller than head length; oral cavity and margin yellow; postcranium subshiny to shiny; setae whitish yellow; antenna brown to dark brown, yellowish pruinose; scape minute; pedicel oval; first flagellomere elliptical, about 1.5X longer than width; second flagellomere very small, about one-half width of first one; stylus absent.
<emphasis id="B9569834FFF88434E4A5FCD86C4CFD64" bold="true" box="[696,785,655,681]" pageId="2" pageNumber="207">Thorax</emphasis>
(
<figureCitation id="131958A3FFF88434E53CFCD86C3AFD64" box="[801,871,655,681]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 3</figureCitation>
). Mesonotum and scutellum dark brown to black, subshiny to shiny, covered with whitish yellow setae; yellow markings on: postpronotal lobe, postalar callus and thin line along transverse suture; pleura dark brown to black, shiny, with yellow markings on: proepimeron; anterior and dorsal narrow margin of anepisternum, dorsal margin of katepisternum and dorsal 1/3 of meron.
<emphasis id="B9569834FFF88434E338FCAC6A00FCD8" bold="true" box="[1317,1373,763,789]" pageId="2" pageNumber="207">Legs</emphasis>
(
<figureCitation id="131958A3FFF88434E372FCAC6FF9FCF4" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 1</figureCitation>
). Dark brown to black; femora with distal extremity dark yellow; tibiae with proximal and distal extremity dark yellow; tarsi dark yellow, last tarsomere darker.
<emphasis id="B9569834FFF88434E4AFFD146DAEFC90" bold="true" box="[690,755,835,861]" pageId="2" pageNumber="207">Wing</emphasis>
(
<figureCitation id="131958A3FFF88434E51EFD146C15FC90" box="[771,840,835,861]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 4</figureCitation>
). Hyaline; costal vein, radial veins and crossvein
<emphasis id="B9569834FFF88434E369FD136AC1FC90" box="[1396,1436,836,861]" italics="true" pageId="2" pageNumber="207">r-m</emphasis>
light brown, remainder veins pale yellow to translucid; costal vein ending at 1/3 distance between veins
<emphasis id="B9569834FFF88434E322FD3F6A36FC45" box="[1343,1387,872,904]" italics="true" pageId="2" pageNumber="207">
R
<subScript id="17A64663FFF88434E34DFD2D6A36FC45" attach="left" box="[1360,1387,890,904]" fontSize="6" pageId="2" pageNumber="207">4+5</subScript>
</emphasis>
and
<emphasis id="B9569834FFF88434E68AFDDB6FEAFC61" box="[151,183,908,940]" italics="true" pageId="2" pageNumber="207">
M
<subScript id="17A64663FFF88434E6B2FDC96FEAFC61" attach="right" box="[175,183,926,940]" fontSize="6" pageId="2" pageNumber="207">1</subScript>
</emphasis>
; vein
<emphasis id="B9569834FFF88434E71FFDDC6E41FC68" box="[258,284,907,933]" italics="true" pageId="2" pageNumber="207">Sc</emphasis>
incomplete, ending at level of
<emphasis id="B9569834FFF88434E495FDDB6DF9FC68" box="[648,676,908,933]" italics="true" pageId="2" pageNumber="207">Rs</emphasis>
origin;
<emphasis id="B9569834FFF88434E51FFDDB6C43FC68" box="[770,798,908,933]" italics="true" pageId="2" pageNumber="207">Rs</emphasis>
evanescent at connection with
<emphasis id="B9569834FFF88434E297FDDB6BF9FC61" box="[1162,1188,908,940]" italics="true" pageId="2" pageNumber="207">
R
<subScript id="17A64663FFF88434E281FDC96BF9FC61" attach="right" box="[1180,1188,926,940]" fontSize="6" pageId="2" pageNumber="207">1</subScript>
</emphasis>
;
<emphasis id="B9569834FFF88434E2A8FDDB6BBFFC61" box="[1205,1250,908,940]" italics="true" pageId="2" pageNumber="207">
R
<subScript id="17A64663FFF88434E2DBFDC96BBFFC61" attach="left" box="[1222,1250,926,940]" fontSize="6" pageId="2" pageNumber="207">4+5</subScript>
</emphasis>
slightly curved, ending near end of
<emphasis id="B9569834FFF88434E76BFDE76EC8FC1D" box="[374,405,944,976]" italics="true" pageId="2" pageNumber="207">
M
