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<document id="20A5F79C2276FF38EB85496DAD04FE06" ID-CLB-Dataset="6627" ID-DOI="10.5281/zenodo.4648924" ID-GBIF-Dataset="1325ca6e-8ac3-4535-b243-0b5fa6b53c53" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648924" ID-ZooBank="3E4FA71B-1C3A-4BBD-A928-EF5A8344B419" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617136750815" checkinUser="carolina" docAuthor="Kovařík, František, Lowe, Graeme, Byronová, Markéta &amp; Šťáhlavský, František" docDate="2020" docId="E91C87A3DD3BE3051F1C437C0352FDFE" docLanguage="en" docName="Euscorpius.326.1-19.pdf" docOrigin="Euscorpius 326" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Euscorpius thracicus Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský 2020, sp. n." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="1525FFDBDD3FE3151E08453A0751FF9D" masterDocTitle="Euscorpius thracicus sp n (Scorpiones: Euscorpiidae) from Bulgaria" masterLastPageNumber="17" masterPageNumber="1" pageNumber="3" updateTime="1698935529255" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<figureCitation id="F98E2A30DD3BE3111F01435D06C9F91D" box="[265,408,1639,1664]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStart-2="Figures 7–16" captionStart-3="Figures 17–22" captionStart-4="Figures 23–31" captionStart-5="Figures 32–39" captionStart-6="Figures 40–43" captionStart-7="Figures 44–49" captionStartId-0="3.[113,190,2016,2038]" captionStartId-1="6.[115,192,1981,2003]" captionStartId-2="7.[115,192,1958,1980]" captionStartId-3="8.[116,193,1988,2010]" captionStartId-4="9.[115,192,1979,2001]" captionStartId-5="10.[113,190,1958,1980]" captionStartId-6="11.[116,193,1198,1220]" captionStartId-7="12.[115,192,1991,2013]" captionTargetBox-0="[113,1474,234,2005]" captionTargetBox-1="[138,1430,274,1952]" captionTargetBox-2="[113,1474,248,1887]" captionTargetBox-3="[127,1464,262,1953]" captionTargetBox-4="[129,1472,285,1928]" captionTargetBox-5="[125,1441,250,1915]" captionTargetBox-6="[113,1474,274,1183]" captionTargetBox-7="[151,1429,222,1983]" captionTargetId-0="figure-27@3.[113,1474,234,2007]" captionTargetId-1="figure-59@6.[116,1477,257,1971]" captionTargetId-2="figure-74@7.[113,1474,248,1911]" captionTargetId-3="figure-69@8.[113,1474,234,1973]" captionTargetId-4="figure-54@9.[115,1476,272,1960]" captionTargetId-5="figure-66@10.[113,1474,244,1951]" captionTargetId-6="figure-530@11.[113,1474,234,1184]" captionTargetId-7="figure-80@12.[113,1474,214,1991]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionText-0="Figures 1–2. E. thracicus sp. n., male (1) and female (2) paratypes in vivo habitus." captionText-1="Figures 3–6: E. thracicus sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm." captionText-2="Figures 7–16: E. thracicus sp. n. Figures 7, 9, 11–15. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16)." captionText-3="Figures 17–22: E. thracicus sp. n. Figures 17–19. Female paratype, metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Figures 20–22. Male holotype, metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Scale bars: 10 mm (17–19, 20–22)." captionText-4="Figures 23–31. E. thracicus sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31)." captionText-5="Figures 32–39. E. thracicus sp. n., female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (32–38)." captionText-6="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." captionText-7="Figures 44–49. Euscorpius thracicus sp. n., left hemispermatophore. Figures 44–45. Whole hemispermatophore in convex (44) and concave (45) views. Figures 46–49. Capsule of left hemispermatophore, 3-dimensional form displayed in cross stereoscopic pairs, in posterior-convex (46), anterior (47), convex (48) and concave (49) views. Paratype No. 1864. Scale bar: 1 mm (Figs. 44–49)." figureDoi-0="http://doi.org/10.5281/zenodo.4648926" figureDoi-1="http://doi.org/10.5281/zenodo.4648928" figureDoi-2="http://doi.org/10.5281/zenodo.4648930" figureDoi-3="http://doi.org/10.5281/zenodo.4648932" figureDoi-4="http://doi.org/10.5281/zenodo.4648934" figureDoi-5="http://doi.org/10.5281/zenodo.4648936" figureDoi-6="http://doi.org/10.5281/zenodo.4648938" figureDoi-7="http://doi.org/10.5281/zenodo.4648940" httpUri-0="https://zenodo.org/record/4648926/files/figure.png" httpUri-1="https://zenodo.org/record/4648928/files/figure.png" httpUri-2="https://zenodo.org/record/4648930/files/figure.png" httpUri-3="https://zenodo.org/record/4648932/files/figure.png" httpUri-4="https://zenodo.org/record/4648934/files/figure.png" httpUri-5="https://zenodo.org/record/4648936/files/figure.png" httpUri-6="https://zenodo.org/record/4648938/files/figure.png" httpUri-7="https://zenodo.org/record/4648940/files/figure.png" pageId="4" pageNumber="3">Figures 1–49</figureCitation>
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<figureCitation id="F98E2A30DD3BE3111FAC435D06B8F9E2" box="[420,489,1639,1663]" captionStart-0="Figures50–58" captionStart-1="Figures 59–60" captionStart-2="Figure 61" captionStartId-0="13.[115,192,1982,2004]" captionStartId-1="14.[116,193,2015,2037]" captionStartId-2="15.[116,184,886,908]" captionTargetBox-0="[115,1472,214,1975]" captionTargetBox-1="[113,1474,268,2003]" captionTargetBox-2="[113,1474,234,877]" captionTargetId-0="figure-136@13.[113,1474,211,1977]" captionTargetId-1="figure-35@14.[113,1474,232,2006]" captionTargetId-2="figure-360@15.[113,1474,234,880]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." captionText-1="Figures 59–60. E. thracicus sp. n., type locality, Bulgaria, Krumovgrad, Bryagovets, 41.6485862°N 25.8132185°E." captionText-2="Figure 61. Map of distribution of Euscorpius populations analysed by DNA (see Table 2)." figureDoi-0="http://doi.org/10.5281/zenodo.4648942" figureDoi-1="http://doi.org/10.5281/zenodo.4648944" figureDoi-2="http://doi.org/10.5281/zenodo.4648946" httpUri-0="https://zenodo.org/record/4648942/files/figure.png" httpUri-1="https://zenodo.org/record/4648944/files/figure.png" httpUri-2="https://zenodo.org/record/4648946/files/figure.png" pageId="4" pageNumber="3">58–61</figureCitation>
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<tableCitation id="2C37030EDD3BE3111FFB435D0539F91D" box="[499,616,1639,1664]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="5.[115,171,993,1014]" captionStartId-1="15.[112,168,1430,1451]" captionTargetBox-1="[117,1471,936,1402]" captionTargetId-0="graphics-542@5.[113,1483,237,975]" captionTargetId-1="graphics-361@15.[111,1476,929,1407]" captionTargetPageId-0="5" captionTargetPageId-1="15" captionText-0="Table 1. Comparative measurements of adults of Euscorpius thracicus sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 2. A matrix of genetic distances among the COI genes of the Euscorpius species from Bulgaria, eastern Serbia, and northern Greece. p-distance (below the diagonal); Kimura 2-parameter distance (above the diagonal). In the brackets are mentioned specimen codes/accession number from GenBank." httpUri-0="http://table.plazi.org/id/35CA663DDD3AE3101E7B46DB055EFB8E" httpUri-1="http://table.plazi.org/id/35CA663DDD30E31A1E7840AC0607FA7B" pageId="4" pageNumber="3" tableUuid-0="35CA663DDD3AE3101E7B46DB055EFB8E" tableUuid-1="35CA663DDD30E31A1E7840AC0607FA7B">Tables 1–2</tableCitation>
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(
<figureCitation id="F98E2A30DD3BE3111E70429D07AFF822" box="[120,254,1959,1983]" captionStart="Figures 59–60" captionStartId="14.[116,193,2015,2037]" captionTargetBox="[113,1474,268,2003]" captionTargetId="figure-35@14.[113,1474,232,2006]" captionTargetPageId="14" captionText="Figures 59–60. E. thracicus sp. n., type locality, Bulgaria, Krumovgrad, Bryagovets, 41.6485862°N 25.8132185°E." figureDoi="http://doi.org/10.5281/zenodo.4648944" httpUri="https://zenodo.org/record/4648944/files/figure.png" pageId="4" pageNumber="3">Figs. 59–60</figureCitation>
, 
<specimenCount id="77B3FD3CDD3BE3111F0742920660F85D" box="[271,305,1960,1984]" pageId="4" pageNumber="3" type="male">
<figureCitation id="F98E2A30DD3BE3111F0742920648F85D" box="[271,281,1960,1984]" captionStart="Figures 1–2" captionStartId="3.[113,190,2016,2038]" captionTargetBox="[113,1474,234,2005]" captionTargetId="figure-27@3.[113,1474,234,2007]" captionTargetPageId="3" captionText="Figures 1–2. E. thracicus sp. n., male (1) and female (2) paratypes in vivo habitus." figureDoi="http://doi.org/10.5281/zenodo.4648926" httpUri="https://zenodo.org/record/4648926/files/figure.png" pageId="4" pageNumber="3">1</figureCitation>
♂
</specimenCount>
(
<typeStatus id="BE0E8817DD3BE3111F4D429D06F2F822" box="[325,419,1959,1983]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
) 
<specimenCount id="77B3FD3CDD3BE3111FB0429D068FF822" box="[440,478,1959,1983]" pageId="4" pageNumber="3" type="male">6♂</specimenCount>
<specimenCount id="77B3FD3CDD3BE3111FD5429D06ABF822" box="[477,506,1959,1983]" pageId="4" pageNumber="3" type="female">8♀</specimenCount>
(
<typeStatus id="BE0E8817DD3BE3111C0742920526F85D" box="[527,631,1960,1984]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
, 
<collectorName id="CC405363DD3BE3111C80429205EBF822" box="[648,698,1960,1983]" pageId="4" pageNumber="3">Nos.</collectorName>
1864, 1865, 1866, 1878, 1879, 1882, 1883), 
<date id="150B1075DD3BE3111C0A42FD0580F87D" box="[514,721,1991,2016]" pageId="4" pageNumber="3" value="2020-09-25">
<collectingDate id="054FE99DDD3BE3111C0A42FD0580F87D" box="[514,721,1991,2016]" pageId="4" pageNumber="3" value="2020-09-25">25 September 2020</collectingDate>
</date>
, leg. 
<collectorName id="CC405363DD3BE3111E7842D207AAF79D" box="[112,251,2024,2048]" pageId="4" pageNumber="3">M. Byronová</collectorName>
et al.,
</materialsCitation>
<collectionCode id="07A4AE70DD3BE3111F4942D206D2F79D" box="[321,387,2024,2048]" pageId="4" pageNumber="3">FKCP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="29AF653EDD3BE3111D2745D1027AFEBE" pageId="4" pageNumber="3" type="etymology">
<paragraph id="610A36B5DD3BE3111D2745D1027AFEBE" blockId="4.[815,1473,235,291]" pageId="4" pageNumber="3">
ETYMOLOGY. Named after Thracia (Thrace), the ancient name of the region where the 
<typeStatus id="BE0E8817DD3BE3111A254431030AFEBE" box="[1069,1115,267,291]" pageId="4" pageNumber="3">type</typeStatus>
locality is situated.
</paragraph>
</subSubSection>
<subSubSection id="29AF653EDD3BE3111D2744710363FCFE" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="610A36B5DD3BE3111D2744710363FCFE" blockId="4.[815,1474,331,867]" pageId="4" pageNumber="3">
DIAGNOSIS (
<specimenCount id="77B3FD3CDD3BE3111DB04471049EFEFE" box="[952,975,331,355]" pageId="4" pageNumber="3" type="male">♂</specimenCount>
<specimenCount id="77B3FD3CDD3BE3111DC74471048EFEFE" box="[975,991,331,355]" pageId="4" pageNumber="3" type="female">♀</specimenCount>
). Total length 
<quantity id="A64D9B50DD3BE3111A9D44710245FEFF" box="[1173,1300,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="4" pageNumber="3" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
. Color reddish yellow to brown, telson yellow. Pedipalp patella external trichobothria numbers: 4 
<emphasis id="53C1EAA7DD3BE3111A3F44B10301FE3E" box="[1079,1104,395,419]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111A6344B103D5FE3E" box="[1131,1156,395,419]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
<subScript id="FD3134F0DD3BE3111A8C44A103DAFE34" attach="left" box="[1156,1163,411,425]" fontSize="6" pageId="4" pageNumber="3">a</subScript>
, 2 
<emphasis id="53C1EAA7DD3BE3111AAE44B1039BFE3E" box="[1190,1226,395,419]" italics="true" pageId="4" pageNumber="3">esb</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111AEC44B10252FE3F" box="[1252,1283,395,418]" italics="true" pageId="4" pageNumber="3">em</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111B1644B1026DFE3F" box="[1310,1340,395,418]" italics="true" pageId="4" pageNumber="3">est</emphasis>
, 6 
<emphasis id="53C1EAA7DD3BE3111B5E44B10238FE3F" box="[1366,1385,395,418]" italics="true" pageId="4" pageNumber="3">et</emphasis>
, ventral aspect of patella with 7 trichobothria. Pectinal teeth number 
<quantity id="A64D9B50DD3BE3111D2744F1042FFE7E" box="[815,894,459,483]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="4" pageNumber="3" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
males, 
<quantity id="A64D9B50DD3BE3111DC544F1035EFE7E" box="[973,1039,459,483]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.286" metricValueMin="1.524" pageId="4" pageNumber="3" unit="in" value="7.5" valueMax="9.0" valueMin="6.0">6–9 in</quantity>
females. Chelicerae yellow, very slightly reticulated. Male with pedipalp finger marginal profile 
<typeStatus id="BE0E8817DD3BE3111B7244D102F9FD9E" box="[1402,1448,491,515]" pageId="4" pageNumber="3">type</typeStatus>
C (KovařÍk &amp; Šťáhlavský, 2020: 2), female with fingers very weakly undulate, almost linear. Dorsal metasomal carinae on segments III– IV irregularly granulated, mainly in male. Dorsolateral carinae on metasomal segments II–IV absent. Ventrolateral carinae on metasomal segments II–IV present or indicated and smooth. Metasoma V ventrally with median carina indicated. Metasoma finely granulated dorsally in male and smooth in female. Chela length/width ratio, 
<quantity id="A64D9B50DD3BE3111B3247D00290FC9E" box="[1338,1473,746,771]" metricMagnitude="-2" metricUnit="m" metricValue="6.1341" metricValueMax="6.2992" metricValueMin="5.969" pageId="4" pageNumber="3" unit="in" value="2.415" valueMax="2.48" valueMin="2.35">2.35–2.48 in</quantity>
male, 
<quantity id="A64D9B50DD3BE3111D7B463004AAFCBE" box="[883,1019,778,803]" metricMagnitude="-2" metricUnit="m" metricValue="6.4135" metricValueMax="6.476999999999999" metricValueMin="6.35" pageId="4" pageNumber="3" unit="in" value="2.525" valueMax="2.55" valueMin="2.5">2.50–2.55 in</quantity>
female. Metasoma IV length/width ratio, 
<quantity id="A64D9B50DD3BE3111D27461104CEFCDE" box="[815,927,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="4" pageNumber="3" unit="in" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 in</quantity>
both sexes. Telson length/depth ratio, 
<quantity id="A64D9B50DD3BE3111B5946100290FCDE" box="[1361,1473,810,835]" metricMagnitude="-2" metricUnit="m" metricValue="6.476999999999999" metricValueMax="6.604" metricValueMin="6.35" pageId="4" pageNumber="3" unit="in" value="2.55" valueMax="2.6" valueMin="2.5">2.5–2.6 in</quantity>
male, 
<quantity id="A64D9B50DD3BE3111D784671048DFCFE" box="[880,988,843,867]" metricMagnitude="-2" metricUnit="m" metricValue="8.001" metricValueMax="8.128" metricValueMin="7.874" pageId="4" pageNumber="3" unit="in" value="3.15" valueMax="3.2" valueMin="3.1">3.1–3.2 in</quantity>
female.
