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in 
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Basionym: 
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s.d., 
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slope, 
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, 
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(epitype, 
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[
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] image seen; isoepitype, KLU-36404 [2 sheets] images seen, 
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00018268]!)
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Syn.: 
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<emphasis id="B956BF3EFFCEBF2DFF00542FCD16FBE3" box="[191,381,1122,1143]" italics="true" pageId="1" pageNumber="162">Pandanus gladiator</emphasis>
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. 
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<collectingCountry id="F33523BCFFCEBF2DFD33542FCF79FBE3" box="[652,786,1122,1143]" name="Indonesia" pageId="1" pageNumber="162">INDONESIA</collectingCountry>
. 
<materialsCitation id="3B4A6971FFCEBF2DFCA6542FC810FBEC" box="[793,1147,1122,1144]" collectingDate="1979-05-11" collectionCode="W, L" collectorName="Mt Roroka Timbu, W" country="Philippines" elevation="1050" latitude="-1.2666667" location="Mt Roroka Timbu" longLatPrecision="1308" longitude="120.3" pageId="1" pageNumber="162" specimenCode="L0332713" specimenCount="1" stateProvince="Sulawesi Tengah" typeStatus="holotype">
<collectingRegion id="49E6ADCEFFCEBF2DFCA6542FC871FBEC" box="[793,1050,1122,1144]" country="Indonesia" name="Sulawesi Tengah" pageId="1" pageNumber="162">Central Sulawesi Province</collectingRegion>
: 
<collectingRegion id="49E6ADCEFFCEBF2DFB9D542FC810FBEC" box="[1058,1147,1122,1144]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="162">Sulawesi</collectingRegion>
</materialsCitation>
</materialsCitation>
, 
<materialsCitation id="3B4A6971FFCEBF2DFB3B542ECE26FB55" collectingDate="1979-05-11" collectionCode="W, L" collectorName="Mt Roroka Timbu, W" country="Philippines" elevation="1050" latitude="-1.2666667" location="Mt Roroka Timbu" longLatPrecision="1308" longitude="120.3" pageId="1" pageNumber="162" specimenCode="L0332713" specimenCount="1" typeStatus="holotype">
<collectorName id="26D706FAFFCEBF2DFB3B542EC93BFBEC" box="[1156,1360,1122,1144]" pageId="1" pageNumber="162">
<location id="8EFD35F7FFCEBF2DFB3B542EC959FBEC" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCEBF2DFB3B542EC959FBEC" box="[1156,1330,1122,1144]" country="Philippines" latitude="-1.2666667" longLatPrecision="1308" longitude="120.3" name="Mt Roroka Timbu" pageId="1" pageNumber="162" stateProvince="Sulawesi Tengah">Mt Roroka Timbu</location>
, 
<collectionCode id="ED33FBE9FFCEBF2DFA85542EC93BFBEC" box="[1338,1360,1123,1144]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="1" pageNumber="162" type="Herbarium">W</collectionCode>
</collectorName>
slope, 
<locationDeviation id="9A2F7A0EFFCEBF2DFA2C542FCD33FB08" location="Palu" pageId="1" pageNumber="162">
<quantity id="4CDACEC9FFCEBF2DFA2C542FCCB7FB08" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="162" unit="km" value="80.0">80 km</quantity>
SSE of Palu
</locationDeviation>
, 
<geoCoordinate id="EE1605EBFFCEBF2DFEDE54CBCDCEFB08" box="[353,421,1158,1180]" degrees="1" direction="south" minutes="16" orientation="latitude" pageId="1" pageNumber="162" precision="925" value="-1.2666667">1°16’S</geoCoordinate>
, 
<geoCoordinate id="EE1605EBFFCEBF2DFE0F54CBCE65FB0F" box="[432,526,1158,1179]" degrees="120" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="162" precision="925" value="120.3">120°18’E</geoCoordinate>
, c. 