<subScript id="17A64663FFF88434E790FD956EC8FC1D" attach="left" box="[397,405,962,976]" fontSize="6" pageId="2" pageNumber="207">1</subScript>
</emphasis>
;
<emphasis id="B9569834FFF88434E7BBFDE76E98FC1D" box="[422,453,944,976]" italics="true" pageId="2" pageNumber="207">
M
<subScript id="17A64663FFF88434E7A0FD956E98FC1D" attach="left" box="[445,453,962,976]" fontSize="6" pageId="2" pageNumber="207">1</subScript>
</emphasis>
slightly curved toward wing margin;
<emphasis id="B9569834FFF88434E56BFDE76CC8FC1D" box="[886,917,944,976]" italics="true" pageId="2" pageNumber="207">
M
<subScript id="17A64663FFF88434E590FD956CC8FC1D" attach="left" box="[909,917,962,976]" fontSize="6" pageId="2" pageNumber="207">2</subScript>
</emphasis>
and
<emphasis id="B9569834FFF88434E5D2FDE76CB2FC1D" box="[975,1007,944,976]" italics="true" pageId="2" pageNumber="207">
M
<subScript id="17A64663FFF88434E5FAFD956CB2FC1D" attach="left" box="[999,1007,962,976]" fontSize="6" pageId="2" pageNumber="207">4</subScript>
</emphasis>
straight to wing margin;
<emphasis id="B9569834FFF88434E30FFDE76A19FC07" box="[1298,1348,944,970]" italics="true" pageId="2" pageNumber="207">CuP</emphasis>
joining
<emphasis id="B9569834FFF88434E68AFD846F94FC20" box="[151,201,979,1005]" italics="true" pageId="2" pageNumber="207">CuA</emphasis>
at wing margin;
<emphasis id="B9569834FFF88434E79AFD846EFCFC39" box="[391,417,979,1012]" italics="true" pageId="2" pageNumber="207">
A
<subScript id="17A64663FFF88434E784FDB16EFCFC39" attach="left" box="[409,417,998,1012]" fontSize="6" pageId="2" pageNumber="207">1</subScript>
</emphasis>
evanescent towards wing margin; cell
<emphasis id="B9569834FFF88434E548FD836C20FC20" box="[853,893,980,1005]" italics="true" pageId="2" pageNumber="207">cua</emphasis>
closed at wing margin. Halter with stem yellow, slightly brownish infuscate, knob yellow with dark brown marks.
<emphasis id="B9569834FFF88434E59CFDA06CABFBDC" bold="true" box="[897,1014,1015,1041]" pageId="2" pageNumber="207">Abdomen</emphasis>
(
<figureCitation id="131958A3FFF88434E21BFDA06B10FBDC" box="[1030,1101,1015,1041]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 2</figureCitation>
). Dark brown to black, shiny, covered with yellowish setae uniform in length; sternite 7 (
<figureCitation id="131958A3FFF88434E537FA4C6C26FBF8" box="[810,891,1051,1077]" captionStart="FIGURES 1719" captionStartId="8.[151,264,637,661]" captionTargetBox="[180,1412,204,598]" captionTargetId="figure-20@8.[151,1436,181,613]" captionTargetPageId="8" captionText="FIGURES 1719. Sternites 7 and 8 of the new species of Glabellula, ventral view. 17, G. catiapatiuae sp. nov.; 18, G. rafaelae sp. nov.; 19, G. sarahae sp. nov. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405098" httpUri="https://zenodo.org/record/6405098/files/figure.png" pageId="2" pageNumber="207">Fig. 17</figureCitation>
) with anterior margin irregular, posterior margin with lateral digitiform projection and shallow concavity medially, setae uniform in length, slightly longer distally; sternite 8 (
<figureCitation id="131958A3FFF88434E70CFA346E3EFBB0" box="[273,355,1123,1149]" captionStart="FIGURES 1719" captionStartId="8.[151,264,637,661]" captionTargetBox="[180,1412,204,598]" captionTargetId="figure-20@8.[151,1436,181,613]" captionTargetPageId="8" captionText="FIGURES 1719. Sternites 7 and 8 of the new species of Glabellula, ventral view. 17, G. catiapatiuae sp. nov.; 18, G. rafaelae sp. nov.; 19, G. sarahae sp. nov. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405098" httpUri="https://zenodo.org/record/6405098/files/figure.png" pageId="2" pageNumber="207">Fig. 17</figureCitation>
) reduced, sub-trapezoidal, with deep concave indentation medially placed on posterior margin.