</paragraph>
</subSubSection>
<subSubSection id="29AF653EDD3BE3051D2746B1067DF9FE" lastPageId="16" lastPageNumber="15" pageId="4" pageNumber="3" type="description">
<paragraph id="610A36B5DD3BE3111D2746B104FCFBBE" blockId="4.[815,1474,907,2057]" pageId="4" pageNumber="3">
DESCRIPTION (
<specimenCount id="77B3FD3CDD3BE3111DC146B10488FC3E" box="[969,985,907,931]" pageId="4" pageNumber="3" type="female">♀</specimenCount>
<specimenCount id="77B3FD3CDD3BE3111DD146B104A1FC3E" box="[985,1008,907,931]" pageId="4" pageNumber="3" type="male">♂</specimenCount>
). Total length 
<quantity id="A64D9B50DD3BE3111A8246B103AAFC3F" box="[1162,1275,907,931]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="4" pageNumber="3" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
in both sexes. The habitus is shown in 
<figureCitation id="F98E2A30DD3BE3111A1846910328FC5E" box="[1040,1145,939,963]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStartId-0="3.[113,190,2016,2038]" captionStartId-1="6.[115,192,1981,2003]" captionTargetBox-0="[113,1474,234,2005]" captionTargetBox-1="[138,1430,274,1952]" captionTargetId-0="figure-27@3.[113,1474,234,2007]" captionTargetId-1="figure-59@6.[116,1477,257,1971]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionText-0="Figures 1–2. E. thracicus sp. n., male (1) and female (2) paratypes in vivo habitus." captionText-1="Figures 3–6: E. thracicus sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4648926" figureDoi-1="http://doi.org/10.5281/zenodo.4648928" httpUri-0="https://zenodo.org/record/4648926/files/figure.png" httpUri-1="https://zenodo.org/record/4648928/files/figure.png" pageId="4" pageNumber="3">Figs. 1–6</figureCitation>
. For position and distribution of trichobothria on pedipalps, see 
<figureCitation id="F98E2A30DD3BE3111ABD46F1026DFC7E" box="[1205,1340,971,995]" captionStart="Figures 32–39" captionStartId="10.[113,190,1958,1980]" captionTargetBox="[125,1441,250,1915]" captionTargetId="figure-66@10.[113,1474,244,1951]" captionTargetPageId="10" captionText="Figures 32–39. E. thracicus sp. n., female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (32–38)." figureDoi="http://doi.org/10.5281/zenodo.4648936" httpUri="https://zenodo.org/record/4648936/files/figure.png" pageId="4" pageNumber="3">Figs. 32–38</figureCitation>
. For sexual dimorphism, see below in the description. For measurements, see 
<tableCitation id="2C37030EDD3BE3111D5F413104F6FBBE" box="[855,935,1035,1059]" captionStart="Table 1" captionStartId="5.[115,171,993,1014]" captionTargetId="graphics-542@5.[113,1483,237,975]" captionTargetPageId="5" captionText="Table 1. Comparative measurements of adults of Euscorpius thracicus sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/35CA663DDD3AE3101E7B46DB055EFB8E" pageId="4" pageNumber="3" tableUuid="35CA663DDD3AE3101E7B46DB055EFB8E">Table 1</tableCitation>
.
</paragraph>
<paragraph id="610A36B5DD3BE3111D274110023CFB1E" blockId="4.[815,1474,907,2057]" pageId="4" pageNumber="3">
<emphasis id="53C1EAA7DD3BE3111D27411004FBFBDF" bold="true" box="[815,938,1066,1090]" pageId="4" pageNumber="3">Coloration</emphasis>
(
<figureCitation id="F98E2A30DD3BE3111DB241110370FBDE" box="[954,1057,1067,1091]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStartId-0="3.[113,190,2016,2038]" captionStartId-1="6.[115,192,1981,2003]" captionTargetBox-0="[113,1474,234,2005]" captionTargetBox-1="[138,1430,274,1952]" captionTargetId-0="figure-27@3.[113,1474,234,2007]" captionTargetId-1="figure-59@6.[116,1477,257,1971]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionText-0="Figures 1–2. E. thracicus sp. n., male (1) and female (2) paratypes in vivo habitus." captionText-1="Figures 3–6: E. thracicus sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4648926" figureDoi-1="http://doi.org/10.5281/zenodo.4648928" httpUri-0="https://zenodo.org/record/4648926/files/figure.png" httpUri-1="https://zenodo.org/record/4648928/files/figure.png" pageId="4" pageNumber="3">Figs. 1–6</figureCitation>
). Base color uniformly reddish yellow to brown including sternites, telson yellow, pedipalps reddish brown. Chelicerae yellow and very slightly reticulate.
</paragraph>
<paragraph id="610A36B5DD3BE3111D2741B004C0FA7E" blockId="4.[815,1474,907,2057]" pageId="4" pageNumber="3">
<emphasis id="53C1EAA7DD3BE3111D2741B0030CFB3E" bold="true" box="[815,1117,1162,1187]" pageId="4" pageNumber="3">Carapace and mesosoma</emphasis>
(
<figureCitation id="F98E2A30DD3BE3111A7E41B103A4FB3E" box="[1142,1269,1163,1187]" captionStart="Figures 7–16" captionStartId="7.[115,192,1958,1980]" captionTargetBox="[113,1474,248,1887]" captionTargetId="figure-74@7.[113,1474,248,1911]" captionTargetPageId="7" captionText="Figures 7–16: E. thracicus sp. n. Figures 7, 9, 11–15. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16)." figureDoi="http://doi.org/10.5281/zenodo.4648930" httpUri="https://zenodo.org/record/4648930/files/figure.png" pageId="4" pageNumber="3">Figs. 7–10</figureCitation>
). Carapace finely granulated with several smooth areas; carinae absent. Anterior margin of carapace straight. Carapace with two lateral eyes. Tergites finely granulated, more so in male, without carinae developed. Tergite VII lacking median and paired lateral carinae. Sternites III–VII smooth and lustrous; VII lacking median and paired lateral carinae. Stigmata small, narrow ellipical. Pectinal teeth number 8–10 (8 x 8, 4 x 9, 2 x 10) in males and 6–9 (1 x 6, 12 x 7, 2 x 8, 1 x 9) in females, fulcra present. Pectines with 3 marginal lamellae and 4–6 middle lamellae.
</paragraph>
<paragraph id="610A36B5DD3BE3111D2740D10342F81E" blockId="4.[815,1474,907,2057]" pageId="4" pageNumber="3">
<emphasis id="53C1EAA7DD3BE3111D2740D1034EF99E" bold="true" box="[815,1055,1515,1539]" pageId="4" pageNumber="3">Metasoma and telson</emphasis>
(
<figureCitation id="F98E2A30DD3BE3111A2440D103FFF99F" box="[1068,1198,1514,1539]" captionStart-0="Figures 7–16" captionStart-1="Figures 17–22" captionStartId-0="7.[115,192,1958,1980]" captionStartId-1="8.[116,193,1988,2010]" captionTargetBox-0="[113,1474,248,1887]" captionTargetBox-1="[127,1464,262,1953]" captionTargetId-0="figure-74@7.[113,1474,248,1911]" captionTargetId-1="figure-69@8.[113,1474,234,1973]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 7–16: E. thracicus sp. n. Figures 7, 9, 11–15. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16)." captionText-1="Figures 17–22: E. thracicus sp. n. Figures 17–19. Female paratype, metasoma and telson lateral (17), ventral (18), and dorsal (19) views. Figures 20–22. Male holotype, metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Scale bars: 10 mm (17–19, 20–22)." figureDoi-0="http://doi.org/10.5281/zenodo.4648930" figureDoi-1="http://doi.org/10.5281/zenodo.4648932" httpUri-0="https://zenodo.org/record/4648930/files/figure.png" httpUri-1="https://zenodo.org/record/4648932/files/figure.png" pageId="4" pageNumber="3">Figs. 15–22</figureCitation>
). Metasoma very sparsely hirsute and smooth. Metasoma I–V finely granulated in male, several fine granules also present on lateral surfaces of metasoma I and V in both sexes. Dorsal carinae on metasomal segments I–V irregularly granulated in male, reduced in female; dorsolateral carinae absent; ventrolateral carinae present or indicated and smooth on segments II–IV, granulated on segment V. Metasoma V ventrally granulated with median carina indicated, metasoma I–IV with ventral median carinae absent.Anal arch with small pigmented granules. Telson rather smooth, elongate in female and swollen in male, with annular ring indicated in female and developed in male. Aculeus short, more curved in male.
</paragraph>
<paragraph id="610A36B5DD3BE3101D2742B0063AFA17" blockId="4.[815,1474,907,2057]" lastBlockId="5.[113,772,1074,2026]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
<emphasis id="53C1EAA7DD3BE3111D2742B004CFF83F" bold="true" box="[815,926,1930,1954]" pageId="4" pageNumber="3">Pedipalps</emphasis>
(
<figureCitation id="F98E2A30DD3BE3111DBC42B1036EF83E" box="[948,1087,1931,1955]" captionStart-0="Figures 23–31" captionStart-1="Figures 32–39" captionStartId-0="9.[115,192,1979,2001]" captionStartId-1="10.[113,190,1958,1980]" captionTargetBox-0="[129,1472,285,1928]" captionTargetBox-1="[125,1441,250,1915]" captionTargetId-0="figure-54@9.[115,1476,272,1960]" captionTargetId-1="figure-66@10.[113,1474,244,1951]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figures 23–31. E. thracicus sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31)." captionText-1="Figures 32–39. E. thracicus sp. n., female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (32–38)." figureDoi-0="http://doi.org/10.5281/zenodo.4648934" figureDoi-1="http://doi.org/10.5281/zenodo.4648936" httpUri-0="https://zenodo.org/record/4648934/files/figure.png" httpUri-1="https://zenodo.org/record/4648936/files/figure.png" pageId="4" pageNumber="3">Figs. 23–39</figureCitation>
). Pedipalps very sparsely hirsute. Patella with 24 (4 
<emphasis id="53C1EAA7DD3BE3111A0842910348F85E" box="[1024,1049,1963,1987]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111A3342910305F85E" box="[1083,1108,1963,1987]" italics="true" pageId="4" pageNumber="3">eb</emphasis>
<subScript id="FD3134F0DD3BE3111A5C4281030AF854" attach="left" box="[1108,1115,1979,1993]" fontSize="6" pageId="4" pageNumber="3">a</subScript>
, 2 
<emphasis id="53C1EAA7DD3BE3111A75429103F0F85E" box="[1149,1185,1963,1987]" italics="true" pageId="4" pageNumber="3">esb</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111ACA429103B0F85F" box="[1218,1249,1963,1986]" italics="true" pageId="4" pageNumber="3">em</emphasis>
, 4 
<emphasis id="53C1EAA7DD3BE3111B0A42910271F85F" box="[1282,1312,1963,1986]" italics="true" pageId="4" pageNumber="3">est</emphasis>
, 6 
<emphasis id="53C1EAA7DD3BE3111B4942910205F85F" box="[1345,1364,1963,1986]" italics="true" pageId="4" pageNumber="3">et</emphasis>
) external and 7 ventral trichobothria. Chela with 4 trichobothria in ventral series, of which 
<emphasis id="53C1EAA7DD3BE3111A7C42D103D5F79F" box="[1140,1156,2027,2050]" italics="true" pageId="4" pageNumber="3">V</emphasis>
<subScript id="FD3134F0DD3BE3111A8C42C103DDF794" attach="left" box="[1156,1164,2043,2057]" fontSize="6" pageId="4" pageNumber="3">4</subScript>
is located external to the ventroexternal carina, on the external surface near 
<emphasis id="53C1EAA7DD3AE3101CDF410805A4FBD7" box="[727,757,1074,1098]" italics="true" pageId="5" pageNumber="4">Eb</emphasis>
<subScript id="FD3134F0DD3AE3101CFD417805ACFBCD" attach="left" box="[757,765,1090,1104]" fontSize="6" pageId="5" pageNumber="4">1</subScript>
. Entire femur finely granulated and patella smooth with fine granulated dorsointernal and ventrointernal margins. Femur with granulated developed carinae; ventroexternal carina incomplete. Patella with 5 complete carinae including irregular wide externomedian carina. Dorsal patellar spur well developed in male. Manus dorsally with fine, rounded granules, which do not form a median carina (mainly in male); only five chelal carinae developed. Male with pedipalp finger undulation profile 
<typeStatus id="BE0E8817DD3AE3101F374068063CFAF7" box="[319,365,1362,1386]" pageId="5" pageNumber="4">type</typeStatus>
C, female with fingers very weakly undulate, almost linear.