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<elevation id="000F841FFFCEBF2DFD9154CBCE12FB08" box="[558,633,1158,1180]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="1" pageNumber="162" unit="m" value="1050.0">1050 m</elevation>
</quantity>
, 
<date id="FF9C45ECFFCEBF2DFD3D54CBCE8DFB0F" box="[642,742,1158,1179]" pageId="1" pageNumber="162" value="1979-05-11">
<collectingDate id="EFD8BC04FFCEBF2DFD3D54CBCE8DFB0F" box="[642,742,1158,1179]" pageId="1" pageNumber="162" value="1979-05-11">11.V.1979</collectingDate>
</date>
, 
<emphasis id="B956BF3EFFCEBF2DFD5054CBCF31FB0F" box="[751,858,1158,1179]" italics="true" pageId="1" pageNumber="162">Vogel 5287</emphasis>
(
<typeStatus id="5499DD8EFFCEBF2DFCD854CBCFD4FB08" box="[871,959,1158,1180]" pageId="1" pageNumber="162" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED33FBE9FFCEBF2DFC7754CACFBDFB08" box="[968,982,1159,1180]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="162" type="Herbarium">L</collectionCode>
[
<specimenCode id="DB84CB57FFCEBF2DFC5E54CBC822FB08" box="[993,1097,1158,1180]" collectionCode="L" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="1" pageNumber="162" type="Herbarium">L0332713</specimenCode>
] image seen; isotype, KLU-36404 [2 sheets] images seen, 
<collectingCountry id="F33523BCFFCEBF2DFE3854E6CDCDFB54" box="[391,422,1195,1216]" name="Philippines" pageId="1" pageNumber="162">PH</collectingCountry>
[
<collectingCountry id="F33523BCFFCEBF2DFE0E54E7CDB8FB55" box="[433,467,1194,1217]" name="Philippines" pageId="1" pageNumber="162">PH</collectingCountry>
00018268]!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33830A7FFCEBF2DFF3754A5CFEBF78E" pageId="1" pageNumber="162" type="description">
<paragraph id="8B9D632CFFCEBF2DFF3754A5CFEBF78E" blockId="1.[136,1452,1256,2075]" pageId="1" pageNumber="162">
<emphasis id="B956BF3EFFCEBF2DFF3754A5CC9AFA96" bold="true" box="[136,241,1256,1282]" pageId="1" pageNumber="162">Notes:—</emphasis>
<taxonomicName id="4C2218AFFFCEBF2DFF4F54A4CDA1FA96" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[240,458,1256,1282]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFF4F54A4CDA1FA96" box="[240,458,1256,1282]" italics="true" pageId="1" pageNumber="162">Benstonea celebica</emphasis>
</taxonomicName>
was regarded as an enigmatic species before its rediscovery by AK in 2005 (
<emphasis id="B956BF3EFFCEBF2DFA8A54A4C9C9FA96" box="[1333,1442,1256,1282]" italics="true" pageId="1" pageNumber="162">Keim 558</emphasis>
) in Sulawesi at Pontolotua Hill during a survey of the Lore-Lindu National Park (
<bibRefCitation id="EFB31EDDFFCEBF2DFBB65541C8F8FAB3" author="Keim, A. P." box="[1033,1171,1292,1319]" pageId="1" pageNumber="162" refId="ref3609" refString="Keim, A. P. (2005) A joint Herbarium Bogoriense &amp; HerbariumWallacea exploration in the Lore Lindu National Park, Central Celebes (including Mount Nokilalaki &amp; Adjacent areas): Pandanaceae. Technical report, LIPI, Bogor, 18 pp." type="book" year="2005">Keim, 2005</bibRefCitation>
). The delimitation of this taxon has remained challenging for a century due to the very fragmentary material of the 
<typeStatus id="5499DD8EFFCEBF2DFB2D557CC8A9FADF" box="[1170,1218,1329,1355]" pageId="1" pageNumber="162">type</typeStatus>
(
<emphasis id="B956BF3EFFCEBF2DFB6A557DC90CFADE" box="[1237,1383,1328,1354]" italics="true" pageId="1" pageNumber="162">Warburg s.n.</emphasis>
). The fruiting material deposited at B was apparently destroyed during World War II, but a photograph exists at FI. 