<emphasis id="B9569834FFF88434E68AFAD06E47FB6C" bold="true" box="[151,282,1159,1185]" pageId="2" pageNumber="207">Terminalia</emphasis>
(
<figureCitation id="131958A3FFF88434E737FAD06E31FB6F" box="[298,364,1159,1186]" captionStart="FIGURES 15" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[196,1390,202,1858]" captionTargetId="figure-19@3.[175,1412,181,1919]" captionTargetPageId="3" captionText="FIGURES 15. Glabellula catiapatiuae sp. nov., female, holotype. 1, habitus, lateral view; 2, abdomen, dorsal view; 3, thorax, dorsal view; 4, left wing, ventral view; 5, terminalia, holotype. Scale 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6405092" httpUri="https://zenodo.org/record/6405092/files/figure.png" pageId="2" pageNumber="207">Fig. 5</figureCitation>
). Genital fork V-shaped, anterior margin arched with small conical projection, posterior margin bifurcate, inner arms convergent with central circular orifice, outer arms divergent, longer and tapered toward apex. Common duct short, about 1/4 length of spermatheca, basal plate hyaline; proximal spermathecal duct reduced or absent; sperm pump tubular-shaped with apical sclerotized collars well developed and formed by two level structure complex, proximally surrounded by membranous fringe and distally by well sclerotized horseshoe-shaped fold; amorphous membranous extroversion arise from sclerotized collars apex; distal spermathecal duct very long and thin, 5.5X longer than sperm pump, with elliptical swelling near base of spermatheca, swelling surrounded by minute, tangled, glandular trichomes; spermatheca as long as sperm pump, sclerotized in distal 1/3, brown, with glandular trichomes at apex.
</paragraph>
<paragraph id="8B9D4426FFF88434E6DAFB9C6ED5FA28" blockId="2.[151,1437,259,1834]" box="[199,392,1483,1509]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAFB9C6E56FA28" bold="true" box="[199,267,1483,1509]" pageId="2" pageNumber="207">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8B9D4426FFF88434E6DAFBB86DE4F9E0" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAFBB86E8EF9C4" bold="true" box="[199,467,1519,1545]" pageId="2" pageNumber="207">Geographical records.</emphasis>
This species is known only from the type-locality, which is inserted in the Cerrado biome of the state of
<collectingRegion id="49E68AC4FFF88434E799F8446EA5F9E0" box="[388,504,1555,1581]" country="Brazil" name="Maranhao" pageId="2" pageNumber="207">Maranhão</collectingRegion>
,
<collectingCountry id="F33504B6FFF88434E419F8446D15F9E0" box="[516,584,1555,1581]" name="Brazil" pageId="2" pageNumber="207">Brazil</collectingCountry>
(
<figureCitation id="131958A3FFF88434E445F8446DF1F9E0" box="[600,684,1555,1581]" captionStart="FIGURE 20" captionStartId="8.[151,250,1536,1560]" captionTargetBox="[178,1417,745,1490]" captionTargetId="figure-63@8.[151,1436,722,1512]" captionTargetPageId="8" captionText="FIGURE 20. Geographical records of new species of Glabellula known from Brazil." figureDoi="http://doi.org/10.5281/zenodo.6405100" httpUri="https://zenodo.org/record/6405100/files/figure.png" pageId="2" pageNumber="207">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33817ADFFF88434E6DAF8606B65F8E4" pageId="2" pageNumber="207" type="materials_examined">
<paragraph id="8B9D4426FFF88434E6DAF8606C1BF954" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAF8606E28F99C" bold="true" box="[199,373,1591,1617]" pageId="2" pageNumber="207">Type material.</emphasis>
<materialsCitation id="3B4A4E7BFFF88434E761F8606C1BF954" ID-GBIF-Occurrence="3746287307" collectingDate="2013-10-10" collectingDateMax="2013-10-20" collectingDateMin="2013-10-10" collectionCode="MZUSP, T" collectorName="F. Limeira-de-Oliveira &amp; L.L.M. Santos &amp; T.L. Rocha" country="Brazil" county="Mirador" elevation="416" latitude="-6.63" location="Base da Geraldina" longLatPrecision="21" longitude="-45.880276" municipality="Mirador" pageId="2" pageNumber="207" specimenCount="1" specimenCount-female="1" stateProvince="Maranhao" typeStatus="holotype">