</paragraph>
<paragraph id="610A36B5DD3AE3101D0D45C2027CFC55" pageId="5" pageNumber="4">
<table id="13B5C415DD3A1CEA1E7F45CD02F8FC56" box="[119,1449,247,971]" gridcols="4" gridrows="19" pageId="5" pageNumber="4">
<tr id="DF8534F7DD3A1CEA1E7F45CD02F8FE8D" box="[119,1449,247,272]" gridrow="0" pageId="5" pageNumber="4" rowspan-0="1" rowspan-1="1">
<th id="9C545D8BDD3A1CEA1D0D45CD0364FE8D" box="[773,1077,247,272]" gridcol="2" gridrow="0" pageId="5" pageNumber="4">
<taxonomicName id="A6B54D36DD3AE3101D0D45C204BCFE92" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[773,1005,247,272]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD3AE3101D0D45C204BCFE92" bold="true" box="[773,1005,247,272]" italics="true" pageId="5" pageNumber="4">Euscorpius thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD3AE3101DF145C20364FE92" box="[1017,1077,248,272]" pageId="5" pageNumber="4" rank="species">
<emphasis id="53C1EAA7DD3AE3101DF145C20343FE8D" bold="true" box="[1017,1042,248,272]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="53C1EAA7DD3AE3101A1745C2037FFE8D" bold="true" box="[1055,1070,248,272]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
</th>
<th id="9C545D8BDD3A1CEA1A7145CD02F8FE8D" box="[1145,1449,247,272]" gridcol="3" gridrow="0" pageId="5" pageNumber="4">
<taxonomicName id="A6B54D36DD3AE3101A7145C20230FE92" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1145,1377,247,272]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD3AE3101A7145C20230FE92" bold="true" box="[1145,1377,247,272]" italics="true" pageId="5" pageNumber="4">Euscorpius thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD3AE3101B6445C202F8FE92" box="[1388,1449,248,272]" pageId="5" pageNumber="4" rank="species">
<emphasis id="53C1EAA7DD3AE3101B6445C202D4FE8D" bold="true" box="[1388,1413,248,272]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="53C1EAA7DD3AE3101B9B45C202F3FE8D" bold="true" box="[1427,1442,248,272]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
</th>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F442402F8FEA5" box="[119,1449,286,312]" gridrow="1" pageId="5" pageNumber="4" rowspan-1="1">
<th id="9C545D8BDD3A1CEA1E7F44240615FEA5" box="[119,324,286,312]" gridcol="0" gridrow="1" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101E7F441A066CFEA5" bold="true" box="[119,317,288,312]" pageId="5" pageNumber="4">Dimensions (mm)</emphasis>
</th>
<td id="9C545D8BDD3A1CEA1D0D44240364FEA5" box="[773,1077,286,312]" gridcol="2" gridrow="1" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101D564424048AFEA5" bold="true" box="[862,987,286,312]" pageId="5" pageNumber="4">♂ holotype</emphasis>
</td>
<td id="9C545D8BDD3A1CEA1A71442402F8FEA5" box="[1145,1449,286,312]" gridcol="3" gridrow="1" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101AD844240200FEA5" bold="true" box="[1232,1361,286,312]" pageId="5" pageNumber="4">♀ paratype</emphasis>
</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F447F02F8FEC2" box="[119,1449,325,351]" gridrow="2" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F447F0615FEC2" box="[119,324,325,351]" gridcol="0" gridrow="2" pageId="5" pageNumber="4">Carapace</th>
<td id="9C545D8BDD3A1CEA1C32447F05CEFEC2" box="[570,671,325,351]" gridcol="1" gridrow="2" pageId="5" pageNumber="4">L / W</td>
<td id="9C545D8BDD3A1CEA1D0D447F0364FEC2" box="[773,1077,325,351]" gridcol="2" gridrow="2" pageId="5" pageNumber="4">4.13 / 3.97</td>
<td id="9C545D8BDD3A1CEA1A71447F02F8FEC2" box="[1145,1449,325,351]" gridcol="3" gridrow="2" pageId="5" pageNumber="4">4.00 / 4.00</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F445602F8FE18" box="[119,1449,364,389]" gridrow="3" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F44560615FE18" box="[119,324,364,389]" gridcol="0" gridrow="3" pageId="5" pageNumber="4">Mesosoma</th>
<td id="9C545D8BDD3A1CEA1C32445605CEFE18" box="[570,671,364,389]" gridcol="1" gridrow="3" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D44560364FE18" box="[773,1077,364,389]" gridcol="2" gridrow="3" pageId="5" pageNumber="4">9.92</td>
<td id="9C545D8BDD3A1CEA1A71445602F8FE18" box="[1145,1449,364,389]" gridcol="3" gridrow="3" pageId="5" pageNumber="4">13.58</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F44A902F8FE30" box="[119,1449,403,429]" gridrow="4" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F44A90615FE30" box="[119,324,403,429]" gridcol="0" gridrow="4" pageId="5" pageNumber="4">Tergite VII</th>
<td id="9C545D8BDD3A1CEA1C3244A905CEFE30" box="[570,671,403,429]" gridcol="1" gridrow="4" pageId="5" pageNumber="4">L / W</td>
<td id="9C545D8BDD3A1CEA1D0D44A90364FE30" box="[773,1077,403,429]" gridcol="2" gridrow="4" pageId="5" pageNumber="4">1.73 / 3.31</td>
<td id="9C545D8BDD3A1CEA1A7144A902F8FE30" box="[1145,1449,403,429]" gridcol="3" gridrow="4" pageId="5" pageNumber="4">2.23 / 3.73</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F448302F8FE4E" box="[119,1449,441,467]" gridrow="5" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F44830615FE4E" box="[119,324,441,467]" gridcol="0" gridrow="5" pageId="5" pageNumber="4">Metasoma + telson</th>
<td id="9C545D8BDD3A1CEA1C32448305CEFE4E" box="[570,671,441,467]" gridcol="1" gridrow="5" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D44830364FE4E" box="[773,1077,441,467]" gridcol="2" gridrow="5" pageId="5" pageNumber="4">15.26</td>
<td id="9C545D8BDD3A1CEA1A71448302F8FE4E" box="[1145,1449,441,467]" gridcol="3" gridrow="5" pageId="5" pageNumber="4">13.00</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F44DA02F8FE67" box="[119,1449,480,506]" gridrow="6" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F44DA0615FE67" box="[119,324,480,506]" gridcol="0" gridrow="6" pageId="5" pageNumber="4">Segment I</th>
<td id="9C545D8BDD3A1CEA1C3244DA05CEFE67" box="[570,671,480,506]" gridcol="1" gridrow="6" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D44DA0364FE67" box="[773,1077,480,506]" gridcol="2" gridrow="6" pageId="5" pageNumber="4">1.61 / 1.51 / 1.23</td>
<td id="9C545D8BDD3A1CEA1A7144DA02F8FE67" box="[1145,1449,480,506]" gridcol="3" gridrow="6" pageId="5" pageNumber="4">1.32 / 1.42 / 1.27</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F473D02F8FDBC" box="[119,1449,519,545]" gridrow="7" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F473D0615FDBC" box="[119,324,519,545]" gridcol="0" gridrow="7" pageId="5" pageNumber="4">Segment II</th>
<td id="9C545D8BDD3A1CEA1C32473D05CEFDBC" box="[570,671,519,545]" gridcol="1" gridrow="7" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D473D0364FDBC" box="[773,1077,519,545]" gridcol="2" gridrow="7" pageId="5" pageNumber="4">1.76 / 1.33 / 1.21</td>
<td id="9C545D8BDD3A1CEA1A71473D02F8FDBC" box="[1145,1449,519,545]" gridcol="3" gridrow="7" pageId="5" pageNumber="4">1.54 / 1.20 / 1.13</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F471702F8FDDA" box="[119,1449,557,583]" gridrow="8" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F47170615FDDA" box="[119,324,557,583]" gridcol="0" gridrow="8" pageId="5" pageNumber="4">Segment III</th>
<td id="9C545D8BDD3A1CEA1C32471705CEFDDA" box="[570,671,557,583]" gridcol="1" gridrow="8" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D47170364FDDA" box="[773,1077,557,583]" gridcol="2" gridrow="8" pageId="5" pageNumber="4">1.90 / 1.25 / 1.18</td>
<td id="9C545D8BDD3A1CEA1A71471702F8FDDA" box="[1145,1449,557,583]" gridcol="3" gridrow="8" pageId="5" pageNumber="4">1.66 / 1.10 / 1.13</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F476E02F8FDF3" box="[119,1449,596,622]" gridrow="9" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F476E0615FDF3" box="[119,324,596,622]" gridcol="0" gridrow="9" pageId="5" pageNumber="4">Segment IV</th>
<td id="9C545D8BDD3A1CEA1C32476E05CEFDF3" box="[570,671,596,622]" gridcol="1" gridrow="9" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D476E0364FDF3" box="[773,1077,596,622]" gridcol="2" gridrow="9" pageId="5" pageNumber="4">2.23 / 1.17 / 1.18</td>
<td id="9C545D8BDD3A1CEA1A71476E02F8FDF3" box="[1145,1449,596,622]" gridcol="3" gridrow="9" pageId="5" pageNumber="4">1.97 / 1.06 / 1.08</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F474102F8FD08" box="[119,1449,635,661]" gridrow="10" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F47410615FD08" box="[119,324,635,661]" gridcol="0" gridrow="10" pageId="5" pageNumber="4">Segment V</th>
<td id="9C545D8BDD3A1CEA1C32474105CEFD08" box="[570,671,635,661]" gridcol="1" gridrow="10" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D47410364FD08" box="[773,1077,635,661]" gridcol="2" gridrow="10" pageId="5" pageNumber="4">3.54 / 1.15 / 1.23</td>
<td id="9C545D8BDD3A1CEA1A71474102F8FD08" box="[1145,1449,635,661]" gridcol="3" gridrow="10" pageId="5" pageNumber="4">3.09 / 1.00 / 1.01</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F479B02F8FD21" box="[119,1449,673,700]" gridrow="11" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F479B0615FD21" box="[119,324,673,700]" gridcol="0" gridrow="11" pageId="5" pageNumber="4">Telson</th>
<td id="9C545D8BDD3A1CEA1C32479B05CEFD21" box="[570,671,673,700]" gridcol="1" gridrow="11" pageId="5" pageNumber="4">L / W / D</td>
<td id="9C545D8BDD3A1CEA1D0D479B0364FD21" box="[773,1077,673,700]" gridcol="2" gridrow="11" pageId="5" pageNumber="4">4.22 / 1.64 / 1.63</td>
<td id="9C545D8BDD3A1CEA1A71479B02F8FD21" box="[1145,1449,673,700]" gridcol="3" gridrow="11" pageId="5" pageNumber="4">3.42 / 1.07 / 1.06</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F47F202F8FD7F" box="[119,1449,712,738]" gridrow="12" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F47F20615FD7F" box="[119,324,712,738]" gridcol="0" gridrow="12" pageId="5" pageNumber="4">Pedipalp</th>
<td id="9C545D8BDD3A1CEA1C3247F205CEFD7F" box="[570,671,712,738]" gridcol="1" gridrow="12" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D47F20364FD7F" box="[773,1077,712,738]" gridcol="2" gridrow="12" pageId="5" pageNumber="4">14.37</td>
<td id="9C545D8BDD3A1CEA1A7147F202F8FD7F" box="[1145,1449,712,738]" gridcol="3" gridrow="12" pageId="5" pageNumber="4">13.85</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F47D502F8FC94" box="[119,1449,751,777]" gridrow="13" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F47D50615FC94" box="[119,324,751,777]" gridcol="0" gridrow="13" pageId="5" pageNumber="4">Femur</th>
<td id="9C545D8BDD3A1CEA1C3247D505CEFC94" box="[570,671,751,777]" gridcol="1" gridrow="13" pageId="5" pageNumber="4">L / W</td>
<td id="9C545D8BDD3A1CEA1D0D47D50364FC94" box="[773,1077,751,777]" gridcol="2" gridrow="13" pageId="5" pageNumber="4">3.52 / 1.34</td>
<td id="9C545D8BDD3A1CEA1A7147D502F8FC94" box="[1145,1449,751,777]" gridcol="3" gridrow="13" pageId="5" pageNumber="4">3.38 / 1.32</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F462C02F8FCAD" box="[119,1449,790,816]" gridrow="14" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F462C0615FCAD" box="[119,324,790,816]" gridcol="0" gridrow="14" pageId="5" pageNumber="4">Patella</th>
<td id="9C545D8BDD3A1CEA1C32462C05CEFCAD" box="[570,671,790,816]" gridcol="1" gridrow="14" pageId="5" pageNumber="4">L / W</td>
<td id="9C545D8BDD3A1CEA1D0D462C0364FCAD" box="[773,1077,790,816]" gridcol="2" gridrow="14" pageId="5" pageNumber="4">3.49 / 1.48</td>
<td id="9C545D8BDD3A1CEA1A71462C02F8FCAD" box="[1145,1449,790,816]" gridcol="3" gridrow="14" pageId="5" pageNumber="4">3.44 / 1.44</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F460602F8FCCB" box="[119,1449,828,854]" gridrow="15" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F46060615FCCB" box="[119,324,828,854]" gridcol="0" gridrow="15" pageId="5" pageNumber="4">Chela</th>
<td id="9C545D8BDD3A1CEA1C32460605CEFCCB" box="[570,671,828,854]" gridcol="1" gridrow="15" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D46060364FCCB" box="[773,1077,828,854]" gridcol="2" gridrow="15" pageId="5" pageNumber="4">7.36</td>
<td id="9C545D8BDD3A1CEA1A71460602F8FCCB" box="[1145,1449,828,854]" gridcol="3" gridrow="15" pageId="5" pageNumber="4">7.03</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F465902F8FCE0" box="[119,1449,867,893]" gridrow="16" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F46590615FCE0" box="[119,324,867,893]" gridcol="0" gridrow="16" pageId="5" pageNumber="4">Manus</th>
<td id="9C545D8BDD3A1CEA1C32465905CEFCE0" box="[570,671,867,893]" gridcol="1" gridrow="16" pageId="5" pageNumber="4">W / D</td>
<td id="9C545D8BDD3A1CEA1D0D46590364FCE0" box="[773,1077,867,893]" gridcol="2" gridrow="16" pageId="5" pageNumber="4">2.98 / 2.15</td>
<td id="9C545D8BDD3A1CEA1A71465902F8FCE0" box="[1145,1449,867,893]" gridcol="3" gridrow="16" pageId="5" pageNumber="4">2.78 / 1.77</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F46B002F8FC39" box="[119,1449,906,932]" gridrow="17" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F46B00615FC39" box="[119,324,906,932]" gridcol="0" gridrow="17" pageId="5" pageNumber="4">Movable finger</th>
<td id="9C545D8BDD3A1CEA1C3246B005CEFC39" box="[570,671,906,932]" gridcol="1" gridrow="17" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D46B00364FC39" box="[773,1077,906,932]" gridcol="2" gridrow="17" pageId="5" pageNumber="4">4.18</td>
<td id="9C545D8BDD3A1CEA1A7146B002F8FC39" box="[1145,1449,906,932]" gridcol="3" gridrow="17" pageId="5" pageNumber="4">3.86</td>
</tr>
<tr id="DF8534F7DD3A1CEA1E7F468A02F8FC56" box="[119,1449,944,971]" gridrow="18" pageId="5" pageNumber="4">
<th id="9C545D8BDD3A1CEA1E7F468A0615FC56" box="[119,324,944,971]" gridcol="0" gridrow="18" pageId="5" pageNumber="4">Total</th>
<td id="9C545D8BDD3A1CEA1C32468A05CEFC56" box="[570,671,944,971]" gridcol="1" gridrow="18" pageId="5" pageNumber="4">L</td>
<td id="9C545D8BDD3A1CEA1D0D468A0364FC56" box="[773,1077,944,971]" gridcol="2" gridrow="18" pageId="5" pageNumber="4">29.31</td>
<td id="9C545D8BDD3A1CEA1A71468A02F8FC56" box="[1145,1449,944,971]" gridcol="3" gridrow="18" pageId="5" pageNumber="4">30.58</td>
</tr>
</table>
</paragraph>
<caption id="35CA663DDD3AE3101E7B46DB055EFB8E" ID-Table-UUID="35CA663DDD3AE3101E7B46DB055EFB8E" httpUri="http://table.plazi.org/id/35CA663DDD3AE3101E7B46DB055EFB8E" pageId="5" pageNumber="4" startId="5.[115,171,993,1014]" targetBox="[119,1449,247,971]" targetIsTable="true" targetPageId="5">
<paragraph id="610A36B5DD3AE3101E7B46DB055EFB8E" blockId="5.[115,1360,992,1043]" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101E7B46DB07EFFC68" bold="true" box="[115,190,992,1014]" pageId="5" pageNumber="4">Table 1</emphasis>
. Comparative measurements of adults of 
<taxonomicName id="A6B54D36DD3AE3101C5C46DB0471FC6B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[596,800,992,1014]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD3AE3101C5C46DB0471FC6B" box="[596,800,992,1014]" italics="true" pageId="5" pageNumber="4">Euscorpius thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD3AE3101D2F46DB040FFC6B" box="[807,862,993,1014]" pageId="5" pageNumber="4" rank="species">
<emphasis id="53C1EAA7DD3AE3101D2F46DB046FFC6B" bold="true" box="[807,830,993,1014]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="53C1EAA7DD3AE3101D4346DB0409FC6B" bold="true" box="[843,856,993,1014]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
</paragraph>
</caption>
<paragraph id="610A36B5DD3AE3101E7940A806F6F997" blockId="5.[113,772,1074,2026]" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101E7940A807F7FA37" bold="true" box="[113,166,1426,1450]" pageId="5" pageNumber="4">Legs</emphasis>
(
<figureCitation id="F98E2A30DD3AE3101EB340A80612FA37" box="[187,323,1426,1450]" captionStart="Figures 7–16" captionStartId="7.[115,192,1958,1980]" captionTargetBox="[113,1474,248,1887]" captionTargetId="figure-74@7.[113,1474,248,1911]" captionTargetPageId="7" captionText="Figures 7–16: E. thracicus sp. n. Figures 7, 9, 11–15. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16)." figureDoi="http://doi.org/10.5281/zenodo.4648930" httpUri="https://zenodo.org/record/4648930/files/figure.png" pageId="5" pageNumber="4">Figs. 11–14</figureCitation>
). Both pedal spurs present on all legs, lacking spinelets; tibial spurs absent. Tarsus with single row of spinules on ventral surface, terminating distally with two essentially adjacent spinules.