<bibRefCitation id="EFB31EDDFFCEBF2DFAD35519CCB8FA06" author="Stone, B. C." pageId="1" pageNumber="162" pagination="1 - 74" refId="ref4104" refString="Stone, B. C. (1978) Revisio Pandanacearum, Part I. Pandanus subgenera Coronata and Acrostigma. Flora Malesaina precursores. Federation Museums Journal, new series 23: 1 - 74." type="book chapter" year="1978">Stone (1978)</bibRefCitation>
has even suggested that 
<taxonomicName id="4C2218AFFFCEBF2DFE575534CE08FA06" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[488,611,1400,1426]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFE575534CE08FA06" box="[488,611,1400,1426]" italics="true" pageId="1" pageNumber="162">B. celebica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2218AFFFCEBF2DFD255534C91FFA06" authority="(Merrill 1922: 154) Callmander &amp; al. (2012: 337)" authorityName="Callmander &amp; al." authorityYear="2012" baseAuthorityName="Merrill" baseAuthorityPageNumber="154" baseAuthorityYear="1922" box="[666,1396,1400,1427]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="pachyphylla">
<emphasis id="B956BF3EFFCEBF2DFD255534CF29FA06" box="[666,834,1400,1426]" italics="true" pageId="1" pageNumber="162">B. pachyphylla</emphasis>
(
<bibRefCitation id="EFB31EDDFFCEBF2DFCEF5535C871FA07" author="Merrill, E. D." box="[848,1050,1400,1427]" pageId="1" pageNumber="162" pagination="151 - 201" refId="ref3814" refString="Merrill, E. D. (1922) New or noteworthy Bornean plants, part 1. Journal of the Straits Branch of the Royal Asiatic Society 85: 151 - 201." type="journal article" year="1922">Merrill 1922: 154</bibRefCitation>
) 
<bibRefCitation id="EFB31EDDFFCEBF2DFB965535C91FFA06" author="Callmander, M. W. &amp; Lowry II, P. P. &amp; Forest, F. &amp; Devey, D. S. &amp; Beentje, H. &amp; Buerki, S." box="[1065,1396,1400,1426]" pageId="1" pageNumber="162" pagination="323 - 345" refId="ref3439" refString="Callmander, M. W., Lowry II, P. P., Forest, F., Devey, D. S., Beentje, H. &amp; Buerki, S. (2012) Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species. Candollea 67: 323 - 345." type="journal article" year="2012">Callmander &amp; al. (2012: 337)</bibRefCitation>
</taxonomicName>
may be conspecific. 
<taxonomicName id="4C2218AFFFCEBF2DFE8355D0CE73FA22" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[316,536,1436,1462]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFE8355D0CE73FA22" box="[316,536,1436,1462]" italics="true" pageId="1" pageNumber="162">Benstonea celebica</emphasis>
</taxonomicName>
is in reality very different from 
<taxonomicName id="4C2218AFFFCEBF2DFC2B55D0C854FA22" authorityName="Callmander &amp; al." authorityYear="2012" baseAuthorityName="Merrill" baseAuthorityYear="1922" box="[916,1087,1436,1462]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="pachyphylla">
<emphasis id="B956BF3EFFCEBF2DFC2B55D0C854FA22" box="[916,1087,1436,1462]" italics="true" pageId="1" pageNumber="162">B. pachyphylla</emphasis>
</taxonomicName>
, which is a slender acaulescent shrub bearing a small syncarp with a small infrafoliar near-ground solitary syncarp (see 
<bibRefCitation id="EFB31EDDFFCEBF2DFB3F558DC91AFA4F" author="Keim, A. P. &amp; Rugayah &amp; Rustiami, H." box="[1152,1393,1472,1499]" pageId="1" pageNumber="162" pagination="31 - 62" refId="ref3727" refString="Keim, A. P., Rugayah &amp; Rustiami, H. (2011) The Pandanaceae of the Bukit Baka Bukit Raya National Park and adjacent areas, West and Central Kalimantan, Indonesia, with notes an their nomenclature and the rediscovery of Pandanus aristatus and several new records. Garden's Bulletin Singapore 63: 31 - 62." type="journal article" year="2011">Keim &amp; al. 2011: 52</bibRefCitation>
, Fig. 12; 
<bibRefCitation id="EFB31EDDFFCEBF2DFF0955A9CD95FA6A" author="Callmander, M. W. &amp; Lowry II, P. P. &amp; Forest, F. &amp; Devey, D. S. &amp; Beentje, H. &amp; Buerki, S." box="[182,510,1508,1534]" pageId="1" pageNumber="162" pagination="323 - 345" refId="ref3439" refString="Callmander, M. W., Lowry II, P. P., Forest, F., Devey, D. S., Beentje, H. &amp; Buerki, S. (2012) Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species. Candollea 67: 323 - 345." type="journal article" year="2012">Callmander &amp; al. 2012: 331</bibRefCitation>
, Fig. 4D). 
<taxonomicName id="4C2218AFFFCEBF2DFD3F55A8CF37FA6A" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[640,860,1508,1534]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFD3F55A8CF37FA6A" box="[640,860,1508,1534]" italics="true" pageId="1" pageNumber="162">Benstonea celebica</emphasis>
</taxonomicName>
is a tree up to 
<quantity id="4CDACEC9FFCEBF2DFBAB55A9C829FA6B" box="[1044,1090,1508,1535]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="162" unit="m" value="5.0">5 m</quantity>
with a large spicate pendulous infructescence comprising up to 10 subspherical syncarps (c. 