<emphasis id="B9569834FFF88434E761F8606D46F99C" bold="true" box="[380,539,1591,1617]" pageId="2" pageNumber="207">
<typeStatus id="5499FA84FFF88434E761F8606D46F99C" box="[380,539,1591,1617]" pageId="2" pageNumber="207" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="9D248FAFFFF88434E43CF86F6D6BF99F" box="[545,566,1592,1618]" pageId="2" pageNumber="207" type="female"></specimenCount>
(dissected) deposited in
<collectionCode id="ED33DCE3FFF88434E551F8606CF3F99C" box="[844,942,1591,1617]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="207">MZUSP</collectionCode>
:
<collectingCountry id="F33504B6FFF88434E5A5F8606CA3F99C" box="[952,1022,1591,1617]" name="Brazil" pageId="2" pageNumber="207">Brasil</collectingCountry>
,
<emphasis id="B9569834FFF88434E214F8606B6BF99C" bold="true" box="[1033,1078,1591,1617]" pageId="2" pageNumber="207">MA</emphasis>
[=
<collectingRegion id="49E68AC4FFF88434E240F8616B89F99C" box="[1117,1236,1590,1617]" country="Brazil" name="Maranhao" pageId="2" pageNumber="207">Maranhão</collectingRegion>
],
<collectingCounty id="62FC3CAAFFF88434E2FBF8606A1CF99C" box="[1254,1345,1591,1617]" pageId="2" pageNumber="207">Mirador</collectingCounty>
, Parque Est. [= Estadual do]
<collectingMunicipality id="6BF9DE5CFFF88434E79BF80C6EB9F9B8" box="[390,484,1627,1653]" pageId="2" pageNumber="207">Mirador</collectingMunicipality>
,
<location id="8EFD12FDFFF88434E7ECF80B6D9BF9B8" LSID="urn:lsid:plazi:treatment:038BF530FFF88434E68AFEC06B65F8E4:8EFD12FDFFF88434E7ECF80B6D9BF9B8" box="[497,710,1627,1654]" country="Brazil" county="Mirador" latitude="-6.63" longLatPrecision="21" longitude="-45.880276" municipality="Mirador" name="Base da Geraldina" pageId="2" pageNumber="207" stateProvince="Maranhao">
Base
<collectorName id="26D721F0FFF88434E42CF80C6D9BF9B8" box="[561,710,1627,1653]" pageId="2" pageNumber="207">da Geraldina</collectorName>
</location>
,
<quantity id="4CDAE9C3FFF88434E4C9F80C6C42F9BB" box="[724,799,1627,1654]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="2" pageNumber="207" unit="m" value="416.0">
<elevation id="000FA315FFF88434E4C9F80C6C42F9BB" box="[724,799,1627,1654]" metricMagnitude="2" metricUnit="m" metricValue="4.16" pageId="2" pageNumber="207" unit="m" value="416.0">416 m</elevation>
</quantity>
,
<geoCoordinate id="EE1622E1FFF88434E530F80A6CF3F9B5" box="[813,942,1629,1656]" degrees="06" direction="south" minutes="37" orientation="latitude" pageId="2" pageNumber="207" precision="15" seconds="48" value="-6.63">06º3748″S</geoCoordinate>
,
<geoCoordinate id="EE1622E1FFF88434E5A6F80A6B1EF9B5" box="[955,1091,1629,1656]" degrees="45" direction="west" minutes="52" orientation="longitude" pageId="2" pageNumber="207" precision="15" seconds="49" value="-45.880276">45º5249″W</geoCoordinate>
/
<collectorName id="26D721F0FFF88434E249F80C6A1DF9B8" box="[1108,1344,1627,1653]" pageId="2" pageNumber="207">Armadilha Suspensa</collectorName>
,
<date id="FF9C62E6FFF88434E353F80C6FBAF954" pageId="2" pageNumber="207" value="2013-10-10" valueMax="2013-10-20" valueMin="2013-10-10">
<collectingDate id="EFD89B0EFFF88434E353F80C6FBAF954" pageId="2" pageNumber="207" value="2013-10-10" valueMax="2013-10-20" valueMin="2013-10-10">1020. x.2013</collectingDate>
</date>
,
<collectorName id="26D721F0FFF88434E6EFF8D76EAFF954" box="[242,498,1663,1690]" pageId="2" pageNumber="207">F. Limeira-de-Oliveira</collectorName>
,
<collectorName id="26D721F0FFF88434E7E0F8D76DC2F954" box="[509,671,1663,1690]" pageId="2" pageNumber="207">L.L.M. Santos</collectorName>
&amp;
<collectionCode id="ED33DCE3FFF88434E4DFF8D76D88F957" box="[706,725,1664,1690]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="207">T</collectionCode>
.