</paragraph>
<paragraph id="610A36B5DD3AE3101E79432B03C2FA37" blockId="5.[113,772,1074,2026]" lastBlockId="5.[816,1477,1074,2026]" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101E79432B060DF9B4" bold="true" box="[113,348,1553,1577]" pageId="5" pageNumber="4">Hemispermatophore</emphasis>
(
<figureCitation id="F98E2A30DD3AE3101F7E43280554F9B7" box="[374,517,1554,1578]" captionStart-0="Figures 44–49" captionStart-1="Figures50–58" captionStartId-0="12.[115,192,1991,2013]" captionStartId-1="13.[115,192,1982,2004]" captionTargetBox-0="[151,1429,222,1983]" captionTargetBox-1="[115,1472,214,1975]" captionTargetId-0="figure-80@12.[113,1474,214,1991]" captionTargetId-1="figure-136@13.[113,1474,211,1977]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="Figures 44–49. Euscorpius thracicus sp. n., left hemispermatophore. Figures 44–45. Whole hemispermatophore in convex (44) and concave (45) views. Figures 46–49. Capsule of left hemispermatophore, 3-dimensional form displayed in cross stereoscopic pairs, in posterior-convex (46), anterior (47), convex (48) and concave (49) views. Paratype No. 1864. Scale bar: 1 mm (Figs. 44–49)." captionText-1="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi-0="http://doi.org/10.5281/zenodo.4648940" figureDoi-1="http://doi.org/10.5281/zenodo.4648942" httpUri-0="https://zenodo.org/record/4648940/files/figure.png" httpUri-1="https://zenodo.org/record/4648942/files/figure.png" pageId="5" pageNumber="4">Figs. 44–58</figureCitation>
). Lamelliform. Shape and proportions typical of the genus (Fet &amp; Soleglad, 2002). Distal lamina proximally broad, tapered distally, with strong basal constriction to half maximal width at base. Apex of lamina narrow but blunt, bent posteriorly. Capsule broad, with transverse trough on convex surface. Distal margin of trough strengthened with transverse rib that curves and connects to posterior axial rib of distal lamina. Truncal flexure well developed, separating trunk from capsule. Trunk relatively short, ca. 2/3 of length of distal lamina, gradually tapered towards base. Pedicel almost as long as trunk. Basal carina of capsule weakly sclerotized, terminating in typical crownlike structure (
<emphasis id="53C1EAA7DD3AE3101F1142A80666F837" box="[281,311,1938,1962]" italics="true" pageId="5" pageNumber="4">cls</emphasis>
) with 4–7 hook-like tines. Distal carina of capsule strongly sclerotized, distal external lobe (
<emphasis id="53C1EAA7DD3AE3101CD2428805AAF857" box="[730,763,1970,1994]" italics="true" pageId="5" pageNumber="4">lde</emphasis>
) apparently divided into two cristate ‘sub-lobes’, one broader and one narrower, separated by a deep cleft. Distal internal lobe (
<emphasis id="53C1EAA7DD3AE3101D67416804DAFBF7" box="[879,907,1106,1130]" italics="true" pageId="5" pageNumber="4">ldi</emphasis>
) apparently absent. Basal lobe (
<emphasis id="53C1EAA7DD3AE3101AE2416803AEFBF7" box="[1258,1279,1106,1130]" italics="true" pageId="5" pageNumber="4">bl</emphasis>
) absent. Terminal membrane of sperm duct with numerous fine spicules. Both left and right hemispermatophores from 3 individuals (
<typeStatus id="BE0E8817DD3AE3101B5241A80293FB37" box="[1370,1474,1170,1194]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
No. 1864, 1865 and 1866) displayed similar morphology. Measurements of left hemispermatophore No. 1864 (mm): distal lamina length from base of basal constriction to apex 2.50, maximal width 0.78; trunk length from truncal flexure to pedicel 1.34; pedicel length 1.06. Variation: 
<emphasis id="53C1EAA7DD3AE3101B2040080217FAD7" box="[1320,1350,1330,1354]" italics="true" pageId="5" pageNumber="4">cls</emphasis>
tine counts (with bifid tines counting as 2, trifid as 3, etc.) varied among 3 examined 
<typeStatus id="BE0E8817DD3AE3101D9740480358FA17" box="[927,1033,1394,1418]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
: No. 1864, right 4, left 7; No. 1865, right 4, left 5; No. 1866, right 6, left 4.
</paragraph>
<paragraph id="610A36B5DD3AE3101D384088040AF817" blockId="5.[816,1477,1074,2026]" pageId="5" pageNumber="4">
<emphasis id="53C1EAA7DD3AE3101D38408804F8FA57" bold="true" box="[816,937,1458,1482]" pageId="5" pageNumber="4">Karyotype</emphasis>
(
<figureCitation id="F98E2A30DD3AE3101DC84088031BFA57" box="[960,1098,1458,1482]" captionStart="Figures 40–43" captionStartId="11.[116,193,1198,1220]" captionTargetBox="[113,1474,274,1183]" captionTargetId="figure-530@11.[113,1474,234,1184]" captionTargetPageId="11" captionText="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="5" pageNumber="4">Figs. 40–43</figureCitation>
). We analyzed the chromosomes of four 
<typeStatus id="BE0E8817DD3AE3101D8F40E804B2FA77" box="[903,995,1490,1514]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
males (Nos. 1864, 1965, 1866, 1878). The diploid complement of these samples is composed of 92 chromosomes (
<figureCitation id="F98E2A30DD3AE3101DD94328031EF9B7" box="[977,1103,1554,1578]" captionStart="Figures 40–43" captionStartId="11.[116,193,1198,1220]" captionTargetBox="[113,1474,274,1183]" captionTargetId="figure-530@11.[113,1474,234,1184]" captionTargetPageId="11" captionText="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="5" pageNumber="4">Figs. 40, 41</figureCitation>
). In all observed postpachytene and metaphase I nuclei, we observed 46 bivalents without visible chiasmata (
<figureCitation id="F98E2A30DD3AE3101DA3436804AFF9F7" box="[939,1022,1618,1642]" captionStart="Figures 40–43" captionStartId="11.[116,193,1198,1220]" captionTargetBox="[113,1474,274,1183]" captionTargetId="figure-530@11.[113,1474,234,1184]" captionTargetPageId="11" captionText="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="5" pageNumber="4">Fig. 42</figureCitation>
). The relative length of the chromosomes of the diploid set decreased gradually from 2.23% to 0.37%. The chromosomes exhibited monocentric organization and the morphology of all chromosome pairs was telocentric with only one exception. The chromosomes of the pair number 18 were metacentric. The arm ratio of chromosomes belonging to this pair was 1.17 (the standard deviation is 0.12). These two biarmed chromosomes were clearly visible during postpachytene (
<figureCitation id="F98E2A30DD3AE3101DD142680378F8F7" box="[985,1065,1874,1898]" captionStart="Figures 40–43" captionStartId="11.[116,193,1198,1220]" captionTargetBox="[113,1474,274,1183]" captionTargetId="figure-530@11.[113,1474,234,1184]" captionTargetPageId="11" captionText="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="5" pageNumber="4">Fig. 42</figureCitation>
) as well as during metaphase II (
<figureCitation id="F98E2A30DD3AE3101B9F4268041FF817" captionStart="Figures 40–43" captionStartId="11.[116,193,1198,1220]" captionTargetBox="[113,1474,274,1183]" captionTargetId="figure-530@11.[113,1474,234,1184]" captionTargetPageId="11" captionText="Figures 40–43. Chromosomes of E. thracicus sp. n., karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="5" pageNumber="4">Fig. 43</figureCitation>
).
</paragraph>
<paragraph id="610A36B5DD3AE31E1D3842A80684F9AB" blockId="5.[816,1477,1074,2026]" lastBlockId="11.[116,774,1278,2038]" lastPageId="11" lastPageNumber="10" pageId="5" pageNumber="4">
The number of chromosomes in 
<taxonomicName id="A6B54D36DD3AE3101A9042A8024AF837" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1176,1307,1938,1962]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD3AE3101A9042A8024AF837" box="[1176,1307,1938,1962]" italics="true" pageId="5" pageNumber="4">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD3AE3101B2C42A80232F834" box="[1316,1379,1938,1962]" pageId="5" pageNumber="4" rank="species">
<emphasis id="53C1EAA7DD3AE3101B2C42A8026CF837" bold="true" box="[1316,1341,1938,1962]" pageId="5" pageNumber="4">sp</emphasis>
. 
<emphasis id="53C1EAA7DD3AE3101B4542A8020DF837" bold="true" box="[1357,1372,1938,1962]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
(2n=92) fully corresponds to the known 2n values of two 
<taxonomicName id="A6B54D36DD3AE3101B4242880293F854" authorityName="Thorell" authorityYear="1876" box="[1354,1474,1970,1993]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD3AE3101B4242880293F854" box="[1354,1474,1970,1993]" italics="true" pageId="5" pageNumber="4">Euscorpius</emphasis>
</taxonomicName>
species from the Balkan region (
<taxonomicName id="A6B54D36DD3AE3101AA542E8024AF874" authorityName="Kovarik &amp; Stahlavsky" authorityYear="2020" box="[1197,1307,2002,2025]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="janstai">
<emphasis id="53C1EAA7DD3AE3101AA542E8024AF874" box="[1197,1307,2002,2025]" italics="true" pageId="5" pageNumber="4">E. janstai</emphasis>
</taxonomicName>
from North 
<collectingCountry id="19A27625DD34E31E1E7C41C407BEFA8B" box="[116,239,1278,1302]" name="Macedonia" pageId="11" pageNumber="10">Macedonia</collectingCountry>
: 2n=112 and 
<taxonomicName id="A6B54D36DD34E31E1F9D41C5055CFA8B" authorityName="Kovarik &amp; Stahlavsky" authorityYear="2020" box="[405,525,1278,1302]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="sadileki">
<emphasis id="53C1EAA7DD34E31E1F9D41C5055CFA8B" box="[405,525,1278,1302]" italics="true" pageId="11" pageNumber="10">E. sadileki</emphasis>
</taxonomicName>
from 
<collectingCountry id="19A27625DD34E31E1C5641C405F7FA8B" box="[606,678,1278,1302]" name="Serbia" pageId="11" pageNumber="10">Serbia</collectingCountry>
: 2n=90) (KovařÍk &amp; Šťáhlavský, 2020). However, the karyotypes of these species differ conspicuously from that of 
<taxonomicName id="A6B54D36DD34E31E1C69400505B1FACB" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[609,736,1342,1366]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1C69400505B1FACB" box="[609,736,1342,1366]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1CEE400407DBFAE8" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1CEE400405AEFACB" bold="true" box="[742,767,1342,1366]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1E7C406407D2FAEB" bold="true" box="[116,131,1374,1398]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
by the presence of a higher amount of biarmed chromosomes in their karyotypes. Our cytogenetic results from the Balkan Peninsula document a high karyotype variability in the genus 
<taxonomicName id="A6B54D36DD34E31E1E7C408507BDFA4B" authorityName="Thorell" authorityYear="1876" box="[116,236,1471,1494]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD34E31E1E7C408507BDFA4B" box="[116,236,1471,1494]" italics="true" pageId="11" pageNumber="10">Euscorpius</emphasis>
</taxonomicName>
in this region. We support the application of this marker in taxonomy of the family 
<taxonomicName id="A6B54D36DD34E31E1FF240E405D9FA6B" authorityName="Laurie" authorityYear="1896" box="[506,648,1502,1526]" class="Arachnida" family="Euscorpiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="family">Euscorpiidae</taxonomicName>
, as already implemented in the genus 
<taxonomicName id="A6B54D36DD34E31E1F9240C40599F98B" authorityName="Koch" authorityYear="1837" box="[410,712,1534,1558]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD34E31E1F9240C40570F98B" box="[410,545,1534,1558]" italics="true" pageId="11" pageNumber="10">Alpiscorpius</emphasis>
(Euscorpiidae)
</taxonomicName>
from the Alps (
<bibRefCitation id="05244B44DD34E31E1ED24324069BF9AB" author="STUNDLOVA" box="[218,458,1566,1592]" pageId="11" pageNumber="10" pagination="152 - 163" refId="ref7942" refString="STUNDLOVA, J., J. SMID, P. NGUYEN &amp; F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">Štundlová et al., 2019</bibRefCitation>
).