<quantity id="4CDACEC9FFCEBF2DFC8A5645CFC8F9B7" box="[821,931,1544,1571]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="1" pageNumber="162" unit="cm" value="10.5" valueMax="11.0" valueMin="10.0">10–11 cm</quantity>
in diam) (
<figureCitation id="13197FA9FFCEBF2DFBA85645C804F9B6" box="[1047,1135,1544,1570]" captionStart="FIGURE 1" captionStartId="3.[136,229,1953,1975]" captionTargetBox="[210,1383,188,1915]" captionTargetId="figure-17@3.[188,1398,171,1940]" captionTargetPageId="3" captionText="FIGURE 1. A–D. Benstonea verruculosa. E–G. Benstonea stenocarpa. H. Benstonea celebica. A, E. Axillary syncarp. B. Peduncle covered by prophylls. C, D, F. Details of pileus and stigmas. G. Habit. H. Polysyncarpic pending infructescence. A–C: From Callmander &amp; Lasut 1551 (Photos: M. Callmander). F: Munzinger &amp; Bau 6745 (Photo: J. Munzinger). E, G. Bau s.n. (Photos: B. Bau). H. Purwanto &amp; al. PSU42 (Photo: Y. Purwanto &amp; R. Polosakan)." figureDoi="http://doi.org/10.5281/zenodo.5143879" httpUri="https://zenodo.org/record/5143879/files/figure.png" pageId="1" pageNumber="162">Fig. 1G</figureCitation>
). In the absence of adequate material of 
<taxonomicName id="4C2218AFFFCEBF2DFEB35660CDE2F9D2" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[268,393,1580,1606]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFEB35660CDE2F9D2" box="[268,393,1580,1606]" italics="true" pageId="1" pageNumber="162">B. celebica</emphasis>
</taxonomicName>
and apparently unaware of the presence of the photograph at FI, 
<bibRefCitation id="EFB31EDDFFCEBF2DFBCB5661C96CF9D2" author="Stone, B. C." box="[1140,1287,1580,1606]" pageId="1" pageNumber="162" pagination="205 - 212" refId="ref4141" refString="Stone, B. C. (1984) Some new and critical Pandanus species of subgenus Acrostigma, I. Supplement to Revisio Pandanacearum. Gardens' Bulletin Singapore 36: 205 - 212." type="journal article" year="1984">Stone (1984)</bibRefCitation>
was evidently oblivious to its highly distinctive morphology, and he described 
<taxonomicName id="4C2218AFFFCEBF2DFCD2561DC94CF9FE" authority="Stone (1984: 209)" authorityName="Stone" authorityPageNumber="209" authorityYear="1984" box="[877,1319,1616,1642]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="gladiator">
<emphasis id="B956BF3EFFCEBF2DFCD2561DC824F9FE" box="[877,1103,1616,1642]" italics="true" pageId="1" pageNumber="162">Pandanus gladiator</emphasis>
<bibRefCitation id="EFB31EDDFFCEBF2DFBE7561DC94CF9FE" author="Stone, B. C." box="[1112,1319,1616,1642]" pageId="1" pageNumber="162" pagination="205 - 212" refId="ref4141" refString="Stone, B. C. (1984) Some new and critical Pandanus species of subgenus Acrostigma, I. Supplement to Revisio Pandanacearum. Gardens' Bulletin Singapore 36: 205 - 212." type="journal article" year="1984">Stone (1984: 209)</bibRefCitation>
</taxonomicName>
based on a specimen collected from the same island, Sulawesi, 
<emphasis id="B956BF3EFFCEBF2DFD685639CF3EF91A" box="[727,853,1652,1678]" italics="true" pageId="1" pageNumber="162">Vogel 5287</emphasis>
. The latter species was erroneously considered to be a synonym of 
<taxonomicName id="4C2218AFFFCEBF2DFE9156D4CE43F926" authorityName="Callmander &amp; Buerki" baseAuthorityName="Solms-Laubach" baseAuthorityYear="1883" box="[302,552,1689,1714]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="stenocarpa">
<emphasis id="B956BF3EFFCEBF2DFE9156D4CE43F926" box="[302,552,1689,1714]" italics="true" pageId="1" pageNumber="162">Benstonea stenocarpa</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFB31EDDFFCEBF2DFDE956D5CF1BF926" author="Callmander, M. W. &amp; Lowry II, P. P. &amp; Forest, F. &amp; Devey, D. S. &amp; Beentje, H. &amp; Buerki, S." box="[598,880,1688,1714]" pageId="1" pageNumber="162" pagination="323 - 345" refId="ref3439" refString="Callmander, M. W., Lowry II, P. P., Forest, F., Devey, D. S., Beentje, H. &amp; Buerki, S. (2012) Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species. Candollea 67: 323 - 345." type="journal article" year="2012">Callmander &amp; al. (2012)</bibRefCitation>
and this error is corrected here. Original material of 
<taxonomicName id="4C2218AFFFCEBF2DFF1556F0CD41F942" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[170,298,1724,1750]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFF1556F0CD41F942" box="[170,298,1724,1750]" italics="true" pageId="1" pageNumber="162">B. celebica</emphasis>
</taxonomicName>
comprises only Warburg’s 
<typeStatus id="5499DD8EFFCEBF2DFDD356F0CEF7F943" box="[620,668,1725,1751]" pageId="1" pageNumber="162">type</typeStatus>
gathering from Sulawesi, which is highly fragmentary - consisting now of only of two incomplete leaf portions (at B and FI). The only known syncarp of this collection is believed to have been destroyed (at B), although fortunately a photograph is conserved at FI. 