<collectorName id="26D721F0FFF88434E4C4F8D76C1BF954" box="[729,838,1663,1690]" pageId="2" pageNumber="207">L. Rocha.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B9D4426FFF88434E6DAF8F46B65F8E4" blockId="2.[151,1437,259,1834]" pageId="2" pageNumber="207">
<emphasis id="B9569834FFF88434E6DAF8F46E5BF970" bold="true" box="[199,262,1699,1725]" pageId="2" pageNumber="207">Note.</emphasis>
Closed cell
<emphasis id="B9569834FFF88434E78DF8F36EE5F970" box="[400,440,1700,1725]" italics="true" pageId="2" pageNumber="207">cua</emphasis>
at the wing margin in
<taxonomicName id="4C223FA5FFF88434E4AEF8F36C3CF970" authority="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas, 2022" authorityName="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas" authorityYear="2022" box="[691,865,1700,1726]" class="Insecta" family="Bombyliidae" genus="Glabellula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="catiapatiuae" status="sp. nov.">
<emphasis id="B9569834FFF88434E4AEF8F36C3CF970" box="[691,865,1700,1726]" italics="true" pageId="2" pageNumber="207">G. catiapatiuae</emphasis>
</taxonomicName>
<emphasis id="B9569834FFF88434E575F8F46C9CF970" bold="true" box="[872,961,1699,1725]" pageId="2" pageNumber="207">
<taxonomicNameLabel id="A265254FFFF88434E575F8F46C9CF970" box="[872,961,1699,1725]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is uncommon and unique among the other species found in
<collectingCountry id="F33504B6FFF88434E741F8906EFEF92C" box="[348,419,1735,1761]" name="Brazil" pageId="2" pageNumber="207">Brazil</collectingCountry>
, in which cell
<emphasis id="B9569834FFF88434E44DF89F6D25F92C" box="[592,632,1736,1761]" italics="true" pageId="2" pageNumber="207">cua</emphasis>
is always widely opened. Since this new species is known from only a single female specimen, future investigations of
<taxonomicName id="4C223FA5FFF88434E4FCF8BB6CCEF8C8" authority="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas, 2022" authorityName="Mendes &amp; Evenhuis &amp; Limeira-De-Oliveira &amp; Lamas" authorityYear="2022" box="[737,915,1772,1798]" class="Insecta" family="Bombyliidae" genus="Glabellula" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="207" phylum="Arthropoda" rank="species" species="catiapatiuae" status="sp. nov.">
<emphasis id="B9569834FFF88434E4FCF8BB6CCEF8C8" box="[737,915,1772,1798]" italics="true" pageId="2" pageNumber="207">G. catiapatiuae</emphasis>
</taxonomicName>
<emphasis id="B9569834FFF88434E580F8BC6CA7F8C8" bold="true" box="[925,1018,1771,1797]" pageId="2" pageNumber="207">
<taxonomicNameLabel id="A265254FFFF88434E580F8BC6CA7F8C8" box="[925,1018,1771,1797]" pageId="2" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
samples are necessary to determine whether the closed cell
<emphasis id="B9569834FFF88434E7BDF9476E95F8E4" box="[416,456,1808,1833]" italics="true" pageId="2" pageNumber="207">cua</emphasis>
is truly a diagnostic feature or just individual variation.
</paragraph>
</subSubSection>
</treatment>
</document>