</paragraph>
<caption id="35CA663DDD39E3131E7B428706ABF872" ID-DOI="http://doi.org/10.5281/zenodo.4648928" ID-Zenodo-Dep="4648928" httpUri="https://zenodo.org/record/4648928/files/figure.png" pageId="6" pageNumber="5" startId="6.[115,192,1981,2003]" targetBox="[138,1430,274,1952]" targetPageId="6">
<paragraph id="610A36B5DD39E3131E7B428706ABF872" blockId="6.[115,1473,1981,2032]" pageId="6" pageNumber="5">
<emphasis id="53C1EAA7DD39E3131E7B428707BBF84F" bold="true" box="[115,234,1981,2003]" pageId="6" pageNumber="5">Figures 3–6</emphasis>
: 
<taxonomicName id="A6B54D36DD39E3131EF04287063DF84F" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[248,364,1981,2002]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD39E3131EF04287063DF84F" box="[248,364,1981,2002]" italics="true" pageId="6" pageNumber="5">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD39E3131F7B428706FBF84F" box="[371,426,1981,2002]" pageId="6" pageNumber="5" rank="species">
<emphasis id="53C1EAA7DD39E3131F7B428706DBF84F" bold="true" box="[371,394,1981,2002]" pageId="6" pageNumber="5">sp</emphasis>
. 
<emphasis id="53C1EAA7DD39E3131F9E428706F2F84F" bold="true" box="[406,419,1981,2002]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="53C1EAA7DD39E3131FB842870579F84F" bold="true" box="[432,552,1981,2003]" pageId="6" pageNumber="5">Figures 3–4</emphasis>
. Male holotype in dorsal (3) and ventral (4) views. 
<emphasis id="53C1EAA7DD39E3131A17428703C6F84F" bold="true" box="[1055,1175,1981,2003]" pageId="6" pageNumber="5">Figures 5–6</emphasis>
. Female paratype in dorsal (5) and ventral (6) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="35CA663DDD38E3121E7B429C0510F86B" ID-DOI="http://doi.org/10.5281/zenodo.4648930" ID-Zenodo-Dep="4648930" httpUri="https://zenodo.org/record/4648930/files/figure.png" pageId="7" pageNumber="6" startId="7.[115,192,1958,1980]" targetBox="[113,1474,248,1887]" targetPageId="7">
<paragraph id="610A36B5DD38E3121E7B429C0510F86B" blockId="7.[115,1473,1958,2038]" pageId="7" pageNumber="6">
<emphasis id="53C1EAA7DD38E3121E7B429C07A4F826" bold="true" box="[115,245,1958,1980]" pageId="7" pageNumber="6">Figures 7–16</emphasis>
: 
<taxonomicName id="A6B54D36DD38E3121F09429D0625F826" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[257,372,1958,1980]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD38E3121F09429D0625F826" box="[257,372,1958,1980]" italics="true" pageId="7" pageNumber="6">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD38E3121F72429C06E1F821" box="[378,432,1958,1980]" pageId="7" pageNumber="6" rank="species">
<emphasis id="53C1EAA7DD38E3121F72429C06C0F826" bold="true" box="[378,401,1958,1979]" pageId="7" pageNumber="6">sp</emphasis>
. 
<emphasis id="53C1EAA7DD38E3121F95429C06FBF826" bold="true" box="[413,426,1958,1979]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="53C1EAA7DD38E3121FBD429C0545F826" bold="true" box="[437,532,1958,1980]" pageId="7" pageNumber="6">Figures 7</emphasis>
, 
<emphasis id="53C1EAA7DD38E3121C17429C057AF826" bold="true" box="[543,555,1958,1979]" pageId="7" pageNumber="6">9</emphasis>
, 
<emphasis id="53C1EAA7DD38E3121C3F429C0523F826" bold="true" box="[567,626,1958,1980]" pageId="7" pageNumber="6">11–15</emphasis>
. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). 
<emphasis id="53C1EAA7DD38E3121CAA42F90450F845" bold="true" box="[674,769,1987,2009]" pageId="7" pageNumber="6">Figures 8</emphasis>
, 
<emphasis id="53C1EAA7DD38E3121D0442F90475F845" bold="true" box="[780,804,1987,2008]" pageId="7" pageNumber="6">10</emphasis>
, 
<emphasis id="53C1EAA7DD38E3121D3842F90419F845" bold="true" box="[816,840,1987,2008]" pageId="7" pageNumber="6">16</emphasis>
. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16).
</paragraph>
</caption>
<caption id="35CA663DDD37E31D1E7C42FE02C7F86A" ID-DOI="http://doi.org/10.5281/zenodo.4648932" ID-Zenodo-Dep="4648932" httpUri="https://zenodo.org/record/4648932/files/figure.png" pageId="8" pageNumber="7" startId="8.[116,193,1988,2010]" targetBox="[127,1464,262,1953]" targetPageId="8">
<paragraph id="610A36B5DD37E31D1E7C42FE02C7F86A" blockId="8.[116,1475,1988,2039]" pageId="8" pageNumber="7">
<emphasis id="53C1EAA7DD37E31D1E7C42FE0655F844" bold="true" box="[116,260,1988,2010]" pageId="8" pageNumber="7">Figures 17–22</emphasis>
: 
<taxonomicName id="A6B54D36DD37E31D1F1B42FF06D9F844" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[275,392,1988,2010]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD37E31D1F1B42FF06D9F844" box="[275,392,1988,2010]" italics="true" pageId="8" pageNumber="7">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD37E31D1F9842FE0696F847" box="[400,455,1988,2010]" pageId="8" pageNumber="7" rank="species">
<emphasis id="53C1EAA7DD37E31D1F9842FE06F6F844" bold="true" box="[400,423,1988,2009]" pageId="8" pageNumber="7">sp</emphasis>
. 
<emphasis id="53C1EAA7DD37E31D1FBC42FE0690F844" bold="true" box="[436,449,1988,2009]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
<emphasis id="53C1EAA7DD37E31D1FC742FE050EF844" bold="true" box="[463,607,1988,2010]" pageId="8" pageNumber="7">Figures 17–19</emphasis>
. Female paratype, metasoma and telson lateral (17), ventral (18), and dorsal (19) views. 
<emphasis id="53C1EAA7DD37E31D1E7C42DB0652F86B" bold="true" box="[116,259,2017,2039]" pageId="8" pageNumber="7">Figures 20–22</emphasis>
. Male holotype, metasoma and telson lateral (20), ventral (21), and dorsal (22) views. Scale bars: 10 mm (17–19, 20–22).
</paragraph>
</caption>
<caption id="35CA663DDD36E31C1E7B42810207F873" ID-DOI="http://doi.org/10.5281/zenodo.4648934" ID-Zenodo-Dep="4648934" httpUri="https://zenodo.org/record/4648934/files/figure.png" pageId="9" pageNumber="8" startId="9.[115,192,1979,2001]" targetBox="[129,1472,285,1928]" targetPageId="9">
<paragraph id="610A36B5DD36E31C1E7B42810207F873" blockId="9.[115,1473,1979,2030]" pageId="9" pageNumber="8">
<emphasis id="53C1EAA7DD36E31C1E7B42810653F84D" bold="true" box="[115,258,1979,2001]" pageId="9" pageNumber="8">Figures 23–31</emphasis>
. 
<taxonomicName id="A6B54D36DD36E31C1F07428606D2F84D" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[271,387,1979,2001]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD36E31C1F07428606D2F84D" box="[271,387,1979,2001]" italics="true" pageId="9" pageNumber="8">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD36E31C1F82428106EEF84D" box="[394,447,1979,2000]" pageId="9" pageNumber="8" rank="species">
<emphasis id="53C1EAA7DD36E31C1F82428106F0F84D" bold="true" box="[394,417,1979,2000]" pageId="9" pageNumber="8">sp</emphasis>
. 
<emphasis id="53C1EAA7DD36E31C1FA5428106EBF84D" bold="true" box="[429,442,1979,2000]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31).
</paragraph>
</caption>
<caption id="35CA663DDD35E31F1E79429C0658F86B" ID-DOI="http://doi.org/10.5281/zenodo.4648936" ID-Zenodo-Dep="4648936" httpUri="https://zenodo.org/record/4648936/files/figure.png" pageId="10" pageNumber="9" startId="10.[113,190,1958,1980]" targetBox="[125,1441,250,1915]" targetPageId="10">
<paragraph id="610A36B5DD35E31F1E79429C0658F86B" blockId="10.[113,1472,1958,2038]" pageId="10" pageNumber="9">
<emphasis id="53C1EAA7DD35E31F1E79429C0651F826" bold="true" box="[113,256,1958,1980]" pageId="10" pageNumber="9">Figures 32–39</emphasis>
. 
<taxonomicName id="A6B54D36DD35E31F1F04429D062EF826" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[268,383,1958,1980]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD35E31F1F04429D062EF826" box="[268,383,1958,1980]" italics="true" pageId="10" pageNumber="9">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD35E31F1F8D429C06EBF821" box="[389,442,1958,1980]" pageId="10" pageNumber="9" rank="species">
<emphasis id="53C1EAA7DD35E31F1F8D429C06CDF826" bold="true" box="[389,412,1958,1979]" pageId="10" pageNumber="9">sp</emphasis>
. 
<emphasis id="53C1EAA7DD35E31F1FA0429C06E4F826" bold="true" box="[424,437,1958,1979]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (32–38).
</paragraph>
</caption>
<caption id="35CA663DDD34E31E1E7C419405E8FB7C" ID-DOI="http://doi.org/10.5281/zenodo.4648938" ID-Zenodo-Dep="4648938" httpUri="https://zenodo.org/record/4648938/files/figure.png" pageId="11" pageNumber="10" startId="11.[116,193,1198,1220]" targetBox="[113,1474,274,1183]" targetPageId="11">
<paragraph id="610A36B5DD34E31E1E7C419405E8FB7C" blockId="11.[116,1472,1198,1249]" pageId="11" pageNumber="10">
<emphasis id="53C1EAA7DD34E31E1E7C4194065BFB5E" bold="true" box="[116,266,1198,1220]" pageId="11" pageNumber="10">Figures 40–43.</emphasis>
Chromosomes of 
<taxonomicName id="A6B54D36DD34E31E1FB24195057FFB59" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[442,558,1198,1220]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1FB24195057FFB59" box="[442,558,1198,1220]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1C3C41950538FB59" box="[564,617,1199,1220]" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1C3C4195051AFB59" bold="true" box="[564,587,1199,1220]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1C5F41950535FB59" bold="true" box="[599,612,1199,1220]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, karyotype based on postpachytene (2n=92) (40), mitotic metaphase (41), postpachytene (42), one sister metaphase II (n=46) (43). Scale bars: 10 μm.
</paragraph>
</caption>
<paragraph id="610A36B5DD34E31E1E7C43040587F868" blockId="11.[116,774,1278,2038]" pageId="11" pageNumber="10">
<emphasis id="53C1EAA7DD34E31E1E7C43040641F9CB" bold="true" box="[116,272,1598,1622]" pageId="11" pageNumber="10">DNA analysis</emphasis>
(
<tableCitation id="2C37030EDD34E31E1F2943040629F9CB" box="[289,376,1598,1622]" captionStart="Table 2" captionStartId="15.[112,168,1430,1451]" captionTargetBox="[117,1471,936,1402]" captionTargetId="graphics-361@15.[111,1476,929,1407]" captionTargetPageId="15" captionText="Table 2. A matrix of genetic distances among the COI genes of the Euscorpius species from Bulgaria, eastern Serbia, and northern Greece. p-distance (below the diagonal); Kimura 2-parameter distance (above the diagonal). In the brackets are mentioned specimen codes/accession number from GenBank." httpUri="http://table.plazi.org/id/35CA663DDD30E31A1E7840AC0607FA7B" pageId="11" pageNumber="10" tableUuid="35CA663DDD30E31A1E7840AC0607FA7B">Table 2</tableCitation>
). The comparison of the differences among the sequences of the mitochondrial 16S rRNA or/ and COI genes within the genus 
<taxonomicName id="A6B54D36DD34E31E1FE14345050EF90B" authorityName="Thorell" authorityYear="1876" box="[489,607,1663,1686]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD34E31E1FE14345050EF90B" box="[489,607,1663,1686]" italics="true" pageId="11" pageNumber="10">Euscorpius</emphasis>
</taxonomicName>
represents one of the primary important tools for species delimitation in this morphologically uniform group (e.g. 