<emphasis id="B956BF3EFFCEBF2DFB955749C8C3F88A" box="[1066,1192,1796,1822]" italics="true" pageId="1" pageNumber="162">Vogel 5287</emphasis>
(the 
<typeStatus id="5499DD8EFFCEBF2DFB5B5748C97FF88B" box="[1252,1300,1797,1823]" pageId="1" pageNumber="162">type</typeStatus>
of 
<taxonomicName id="4C2218AFFFCEBF2DFA845749CC9BF8D6" authorityName="Stone" authorityYear="1984" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="gladiator">
<emphasis id="B956BF3EFFCEBF2DFA845749CC9BF8D6" italics="true" pageId="1" pageNumber="162">Pandanus gladiator</emphasis>
</taxonomicName>
) is an excellent fertile collection from Central Sulawesi with duplicates in L, KLU and 
<collectingCountry id="F33523BCFFCEBF2DFABA5764C947F8D7" box="[1285,1324,1833,1859]" name="Philippines" pageId="1" pageNumber="162">PH</collectingCountry>
, each with leaves and representative portions of the infructescence. We consider this specimen to represent the same species as the 
<typeStatus id="5499DD8EFFCEBF2DFF0D573CCC89F81F" box="[178,226,1905,1931]" pageId="1" pageNumber="162">type</typeStatus>
of 
<taxonomicName id="4C2218AFFFCEBF2DFEB6573CCDEDF81E" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[265,390,1904,1930]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFEB6573CCDEDF81E" box="[265,390,1904,1930]" italics="true" pageId="1" pageNumber="162">B. celebica</emphasis>
</taxonomicName>
, and we therefore designate it as an epitype for the species. 
<taxonomicName id="4C2218AFFFCEBF2DFB8C573CC967F81E" authorityName="Callmander &amp; Buerki" baseAuthorityName="Warburg" baseAuthorityYear="1900" box="[1075,1292,1904,1930]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="162" phylum="Tracheophyta" rank="species" species="celebica">
<emphasis id="B956BF3EFFCEBF2DFB8C573CC967F81E" box="[1075,1292,1904,1930]" italics="true" pageId="1" pageNumber="162">Benstonea celebica</emphasis>
</taxonomicName>
has also been collected recently elsewhere on Sulawesi: Central Sulawesi Province, Pontolotua Hill and Sedoa in the vicinity of the Lore-Lindu National Park (
<bibRefCitation id="EFB31EDDFFCEBF2DFE0057F5CE22F847" author="Keim, A. P." box="[447,585,1976,2003]" pageId="1" pageNumber="162" refId="ref3609" refString="Keim, A. P. (2005) A joint Herbarium Bogoriense &amp; HerbariumWallacea exploration in the Lore Lindu National Park, Central Celebes (including Mount Nokilalaki &amp; Adjacent areas): Pandanaceae. Technical report, LIPI, Bogor, 18 pp." type="book" year="2005">Keim, 2005</bibRefCitation>
; 