<bibRefCitation id="05244B44DD34E31E1C4A438407FDF96B" author="GANTENBEIN" pageId="11" pageNumber="10" pagination="49 - 65" refId="ref6987" refString="GANTENBEIN, B., V. FET, C. R. LARGIADER &amp; A. SCHOLL. 1999. First DNA phylogeny of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) and its bearing on taxonomy and biogeography of this genus. Biogeographica (Paris), 75 (2): 49 - 65." type="journal article" year="1999">Gantenbein et al., 1999</bibRefCitation>
; 
<bibRefCitation id="05244B44DD34E31E1EB343E406EFF96B" author="PARMAKELIS" box="[187,446,1758,1782]" pageId="11" pageNumber="10" pagination="728 - 748" refId="ref7648" refString="PARMAKELIS, A., P. KOTSAKIOZI, I. STATHI, S. POULIKARAKOU &amp; V. FET. 2013. Hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Greece revealed by multilocus species-delimitation approaches. Biological Journal of the Linnean Society, London, 110: 728 - 748." type="journal article" year="2013">Parmakelis et al., 2013</bibRefCitation>
; 
<bibRefCitation id="05244B44DD34E31E1FC543E40597F96B" author="STUNDLOVA" box="[461,710,1758,1784]" pageId="11" pageNumber="10" pagination="152 - 163" refId="ref7942" refString="STUNDLOVA, J., J. SMID, P. NGUYEN &amp; F. STAHLAVSKY. 2019. Cryptic diversity and dynamic chromosome evolution in Alpine scorpions (Euscorpiidae: Euscorpius). Molecular Phylogenetics and Evolution, 134: 152 - 163." type="journal article" year="2019">Štundlová et al., 2019</bibRefCitation>
). Our comparison of the genetic distances (p-distances and Kimura 2-parameter distances) (
<tableCitation id="2C37030EDD34E31E1F7342240680F8AB" box="[379,465,1822,1846]" captionStart="Table 2" captionStartId="15.[112,168,1430,1451]" captionTargetBox="[117,1471,936,1402]" captionTargetId="graphics-361@15.[111,1476,929,1407]" captionTargetPageId="15" captionText="Table 2. A matrix of genetic distances among the COI genes of the Euscorpius species from Bulgaria, eastern Serbia, and northern Greece. p-distance (below the diagonal); Kimura 2-parameter distance (above the diagonal). In the brackets are mentioned specimen codes/accession number from GenBank." httpUri="http://table.plazi.org/id/35CA663DDD30E31A1E7840AC0607FA7B" pageId="11" pageNumber="10" tableUuid="35CA663DDD30E31A1E7840AC0607FA7B">Table 2</tableCitation>
) among the species known from the studied region (
<collectingCountry id="19A27625DD34E31E1F7A42040681F8CB" box="[370,464,1854,1878]" name="Bulgaria" pageId="11" pageNumber="10">Bulgaria</collectingCountry>
, Eastern 
<collectingCountry id="19A27625DD34E31E1C2542040523F8CB" box="[557,626,1854,1878]" name="Serbia" pageId="11" pageNumber="10">Serbia</collectingCountry>
, and Northern 
<collectingCountry id="19A27625DD34E31E1E7C42640791F8EB" box="[116,192,1886,1910]" name="Greece" pageId="11" pageNumber="10">Greece</collectingCountry>
) corresponds to the previous analyses of the species from this area (
<bibRefCitation id="05244B44DD34E31E1F1B42440559F80B" author="PARMAKELIS" box="[275,520,1918,1942]" pageId="11" pageNumber="10" pagination="728 - 748" refId="ref7648" refString="PARMAKELIS, A., P. KOTSAKIOZI, I. STATHI, S. POULIKARAKOU &amp; V. FET. 2013. Hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Greece revealed by multilocus species-delimitation approaches. Biological Journal of the Linnean Society, London, 110: 728 - 748." type="journal article" year="2013">Parmakelis et al., 2013</bibRefCitation>
; Fet et al., 2014; 
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a, 
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b) and retrieved values of distances always exceeded 0.046 among all analyzed species. This fact supports a separate position and species status of 
<taxonomicName id="A6B54D36DD34E31E1C1F42E505C4F86B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[535,661,2014,2038]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1C1F42E505C4F86B" box="[535,661,2014,2038]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1C9342E40587F868" box="[667,726,2014,2038]" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1C9342E405E5F86B" bold="true" box="[667,692,2014,2038]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1CC942E40581F86B" bold="true" box="[705,720,2014,2038]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
</paragraph>
<paragraph id="610A36B5DD34E31E1D3B41C4034BF90B" blockId="11.[819,1477,1278,2038]" pageId="11" pageNumber="10">
AFFINITIES. DNA analysis separates 
<taxonomicName id="A6B54D36DD34E31E1AA641C5027CFA8B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1198,1325,1278,1302]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1AA641C5027CFA8B" box="[1198,1325,1278,1302]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1B3A41C4023CFA88" box="[1330,1389,1278,1302]" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1B3A41C4021AFA8B" bold="true" box="[1330,1355,1278,1302]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1B5F41C40237FA8B" bold="true" box="[1367,1382,1278,1302]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. The morphologically most similar species is 
<taxonomicName id="A6B54D36DD34E31E1DAD400503BDFACB" authority="Tropea et al., 2015" authorityName="Tropea" authorityYear="2015" box="[933,1260,1342,1366]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD34E31E1DAD4005035EFACB" box="[933,1039,1343,1366]" italics="true" pageId="11" pageNumber="10">E. popovi</emphasis>
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</taxonomicName>
, from southwestern 
<collectingCountry id="19A27625DD34E31E1D3B406404DEFAEB" box="[819,911,1374,1398]" name="Bulgaria" pageId="11" pageNumber="10">Bulgaria</collectingCountry>
and northeastern 
<collectingCountry id="19A27625DD34E31E1A6C406403E2FAEB" box="[1124,1203,1374,1398]" name="Greece" pageId="11" pageNumber="10">Greece</collectingCountry>
. These two species are differentiated according to the granulation of the dorsal surface of the pedipalp patella, which is well developed in 
<taxonomicName id="A6B54D36DD34E31E1B5540A50294FA2B" box="[1373,1477,1439,1462]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD34E31E1B5540A50294FA2B" box="[1373,1477,1439,1462]" italics="true" pageId="11" pageNumber="10">E. popovi</emphasis>
</taxonomicName>
and reduced to absent in 
<taxonomicName id="A6B54D36DD34E31E1A434085039FFA4B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1099,1230,1470,1494]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1A434085039FFA4B" box="[1099,1230,1470,1494]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1AD040840246FA48" box="[1240,1303,1470,1494]" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1AD0408403A0FA4B" bold="true" box="[1240,1265,1470,1494]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1B0940840241FA4B" bold="true" box="[1281,1296,1470,1494]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="F98E2A30DD34E31E1B2040840226FA4B" box="[1320,1399,1470,1494]" captionStart="Figures 23–31" captionStartId="9.[115,192,1979,2001]" captionTargetBox="[129,1472,285,1928]" captionTargetId="figure-54@9.[115,1476,272,1960]" captionTargetPageId="9" captionText="Figures 23–31. E. thracicus sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31)." figureDoi="http://doi.org/10.5281/zenodo.4648934" httpUri="https://zenodo.org/record/4648934/files/figure.png" pageId="11" pageNumber="10">Fig. 26</figureCitation>
versus fig. 
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a: 5). The male of 
<taxonomicName id="A6B54D36DD34E31E1B1440E502CCFA6B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1308,1437,1502,1526]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1B1440E502CCFA6B" box="[1308,1437,1502,1526]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1BAD40E40418F988" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1BAD40E402EFFA6B" bold="true" box="[1445,1470,1502,1526]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1D3B40C40413F98B" bold="true" box="[819,834,1534,1558]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
has pedipalp finger margin profile 
<typeStatus id="BE0E8817DD34E31E1ACE40C403A5F98B" box="[1222,1268,1534,1558]" pageId="11" pageNumber="10">type</typeStatus>
C versus 
<typeStatus id="BE0E8817DD34E31E1B6A40C402C1F98B" box="[1378,1424,1534,1558]" pageId="11" pageNumber="10">type</typeStatus>
A in 
<taxonomicName id="A6B54D36DD34E31E1D3B432504C8F9AB" box="[819,921,1567,1590]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD34E31E1D3B432504C8F9AB" box="[819,921,1567,1590]" italics="true" pageId="11" pageNumber="10">E. popovi</emphasis>
</taxonomicName>
. Females have fingers very weakly undulate, almost linear in 
<taxonomicName id="A6B54D36DD34E31E1D9D43050349F9CB" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[917,1048,1598,1622]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1D9D43050349F9CB" box="[917,1048,1598,1622]" italics="true" pageId="11" pageNumber="10">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1A284304030FF9C8" box="[1056,1118,1598,1622]" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1A2843040368F9CB" bold="true" box="[1056,1081,1598,1622]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1A4043040306F9CB" bold="true" box="[1096,1111,1598,1622]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
and undulate in 
<taxonomicName id="A6B54D36DD34E31E1B1F4305022EF9CB" box="[1303,1407,1599,1622]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD34E31E1B1F4305022EF9CB" box="[1303,1407,1599,1622]" italics="true" pageId="11" pageNumber="10">E. popovi</emphasis>
</taxonomicName>
(
<figureCitation id="F98E2A30DD34E31E1B8743040401F9EB" captionStart="Figures 23–31" captionStartId="9.[115,192,1979,2001]" captionTargetBox="[129,1472,285,1928]" captionTargetId="figure-54@9.[115,1476,272,1960]" captionTargetPageId="9" captionText="Figures 23–31. E. thracicus sp. n., male holotype, pedipalp segments. Chela dorsal (23), external (24) and ventral (25) views. Patella dorsal (26), external (27) and ventral (28) views. Trochanter and femur dorsal (29) and ventral (30) views. Movable finger dentition (31)." figureDoi="http://doi.org/10.5281/zenodo.4648934" httpUri="https://zenodo.org/record/4648934/files/figure.png" pageId="11" pageNumber="10">Figs. 24</figureCitation>
, 
<figureCitation id="F98E2A30DD34E31E1D5243640424F9EB" box="[858,885,1630,1654]" captionStart="Figures 32–39" captionStartId="10.[113,190,1958,1980]" captionTargetBox="[125,1441,250,1915]" captionTargetId="figure-66@10.[113,1474,244,1951]" captionTargetPageId="10" captionText="Figures 32–39. E. thracicus sp. n., female paratype, pedipalp segments. Chela dorsal (32), external (33) and ventral (34) views. Patella dorsal (35), external (36) and ventral (37) views. Trochanter and femur dorsal (38) and ventral (39) views. Trichobothrial pattern is indicated by white circles (32–38)." figureDoi="http://doi.org/10.5281/zenodo.4648936" httpUri="https://zenodo.org/record/4648936/files/figure.png" pageId="11" pageNumber="10">33</figureCitation>
vs. 
<figureCitation id="F98E2A30DD34E31E1DA84364035EF9EB" box="[928,1039,1630,1654]" captionStart="Figures 7–16" captionStartId="7.[115,192,1958,1980]" captionTargetBox="[113,1474,248,1887]" captionTargetId="figure-74@7.[113,1474,248,1911]" captionTargetPageId="7" captionText="Figures 7–16: E. thracicus sp. n. Figures 7, 9, 11–15. Male holotype, carapace and tergites I–IV (7), coxosternal area (9), left legs I–IV, retrolateral aspect (11–14 respectively), and telson lateral (15). Figures 8, 10, 16. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–IV (10), and telson lateral (16)." figureDoi="http://doi.org/10.5281/zenodo.4648930" httpUri="https://zenodo.org/record/4648930/files/figure.png" pageId="11" pageNumber="10">Figs. 7, 11</figureCitation>
in 
<bibRefCitation id="05244B44DD34E31E1A27436403A8F9EB" author="TROPEA" box="[1071,1273,1630,1654]" pageId="11" pageNumber="10" pagination="1 - 16" refId="ref7987" refString="TROPEA, G. &amp; V. FET. 2015. Two new Euscorpius species from central-western Greece (Scorpiones: Euscorpiidae). Euscorpius, 199: 1 - 16." type="journal article" year="2015">Tropea et al., 2015</bibRefCitation>
a: 5; see KovařÍk &amp; Šťáhlavský, 2020: 2).
</paragraph>
<paragraph id="610A36B5DD34E31A1D6B43A40634F92D" blockId="11.[819,1477,1278,2038]" lastBlockId="15.[112,1474,1430,2033]" lastPageId="15" lastPageNumber="14" pageId="11" pageNumber="10">
The hemispermatophore distal carina of 
<taxonomicName id="A6B54D36DD34E31E1B1243A502CDF92B" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[1306,1436,1694,1718]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD34E31E1B1243A5027BF92B" box="[1306,1322,1695,1718]" italics="true" pageId="11" pageNumber="10">E</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1B3143A402CDF92B" box="[1337,1436,1694,1718]" italics="true" pageId="11" pageNumber="10">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD34E31E1BAD43A40418F948" pageId="11" pageNumber="10" rank="species">
<emphasis id="53C1EAA7DD34E31E1BAD43A402EFF92B" bold="true" box="[1445,1470,1694,1718]" pageId="11" pageNumber="10">sp</emphasis>
. 
<emphasis id="53C1EAA7DD34E31E1D3B43840413F94B" bold="true" box="[819,834,1726,1750]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
differed from those of other examined species of related euscorpiids (cf. 
<figureCitation id="F98E2A30DD34E31E1DD643E4037BF96B" box="[990,1066,1758,1782]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="11" pageNumber="10">Fig. 58</figureCitation>
vs. 
<figureCitation id="F98E2A30DD34E31E1A5E43E40384F96B" box="[1110,1237,1758,1782]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="11" pageNumber="10">Figs. 50–57</figureCitation>
) in: (i) lacking a basal lobe (
<emphasis id="53C1EAA7DD34E31E1D7A43C404D6F88B" box="[882,903,1790,1814]" italics="true" pageId="11" pageNumber="10">bl</emphasis>
), (ii) having the distal external lobe (
<emphasis id="53C1EAA7DD34E31E1B2A43C40212F88B" box="[1314,1347,1790,1814]" italics="true" pageId="11" pageNumber="10">lde</emphasis>
) apparently divided into two separate sub-lobes, and (iii) lacking the distal internal lobe (
<emphasis id="53C1EAA7DD34E31E1A1A4204037FF8CB" box="[1042,1070,1854,1878]" italics="true" pageId="11" pageNumber="10">ldi</emphasis>
). An alternative interpretation of the anatomy is that the broader distal sub-lobe corresponds to the 
<emphasis id="53C1EAA7DD34E31E1D3B42440405F80B" box="[819,852,1918,1942]" italics="true" pageId="11" pageNumber="10">lde</emphasis>
, and the narrower distal sub-lobe to the 
<emphasis id="53C1EAA7DD34E31E1B1C42440261F80B" box="[1300,1328,1918,1942]" italics="true" pageId="11" pageNumber="10">ldi</emphasis>
. However, in other species the 
<emphasis id="53C1EAA7DD34E31E1DE542A40358F82B" box="[1005,1033,1950,1974]" italics="true" pageId="11" pageNumber="10">ldi</emphasis>
is always directly joined to the 
<emphasis id="53C1EAA7DD34E31E1B6A42A402D2F82B" box="[1378,1411,1950,1974]" italics="true" pageId="11" pageNumber="10">lde</emphasis>
by an oblique non-terminal suture (which may be partial or limited to a short notch), and its axis is distinctly slanted relative to the 
<emphasis id="53C1EAA7DD30E31A1EB040C20788F98D" box="[184,217,1528,1552]" italics="true" pageId="15" pageNumber="14">lde</emphasis>
axis. In contrast, the two sub-lobes observed here were strongly separated from each other with their axes nearly parallel. We also reject the hypothesis that the narrower distal sub-lobe is homologous to the basal lobe (
<emphasis id="53C1EAA7DD30E31A1C5943620537F9ED" box="[593,614,1624,1648]" italics="true" pageId="15" pageNumber="14">bl</emphasis>
), since the bl invariably arises from a more basal position on the distal carina in other species.
</paragraph>
<caption id="35CA663DDD33E3191E7B42FD032FF78A" ID-DOI="http://doi.org/10.5281/zenodo.4648940" ID-Zenodo-Dep="4648940" httpUri="https://zenodo.org/record/4648940/files/figure.png" pageId="12" pageNumber="11" startId="12.[115,192,1991,2013]" targetBox="[151,1429,222,1983]" targetPageId="12">
<paragraph id="610A36B5DD33E3191E7B42FD032FF78A" blockId="12.[115,1473,1991,2071]" pageId="12" pageNumber="11">
<emphasis id="53C1EAA7DD33E3191E7B42FD0650F841" bold="true" box="[115,257,1991,2013]" pageId="12" pageNumber="11">Figures 44–49</emphasis>
. 
<taxonomicName id="A6B54D36DD33E3191F1B42F2068CF841" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[275,477,1991,2013]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD33E3191F1B42F2068CF841" box="[275,477,1991,2013]" italics="true" pageId="12" pageNumber="11">Euscorpius thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD33E3191FEB42FD0549F841" box="[483,536,1991,2012]" pageId="12" pageNumber="11" rank="species">
<emphasis id="53C1EAA7DD33E3191FEB42FD06ABF841" bold="true" box="[483,506,1991,2012]" pageId="12" pageNumber="11">sp</emphasis>
. 
<emphasis id="53C1EAA7DD33E3191C0E42FD0542F841" bold="true" box="[518,531,1991,2012]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, left hemispermatophore. 
<emphasis id="53C1EAA7DD33E3191D1D42FD04F2F841" bold="true" box="[789,931,1991,2013]" pageId="12" pageNumber="11">Figures 44–45</emphasis>
. Whole hemispermatophore in convex (44) and concave (45) views. 
<emphasis id="53C1EAA7DD33E3191EEC42D90622F864" bold="true" box="[228,371,2019,2041]" pageId="12" pageNumber="11">Figures 46–49</emphasis>
. Capsule of left hemispermatophore, 3-dimensional form displayed in cross stereoscopic pairs, in posterior-convex (46), anterior (47), convex (48) and concave (49) views. Paratype No. 1864. Scale bar: 1 mm (Figs. 44–49).
</paragraph>
</caption>
<caption id="35CA663DDD32E3181E7B42840524F7DA" ID-DOI="http://doi.org/10.5281/zenodo.4648942" ID-Zenodo-Dep="4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="13" pageNumber="12" startId="13.[115,192,1982,2004]" targetBox="[115,1472,214,1975]" targetPageId="13">
<paragraph id="610A36B5DD32E3181E7B42840524F7DA" blockId="13.[115,1474,1982,2120]" pageId="13" pageNumber="12">
<emphasis id="53C1EAA7DD32E3181E7B428407AEF84E" bold="true" box="[115,255,1982,2004]" pageId="13" pageNumber="12">Figures 50–58</emphasis>
. Comparison of the capsule lobes of the hemispermatophores of nine euscorpiids (anterior-convex views). 
<emphasis id="53C1EAA7DD32E3181AEC428403ADF84E" bold="true" box="[1252,1276,1982,2003]" pageId="13" pageNumber="12">50</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181B0E4284078AF86D" baseAuthorityName="De Geer" baseAuthorityYear="1778" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD32E3181B0E4284078AF86D" italics="true" pageId="13" pageNumber="12">Tetratrichobothrius flavicaudis</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181EE242E10653F86D" bold="true" box="[234,258,2011,2032]" pageId="13" pageNumber="12">51</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181F1942E6062CF86C" authorityName="Thorell" authorityYear="1876" box="[273,381,2012,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD32E3181F1942E6062CF86C" box="[273,381,2012,2033]" italics="true" pageId="13" pageNumber="12">Euscorpius</emphasis>
</taxonomicName>
sp., nr 
<taxonomicName id="A6B54D36DD32E3181FC542E60564F86C" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[461,565,2012,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="53C1EAA7DD32E3181FC542E60564F86C" box="[461,565,2012,2033]" italics="true" pageId="13" pageNumber="12">tergestinus</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181C4D42E1050CF86D" bold="true" box="[581,605,2011,2032]" pageId="13" pageNumber="12">52</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181C6442E605BAF86C" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[620,747,2012,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="53C1EAA7DD32E3181C6442E6052AF86C" box="[620,635,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181C8242E605BAF86C" box="[650,747,2012,2033]" italics="true" pageId="13" pageNumber="12">concinnus</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181CF242E10443F86D" bold="true" box="[762,786,2011,2032]" pageId="13" pageNumber="12">53</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181D2942E604C6F86D" box="[801,919,2011,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD32E3181D2942E60461F86C" box="[801,816,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181D3742E104C6F86D" box="[831,919,2011,2032]" italics="true" pageId="13" pageNumber="12">solegladi</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181DAE42E104EFF86D" bold="true" box="[934,958,2011,2032]" pageId="13" pageNumber="12">54</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181DC542E60363F86D" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[973,1074,2011,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="53C1EAA7DD32E3181DC542E6048DF86C" box="[973,988,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181DE342E10363F86D" box="[1003,1074,2011,2032]" italics="true" pageId="13" pageNumber="12">italicus</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181A4842E10309F86D" bold="true" box="[1088,1112,2011,2032]" pageId="13" pageNumber="12">55</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181A6F42E6038BF86D" box="[1127,1242,2011,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD32E3181A6F42E60327F86C" box="[1127,1142,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181A8D42E1038BF86D" box="[1157,1242,2011,2032]" italics="true" pageId="13" pageNumber="12">deltshevi</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181AE142E10250F86D" bold="true" box="[1257,1281,2011,2032]" pageId="13" pageNumber="12">56</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181B1842E60227F86C" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[1296,1398,2012,2033]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="53C1EAA7DD32E3181B1842E6024EF86C" box="[1296,1311,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181B2642E60227F86C" box="[1326,1398,2012,2033]" italics="true" pageId="13" pageNumber="12">sicanus</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181B8D42E102CCF86D" bold="true" box="[1413,1437,2011,2032]" pageId="13" pageNumber="12">57</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181BA442E60796F790" authorityName="Birula" authorityYear="1903" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="candiota">
<emphasis id="53C1EAA7DD32E3181BA442E602EAF86C" box="[1452,1467,2012,2033]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181E7B42C20796F790" box="[115,199,2040,2061]" italics="true" pageId="13" pageNumber="12">candiota</emphasis>
</taxonomicName>
. 
<emphasis id="53C1EAA7DD32E3181EDA42C207BBF790" bold="true" box="[210,234,2040,2061]" pageId="13" pageNumber="12">58</emphasis>
. 
<taxonomicName id="A6B54D36DD32E3181EFE42C30638F790" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[246,361,2040,2062]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD32E3181EFE42C30654F793" box="[246,261,2041,2062]" italics="true" pageId="13" pageNumber="12">E</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181F1842C20638F790" box="[272,361,2040,2061]" italics="true" pageId="13" pageNumber="12">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD32E3181F6642C206F2F790" box="[366,419,2040,2061]" pageId="13" pageNumber="12" rank="species">
<emphasis id="53C1EAA7DD32E3181F6642C206D4F790" bold="true" box="[366,389,2040,2061]" pageId="13" pageNumber="12">sp</emphasis>
. 
<emphasis id="53C1EAA7DD32E3181F9842C206CCF790" bold="true" box="[400,413,2040,2061]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
, paratype No. 1864 Abbreviations: 
<emphasis id="53C1EAA7DD32E3181CF642C2044AF790" box="[766,795,2040,2061]" italics="true" pageId="13" pageNumber="12">lde</emphasis>
: distal external lobe; 
<emphasis id="53C1EAA7DD32E3181DE042C20350F790" box="[1000,1025,2040,2061]" italics="true" pageId="13" pageNumber="12">ldi</emphasis>
: distal internal lobe; 
<emphasis id="53C1EAA7DD32E3181AC342C2038FF790" box="[1227,1246,2040,2061]" italics="true" pageId="13" pageNumber="12">bl</emphasis>
: basal lobe; 
<emphasis id="53C1EAA7DD32E3181B5E42C20220F790" box="[1366,1393,2040,2061]" italics="true" pageId="13" pageNumber="12">cls</emphasis>
: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58.
</paragraph>
</caption>
<caption id="35CA663DDD31E31B1E7C42E50380F868" ID-DOI="http://doi.org/10.5281/zenodo.4648944" ID-Zenodo-Dep="4648944" box="[116,1233,2015,2037]" httpUri="https://zenodo.org/record/4648944/files/figure.png" pageId="14" pageNumber="13" startId="14.[116,193,2015,2037]" targetBox="[113,1474,268,2003]" targetPageId="14">
<paragraph id="610A36B5DD31E31B1E7C42E50380F868" blockId="14.[116,1233,2015,2037]" box="[116,1233,2015,2037]" pageId="14" pageNumber="13">
<emphasis id="53C1EAA7DD31E31B1E7C42E50652F869" bold="true" box="[116,259,2015,2037]" pageId="14" pageNumber="13">Figures 59–60</emphasis>
. 
<taxonomicName id="A6B54D36DD31E31B1F0742E506D2F869" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[271,387,2015,2036]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD31E31B1F0742E506D2F869" box="[271,387,2015,2036]" italics="true" pageId="14" pageNumber="13">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD31E31B1F8142E506EFF869" box="[393,446,2015,2036]" pageId="14" pageNumber="13" rank="species">
<emphasis id="53C1EAA7DD31E31B1F8142E506F1F869" bold="true" box="[393,416,2015,2036]" pageId="14" pageNumber="13">sp</emphasis>
. 
<emphasis id="53C1EAA7DD31E31B1FA442E506E8F869" bold="true" box="[428,441,2015,2036]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, type locality, Bulgaria, Krumovgrad, Bryagovets, 41.6485862°N 25.8132185°E.
</paragraph>
</caption>
<caption id="35CA663DDD30E31A1E7C464C0487FC10" ID-DOI="http://doi.org/10.5281/zenodo.4648946" ID-Zenodo-Dep="4648946" box="[116,982,886,909]" httpUri="https://zenodo.org/record/4648946/files/figure.png" pageId="15" pageNumber="14" startId="15.[116,184,886,908]" targetBox="[113,1474,234,877]" targetPageId="15">
<paragraph id="610A36B5DD30E31A1E7C464C0487FC10" blockId="15.[116,982,886,909]" box="[116,982,886,909]" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7C464C0784FC16" bold="true" box="[116,213,886,908]" pageId="15" pageNumber="14">Figure 61</emphasis>
. Map of distribution of 
<taxonomicName id="A6B54D36DD30E31A1FB4464D0579FC11" authorityName="Thorell" authorityYear="1876" box="[444,552,887,908]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD30E31A1FB4464D0579FC11" box="[444,552,887,908]" italics="true" pageId="15" pageNumber="14">Euscorpius</emphasis>
</taxonomicName>
populations analysed by DNA (see Table 2).