<bibRefCitation id="EFB31EDDFFCEBF2DFDE957F5CF01F846" author="Keim, A. P. &amp; Rustiami, H." box="[598,874,1976,2002]" pageId="1" pageNumber="162" pagination="375 - 389" refId="ref3686" refString="Keim, A. P. &amp; Rustiami, H. (2007) Kenakeragaman suku Pandanaceae di pegunungan sekitar desa Sedoa, kawasan Taman Nasional Lore Lindu Kabupaten Poso-Propinsi Sulawesi Tengah. Berita Biologi 8: 375 - 389." type="journal article" year="2007">Keim &amp; Rustiami, 2007</bibRefCitation>
) and in Northern Sulawesi Province, Ambang Hill in the vicinity of Boganinani Wartabone National Park and Bolaang Mongondow, Minahassa (
<bibRefCitation id="EFB31EDDFFCEBF2DFB105791C9CCF862" author="Polosakan, R. &amp; Susiarti, S. &amp; Purwanto, Y." box="[1199,1447,2012,2038]" pageId="1" pageNumber="162" pagination="748 - 753" refId="ref3848" refString="Polosakan, R., Susiarti, S. &amp; Purwanto, Y. (2009) Habitat Pandanus di Gunung Ambang Taman Nasional Boganinani Wartabone dan pemanfaatan oleh masyarakat sekitarnya. In: Nuryanto, A. &amp; Budisantoso, I. (Eds.) Prosiding Seminar Nasional Biologi Peran Biosistematika dalam Pengelolaan Sumberdaya Hayati Indonesia. General Soedirman University, Purwokerto, pp. 748 - 753." type="book chapter" year="2009">Polosakan &amp; al. 2009</bibRefCitation>
; 
<bibRefCitation id="EFB31EDDFFCEBF2DFF37584CCD18F78E" author="Purwanto, Y. &amp; Polosakan, R. &amp; Susiarti, S." box="[136,371,2048,2075]" pageId="1" pageNumber="162" pagination="151 - 167" refId="ref3919" refString="Purwanto, Y., Polosakan, R. &amp; Susiarti, S. (2009) Studi etnobiologi Pandanaceae masyarakat Minahassa &amp; Bolaang Mongondow. Prosiding Seminar Nasional Etnobotani 4: 151 - 167." type="journal article" year="2009">Purwanto &amp; al. 2009</bibRefCitation>
) and is considered to be endemic to the island.
</paragraph>
</subSubSection>
<subSubSection id="C33830A7FFCDBF2EFF0250D3CFF8FEDC" pageId="2" pageNumber="163" type="materials_examined">
<paragraph id="8B9D632CFFCDBF2EFF0250D3CFF8FEDC" blockId="2.[136,1452,158,329]" pageId="2" pageNumber="163">
<emphasis id="B956BF3EFFCDBF2EFF0250D3CD94FF2C" bold="true" box="[189,511,158,184]" pageId="2" pageNumber="163">Additional specimens seen:</emphasis>
— 
<materialsCitation id="3B4A6971FFCDBF2EFDA350D3C89CFF48" collectingDate="2005-05-31" collectingDateMax="2005-09-11" collectingDateMin="2005-05-31" collectionCode="WAL, BO" country="Indonesia" county="Toro" elevation="900" location="Hill" municipality="Pono" pageId="2" pageNumber="163" specimenCount="1" stateProvince="Sulawesi Tengah">
<collectingCountry id="F33523BCFFCDBF2EFDA350D3CEDCFF2C" box="[540,695,158,184]" name="Indonesia" pageId="2" pageNumber="163">INDONESIA</collectingCountry>
. 
<collectingRegion id="49E6ADCEFFCDBF2EFD7F50D3CF80FF2C" box="[704,1003,158,184]" country="Indonesia" name="Sulawesi Tengah" pageId="2" pageNumber="163">Central Sulawesi Province</collectingRegion>
: 
<collectingRegion id="49E6ADCEFFCDBF2EFC4A50D3C836FF2C" box="[1013,1117,158,184]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="163">Sulawesi</collectingRegion>
, 
<collectingCounty id="62FC1BA0FFCDBF2EFBD950D2C8F6FF2D" box="[1126,1181,159,185]" pageId="2" pageNumber="163">Toro</collectingCounty>
, 
<collectingMunicipality id="6BF9F956FFCDBF2EFB1850D2C88AFF2D" box="[1191,1249,159,185]" pageId="2" pageNumber="163">Pono</collectingMunicipality>
(Pontolotua) 
<location id="8EFD35F7FFCDBF2EFAC650D3C9CCFF2C" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFAC650D3C9CCFF2C" box="[1401,1447,158,184]" country="Indonesia" county="Toro" municipality="Pono" name="Hill" pageId="2" pageNumber="163" stateProvince="Sulawesi Tengah">Hill</location>
, 
<quantity id="4CDACEC9FFCDBF2EFF37508FCCA4FF49" box="[136,207,194,221]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="163" unit="m" value="900.0">
<elevation id="000F841FFFCDBF2EFF37508FCCA4FF49" box="[136,207,194,221]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="163" unit="m" value="900.0">900 m</elevation>
</quantity>
, 
<date id="FF9C45ECFFCDBF2EFF66508FCD05FF49" box="[217,366,194,221]" pageId="2" pageNumber="163" value="2005-05-31">
<collectingDate id="EFD8BC04FFCDBF2EFF66508FCD05FF49" box="[217,366,194,221]" pageId="2" pageNumber="163" value="2005-05-31">31 May 2005</collectingDate>
</date>
, 
<emphasis id="B956BF3EFFCDBF2EFEC7508ECD89FF48" box="[376,482,194,220]" italics="true" pageId="2" pageNumber="163">Keim 558</emphasis>
(
<collectionCode id="ED33FBE9FFCDBF2EFE50508FCE58FF48" box="[495,563,194,220]" country="USA" httpUri="http://grbio.org/cool/vv8r-qmmj" name="Wadsworth Anaerobe Laboratory, Wadsworth Hospital Center" pageId="2" pageNumber="163">WAL</collectionCode>
); Poso, Sedoa, Siboala Hill, 
<date id="FF9C45ECFFCDBF2EFCD7508FC855FF49" box="[872,1086,194,221]" pageId="2" pageNumber="163" value="2005-09-11">
<collectingDate id="EFD8BC04FFCDBF2EFCD7508FC855FF49" box="[872,1086,194,221]" pageId="2" pageNumber="163" value="2005-09-11">11 September 2005</collectingDate>
</date>
, 
<emphasis id="B956BF3EFFCDBF2EFBF5508EC8DFFF48" box="[1098,1204,194,220]" italics="true" pageId="2" pageNumber="163">Keim 683</emphasis>
(
<collectionCode id="ED33FBE9FFCDBF2EFB7D508FC885FF48" box="[1218,1262,194,220]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="2" pageNumber="163" type="Herbarium">BO</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3B4A6971FFCDBF2EFABF508FCFFBFEDC" collectingDate="2006-03-27" collectingDateMax="2009-11-23" collectingDateMin="2006-03-27" collectionCode="WAL, BO" country="Indonesia" county="Mount Ambang Nature Reserve" elevation="1271" latitude="0.74583334" location="Kinho" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" pageId="2" pageNumber="163" specimenCode="PSU42" specimenCount="1" stateProvince="Sulawesi Utara">
<collectingRegion id="49E6ADCEFFCDBF2EFABF508FCC9BFE94" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="163">North Sulawesi Province</collectingRegion>
: 
<collectingRegion id="49E6ADCEFFCDBF2EFF4550ABCD09FE94" box="[250,354,230,256]" country="Indonesia" name="Sulawesi Utara" pageId="2" pageNumber="163">Sulawesi</collectingRegion>
, 
<collectingCounty id="62FC1BA0FFCDBF2EFED250AACEBEFE95" box="[365,725,230,257]" pageId="2" pageNumber="163">Mount Ambang Nature Reserve</collectingCounty>
, near 
<collectingMunicipality id="6BF9F956FFCDBF2EFCA850ABCFA5FE94" box="[791,974,230,256]" pageId="2" pageNumber="163">Kotulidak River</collectingMunicipality>
, 
<geoCoordinate id="EE1605EBFFCDBF2EFC6650ABC80EFE94" box="[985,1125,230,256]" degrees="00" direction="north" minutes="47" orientation="latitude" pageId="2" pageNumber="163" precision="15" seconds="29" value="0.7913889">00°47’29”N</geoCoordinate>
, 
<geoCoordinate id="EE1605EBFFCDBF2EFBCF50ABC96DFE94" box="[1136,1286,230,256]" degrees="124" direction="east" minutes="22" orientation="longitude" pageId="2" pageNumber="163" precision="15" seconds="39" value="124.3775">124°22’39”E</geoCoordinate>
, 
<date id="FF9C45ECFFCDBF2EFAAD50ABCCA8FEB0" pageId="2" pageNumber="163" value="2009-11-23">
<collectingDate id="EFD8BC04FFCDBF2EFAAD50ABCCA8FEB0" pageId="2" pageNumber="163" value="2009-11-23">23 November 2009</collectingDate>
</date>
, 
<emphasis id="B956BF3EFFCDBF2EFF6F5147CD7FFEB0" box="[208,276,266,292]" italics="true" pageId="2" pageNumber="163">
<location id="8EFD35F7FFCDBF2EFF6F5147CD7FFEB0" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFF6F5147CD7FFEB0" box="[208,276,266,292]" country="Indonesia" county="Mount Ambang Nature Reserve" latitude="0.74583334" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" name="Kinho" pageId="2" pageNumber="163" stateProvince="Sulawesi Utara">Kinho</location>
</emphasis>
267 (
<collectionCode id="ED33FBE9FFCDBF2EFEE75147CDF7FEB0" box="[344,412,266,292]" country="USA" httpUri="http://grbio.org/cool/vv8r-qmmj" name="Wadsworth Anaerobe Laboratory, Wadsworth Hospital Center" pageId="2" pageNumber="163">WAL</collectionCode>
); 
<location id="8EFD35F7FFCDBF2EFE0E5147CECCFEB0" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFE0E5147CECCFEB0" box="[433,679,266,292]" country="Indonesia" county="Mount Ambang Nature Reserve" latitude="0.74583334" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" name="Bolaang Mongondow" pageId="2" pageNumber="163" stateProvince="Sulawesi Utara">Bolaang Mongondow</location>
, 
<location id="8EFD35F7FFCDBF2EFD0B5147CF37FEB0" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFD0B5147CF37FEB0" box="[692,860,266,292]" country="Indonesia" county="Mount Ambang Nature Reserve" latitude="0.74583334" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" name="Kolangkangon" pageId="2" pageNumber="163" stateProvince="Sulawesi Utara">Kolangkangon</location>
, around 
<location id="8EFD35F7FFCDBF2EFC015147C8FEFEB0" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFC015147C8FEFEB0" box="[958,1173,266,292]" country="Indonesia" county="Mount Ambang Nature Reserve" latitude="0.74583334" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" name="Ambang mountain" pageId="2" pageNumber="163" stateProvince="Sulawesi Utara">Ambang mountain</location>
, 
<location id="8EFD35F7FFCDBF2EFB1E5147C965FEB0" LSID="urn:lsid:plazi:treatment:038BD23AFFCEBF2EFF3753C3CFF8FEDC:8EFD35F7FFCDBF2EFB1E5147C965FEB0" box="[1185,1294,266,292]" country="Indonesia" county="Mount Ambang Nature Reserve" latitude="0.74583334" longLatPrecision="21" longitude="124.43528" municipality="Kotulidak River" name="Modayag" pageId="2" pageNumber="163" stateProvince="Sulawesi Utara">Modayag</location>
, 
<geoCoordinate id="EE1605EBFFCDBF2EFAA45147C9CCFEB1" box="[1307,1447,266,293]" degrees="00" direction="north" minutes="44" orientation="latitude" pageId="2" pageNumber="163" precision="15" seconds="45" value="0.74583334">00°44’45”N</geoCoordinate>
, 
<geoCoordinate id="EE1605EBFFCDBF2EFF375163CD75FEDC" box="[136,286,302,328]" degrees="124" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="163" precision="15" seconds="07" value="124.43528">124°26’07”E</geoCoordinate>
, 
<quantity id="4CDACEC9FFCDBF2EFE955163CDEAFEDD" box="[298,385,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.271" pageId="2" pageNumber="163" unit="m" value="1271.0">
<elevation id="000F841FFFCDBF2EFE955163CDEAFEDD" box="[298,385,302,329]" metricMagnitude="3" metricUnit="m" metricValue="1.271" pageId="2" pageNumber="163" unit="m" value="1271.0">1271 m</elevation>
</quantity>
, 
<date id="FF9C45ECFFCDBF2EFE325163CE50FEDC" box="[397,571,302,328]" pageId="2" pageNumber="163" value="2006-03-27">
<collectingDate id="EFD8BC04FFCDBF2EFE325163CE50FEDC" box="[397,571,302,328]" pageId="2" pageNumber="163" value="2006-03-27">27 March 2006</collectingDate>
</date>
, 
<emphasis id="B956BF3EFFCDBF2EFDF95162CF20FEDC" box="[582,843,302,329]" italics="true" pageId="2" pageNumber="163">
Purwanto &amp; al. 
<specimenCode id="DB84CB57FFCDBF2EFD435163CF20FEDC" box="[764,843,302,328]" collectionCode="PSU" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="2" pageNumber="163" type="Museum">PSU42</specimenCode>
</emphasis>
(
<collectionCode id="ED33FBE9FFCDBF2EFCE45163CFECFEDC" box="[859,903,302,328]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="2" pageNumber="163" type="Herbarium">BO</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>