</paragraph>
</caption>
<paragraph id="610A36B5DD30E31A1C30469202F8FAE7" pageId="15" pageNumber="14">
<table id="13B5C415DD301CEA1E7D469202EEFAE7" box="[117,1471,936,1402]" gridcols="17" gridrows="16" pageId="15" pageNumber="14">
<tr id="DF8534F7DD301CEA1E7D469202EEFC21" box="[117,1471,936,956]" gridrow="0" pageId="15" pageNumber="14" rowspan-0="1" rowspan-1="1">
<th id="9C545D8BDD301CEA1C2B46920509FC21" box="[547,600,936,956]" gridcol="2" gridrow="0" pageId="15" pageNumber="14">1</th>
<th id="9C545D8BDD301CEA1C69469205C7FC21" box="[609,662,936,956]" gridcol="3" gridrow="0" pageId="15" pageNumber="14">2</th>
<th id="9C545D8BDD301CEA1C9746920585FC21" box="[671,724,936,956]" gridcol="4" gridrow="0" pageId="15" pageNumber="14">3</th>
<th id="9C545D8BDD301CEA1CD546920443FC21" box="[733,786,936,956]" gridcol="5" gridrow="0" pageId="15" pageNumber="14">4</th>
<th id="9C545D8BDD301CEA1D1546920403FC21" box="[797,850,936,956]" gridcol="6" gridrow="0" pageId="15" pageNumber="14">5</th>
<th id="9C545D8BDD301CEA1D53469204C1FC21" box="[859,912,936,956]" gridcol="7" gridrow="0" pageId="15" pageNumber="14">6</th>
<th id="9C545D8BDD301CEA1D924692049EFC21" box="[922,975,936,956]" gridcol="8" gridrow="0" pageId="15" pageNumber="14">7</th>
<th id="9C545D8BDD301CEA1DD04692035CFC21" box="[984,1037,936,956]" gridcol="9" gridrow="0" pageId="15" pageNumber="14">8</th>
<th id="9C545D8BDD301CEA1A1E4692031AFC21" box="[1046,1099,936,956]" gridcol="10" gridrow="0" pageId="15" pageNumber="14">9</th>
<th id="9C545D8BDD301CEA1A5C469203D8FC21" box="[1108,1161,936,956]" gridcol="11" gridrow="0" pageId="15" pageNumber="14">10</th>
<th id="9C545D8BDD301CEA1A9A46920396FC21" box="[1170,1223,936,956]" gridcol="12" gridrow="0" pageId="15" pageNumber="14">11</th>
<th id="9C545D8BDD301CEA1AD846920254FC21" box="[1232,1285,936,956]" gridcol="13" gridrow="0" pageId="15" pageNumber="14">12</th>
<th id="9C545D8BDD301CEA1B0646920212FC21" box="[1294,1347,936,956]" gridcol="14" gridrow="0" pageId="15" pageNumber="14">13</th>
<th id="9C545D8BDD301CEA1B44469202D0FC21" box="[1356,1409,936,956]" gridcol="15" gridrow="0" pageId="15" pageNumber="14">14</th>
<th id="9C545D8BDD301CEA1B82469202EEFC21" box="[1418,1471,936,956]" gridcol="16" gridrow="0" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1B91469202E0FC21" bold="true" box="[1433,1457,936,956]" pageId="15" pageNumber="14">15</emphasis>
</th>
</tr>
<tr id="DF8534F7DD301CEA1E7D46FC02EEFC47" box="[117,1471,966,986]" gridrow="1" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D46FC07DCFC47" box="[117,141,966,986]" gridcol="0" gridrow="1" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E8846FC07DDFC47" bold="true" box="[128,140,966,986]" pageId="15" pageNumber="14">1</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D46FC055DFC47" box="[149,524,966,986]" gridcol="1" gridrow="1" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D46FC0655FC47" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[149,260,966,986]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD30E31A1E9D46FC0655FC47" box="[149,260,966,986]" italics="true" pageId="15" pageNumber="14">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD30E31A1F0646FC0617FC47" box="[270,326,966,986]" pageId="15" pageNumber="14" rank="species">
<emphasis id="53C1EAA7DD30E31A1F0646FC0676FC47" bold="true" box="[270,295,966,986]" pageId="15" pageNumber="14">sp</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1F3B46FC0611FC47" bold="true" box="[307,320,966,986]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
(S1865/ 
<accessionNumber id="7EE6AB56DD30E31A1F9D46FC0556FC47" box="[405,519,966,986]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW291129" pageId="15" pageNumber="14">MW291129</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B46FC0509FC47" box="[547,600,966,986]" gridcol="2" gridrow="1" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1C6946FC05C7FC47" box="[609,662,966,986]" gridcol="3" gridrow="1" pageId="15" pageNumber="14">0.002</td>
<td id="9C545D8BDD301CEA1C9746FC0585FC47" box="[671,724,966,986]" gridcol="4" gridrow="1" pageId="15" pageNumber="14">0.094</td>
<td id="9C545D8BDD301CEA1CD546FC0443FC47" box="[733,786,966,986]" gridcol="5" gridrow="1" pageId="15" pageNumber="14">0.102</td>
<td id="9C545D8BDD301CEA1D1546FC0403FC47" box="[797,850,966,986]" gridcol="6" gridrow="1" pageId="15" pageNumber="14">0.099</td>
<td id="9C545D8BDD301CEA1D5346FC04C1FC47" box="[859,912,966,986]" gridcol="7" gridrow="1" pageId="15" pageNumber="14">0.099</td>
<td id="9C545D8BDD301CEA1D9246FC049EFC47" box="[922,975,966,986]" gridcol="8" gridrow="1" pageId="15" pageNumber="14">0.064</td>
<td id="9C545D8BDD301CEA1DD046FC035CFC47" box="[984,1037,966,986]" gridcol="9" gridrow="1" pageId="15" pageNumber="14">0.066</td>
<td id="9C545D8BDD301CEA1A1E46FC031AFC47" box="[1046,1099,966,986]" gridcol="10" gridrow="1" pageId="15" pageNumber="14">0.078</td>
<td id="9C545D8BDD301CEA1A5C46FC03D8FC47" box="[1108,1161,966,986]" gridcol="11" gridrow="1" pageId="15" pageNumber="14">0.060</td>
<td id="9C545D8BDD301CEA1A9A46FC0396FC47" box="[1170,1223,966,986]" gridcol="12" gridrow="1" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1AD846FC0254FC47" box="[1232,1285,966,986]" gridcol="13" gridrow="1" pageId="15" pageNumber="14">0.049</td>
<td id="9C545D8BDD301CEA1B0646FC0212FC47" box="[1294,1347,966,986]" gridcol="14" gridrow="1" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1B4446FC02D0FC47" box="[1356,1409,966,986]" gridcol="15" gridrow="1" pageId="15" pageNumber="14">0.048</td>
<td id="9C545D8BDD301CEA1B8246FC02EEFC47" box="[1418,1471,966,986]" gridcol="16" gridrow="1" pageId="15" pageNumber="14">0.067</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D46DE02EEFC65" box="[117,1471,996,1016]" gridrow="2" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D46DE07DCFC65" box="[117,141,996,1016]" gridcol="0" gridrow="2" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E8846DE07DDFC65" bold="true" box="[128,140,996,1016]" pageId="15" pageNumber="14">2</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D46DE055DFC65" box="[149,524,996,1016]" gridcol="1" gridrow="2" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D46DE0655FC65" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[149,260,996,1016]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD30E31A1E9D46DE0655FC65" box="[149,260,996,1016]" italics="true" pageId="15" pageNumber="14">E. thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD30E31A1F0646DE0616FC65" box="[270,327,996,1016]" pageId="15" pageNumber="14" rank="species">
<emphasis id="53C1EAA7DD30E31A1F0646DE0676FC65" bold="true" box="[270,295,996,1016]" pageId="15" pageNumber="14">sp</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1F3B46DE0611FC65" bold="true" box="[307,320,996,1016]" pageId="15" pageNumber="14">n</emphasis>
.
</taxonomicNameLabel>
(S1866/ 
<accessionNumber id="7EE6AB56DD30E31A1F9D46DE0556FC65" box="[405,519,996,1016]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW291114" pageId="15" pageNumber="14">MW291114</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B46DE0509FC65" box="[547,600,996,1016]" gridcol="2" gridrow="2" pageId="15" pageNumber="14">0.002</td>
<td id="9C545D8BDD301CEA1C6946DE05C7FC65" box="[609,662,996,1016]" gridcol="3" gridrow="2" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1C9746DE0585FC65" box="[671,724,996,1016]" gridcol="4" gridrow="2" pageId="15" pageNumber="14">0.096</td>
<td id="9C545D8BDD301CEA1CD546DE0443FC65" box="[733,786,996,1016]" gridcol="5" gridrow="2" pageId="15" pageNumber="14">0.104</td>
<td id="9C545D8BDD301CEA1D1546DE0403FC65" box="[797,850,996,1016]" gridcol="6" gridrow="2" pageId="15" pageNumber="14">0.101</td>
<td id="9C545D8BDD301CEA1D5346DE04C1FC65" box="[859,912,996,1016]" gridcol="7" gridrow="2" pageId="15" pageNumber="14">0.101</td>
<td id="9C545D8BDD301CEA1D9246DE049EFC65" box="[922,975,996,1016]" gridcol="8" gridrow="2" pageId="15" pageNumber="14">0.067</td>
<td id="9C545D8BDD301CEA1DD046DE035CFC65" box="[984,1037,996,1016]" gridcol="9" gridrow="2" pageId="15" pageNumber="14">0.068</td>
<td id="9C545D8BDD301CEA1A1E46DE031AFC65" box="[1046,1099,996,1016]" gridcol="10" gridrow="2" pageId="15" pageNumber="14">0.080</td>
<td id="9C545D8BDD301CEA1A5C46DE03D8FC65" box="[1108,1161,996,1016]" gridcol="11" gridrow="2" pageId="15" pageNumber="14">0.062</td>
<td id="9C545D8BDD301CEA1A9A46DE0396FC65" box="[1170,1223,996,1016]" gridcol="12" gridrow="2" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1AD846DE0254FC65" box="[1232,1285,996,1016]" gridcol="13" gridrow="2" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1B0646DE0212FC65" box="[1294,1347,996,1016]" gridcol="14" gridrow="2" pageId="15" pageNumber="14">0.054</td>
<td id="9C545D8BDD301CEA1B4446DE02D0FC65" box="[1356,1409,996,1016]" gridcol="15" gridrow="2" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1B8246DE02EEFC65" box="[1418,1471,996,1016]" gridcol="16" gridrow="2" pageId="15" pageNumber="14">0.069</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D413B02EEFB8B" box="[117,1471,1025,1046]" gridrow="3" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D413B07DCFB8B" box="[117,141,1025,1046]" gridcol="0" gridrow="3" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88413807DDFB8B" bold="true" box="[128,140,1026,1046]" pageId="15" pageNumber="14">3</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D413B055DFB8B" box="[149,524,1025,1046]" gridcol="1" gridrow="3" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D413B0652FB88" box="[149,259,1025,1045]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD30E31A1E9D413B0652FB88" box="[149,259,1025,1045]" italics="true" pageId="15" pageNumber="14">E. deltshevi</emphasis>
</taxonomicName>
(VF-0746-1/ 
<accessionNumber id="7EE6AB56DD30E31A1F73413B06B6FB88" box="[379,487,1025,1045]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM111244" pageId="15" pageNumber="14">KM111244</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B413B0509FB8B" box="[547,600,1025,1046]" gridcol="2" gridrow="3" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1C69413B05C7FB8B" box="[609,662,1025,1046]" gridcol="3" gridrow="3" pageId="15" pageNumber="14">0.089</td>
<td id="9C545D8BDD301CEA1C97413B0585FB8B" box="[671,724,1025,1046]" gridcol="4" gridrow="3" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1CD5413B0443FB8B" box="[733,786,1025,1046]" gridcol="5" gridrow="3" pageId="15" pageNumber="14">0.047</td>
<td id="9C545D8BDD301CEA1D15413B0403FB8B" box="[797,850,1025,1046]" gridcol="6" gridrow="3" pageId="15" pageNumber="14">0.079</td>
<td id="9C545D8BDD301CEA1D53413B04C1FB8B" box="[859,912,1025,1046]" gridcol="7" gridrow="3" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1D92413B049EFB8B" box="[922,975,1025,1046]" gridcol="8" gridrow="3" pageId="15" pageNumber="14">0.086</td>
<td id="9C545D8BDD301CEA1DD0413B035CFB8B" box="[984,1037,1025,1046]" gridcol="9" gridrow="3" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1A1E413B031AFB8B" box="[1046,1099,1025,1046]" gridcol="10" gridrow="3" pageId="15" pageNumber="14">0.101</td>
<td id="9C545D8BDD301CEA1A5C413B03D8FB8B" box="[1108,1161,1025,1046]" gridcol="11" gridrow="3" pageId="15" pageNumber="14">0.089</td>
<td id="9C545D8BDD301CEA1A9A413B0396FB8B" box="[1170,1223,1025,1046]" gridcol="12" gridrow="3" pageId="15" pageNumber="14">0.075</td>
<td id="9C545D8BDD301CEA1AD8413B0254FB8B" box="[1232,1285,1025,1046]" gridcol="13" gridrow="3" pageId="15" pageNumber="14">0.079</td>
<td id="9C545D8BDD301CEA1B06413B0212FB8B" box="[1294,1347,1025,1046]" gridcol="14" gridrow="3" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1B44413B02D0FB8B" box="[1356,1409,1025,1046]" gridcol="15" gridrow="3" pageId="15" pageNumber="14">0.076</td>
<td id="9C545D8BDD301CEA1B82413B02EEFB8B" box="[1418,1471,1025,1046]" gridcol="16" gridrow="3" pageId="15" pageNumber="14">0.084</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D412502EEFBAE" box="[117,1471,1055,1075]" gridrow="4" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D412507DCFBAE" box="[117,141,1055,1075]" gridcol="0" gridrow="4" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88412507DDFBAE" bold="true" box="[128,140,1055,1075]" pageId="15" pageNumber="14">4</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4125055DFBAE" box="[149,524,1055,1075]" gridcol="1" gridrow="4" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D41250652FBAE" box="[149,259,1055,1075]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD30E31A1E9D41250652FBAE" box="[149,259,1055,1075]" italics="true" pageId="15" pageNumber="14">E. deltshevi</emphasis>
</taxonomicName>
(VF-0821-1/ 
<accessionNumber id="7EE6AB56DD30E31A1F71412506B6FBAE" box="[377,487,1055,1075]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM111243" pageId="15" pageNumber="14">KM111243</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B41250509FBAE" box="[547,600,1055,1075]" gridcol="2" gridrow="4" pageId="15" pageNumber="14">0.094</td>
<td id="9C545D8BDD301CEA1C69412505C7FBAE" box="[609,662,1055,1075]" gridcol="3" gridrow="4" pageId="15" pageNumber="14">0.096</td>
<td id="9C545D8BDD301CEA1C9741250585FBAE" box="[671,724,1055,1075]" gridcol="4" gridrow="4" pageId="15" pageNumber="14">0.045</td>
<td id="9C545D8BDD301CEA1CD541250443FBAE" box="[733,786,1055,1075]" gridcol="5" gridrow="4" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1D1541250403FBAE" box="[797,850,1055,1075]" gridcol="6" gridrow="4" pageId="15" pageNumber="14">0.089</td>
<td id="9C545D8BDD301CEA1D53412504C1FBAE" box="[859,912,1055,1075]" gridcol="7" gridrow="4" pageId="15" pageNumber="14">0.089</td>
<td id="9C545D8BDD301CEA1D924125049EFBAE" box="[922,975,1055,1075]" gridcol="8" gridrow="4" pageId="15" pageNumber="14">0.097</td>
<td id="9C545D8BDD301CEA1DD04125035CFBAE" box="[984,1037,1055,1075]" gridcol="9" gridrow="4" pageId="15" pageNumber="14">0.093</td>
<td id="9C545D8BDD301CEA1A1E4125031AFBAE" box="[1046,1099,1055,1075]" gridcol="10" gridrow="4" pageId="15" pageNumber="14">0.106</td>
<td id="9C545D8BDD301CEA1A5C412503D8FBAE" box="[1108,1161,1055,1075]" gridcol="11" gridrow="4" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1A9A41250396FBAE" box="[1170,1223,1055,1075]" gridcol="12" gridrow="4" pageId="15" pageNumber="14">0.092</td>
<td id="9C545D8BDD301CEA1AD841250254FBAE" box="[1232,1285,1055,1075]" gridcol="13" gridrow="4" pageId="15" pageNumber="14">0.097</td>
<td id="9C545D8BDD301CEA1B0641250212FBAE" box="[1294,1347,1055,1075]" gridcol="14" gridrow="4" pageId="15" pageNumber="14">0.099</td>
<td id="9C545D8BDD301CEA1B44412502D0FBAE" box="[1356,1409,1055,1075]" gridcol="15" gridrow="4" pageId="15" pageNumber="14">0.093</td>
<td id="9C545D8BDD301CEA1B82412502EEFBAE" box="[1418,1471,1055,1075]" gridcol="16" gridrow="4" pageId="15" pageNumber="14">0.086</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D410702EEFBCC" box="[117,1471,1085,1105]" gridrow="5" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D410707DCFBCC" box="[117,141,1085,1105]" gridcol="0" gridrow="5" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88410707DDFBCC" bold="true" box="[128,140,1085,1105]" pageId="15" pageNumber="14">5</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4107055DFBCC" box="[149,524,1085,1105]" gridcol="1" gridrow="5" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D41070652FBCC" box="[149,259,1085,1105]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD30E31A1E9D41070652FBCC" box="[149,259,1085,1105]" italics="true" pageId="15" pageNumber="14">E. solegladi</emphasis>
</taxonomicName>
(VF-0801-1/ 
<accessionNumber id="7EE6AB56DD30E31A1F71410706B7FBCC" box="[377,486,1085,1105]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM111247" pageId="15" pageNumber="14">KM111247</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B41070509FBCC" box="[547,600,1085,1105]" gridcol="2" gridrow="5" pageId="15" pageNumber="14">0.093</td>
<td id="9C545D8BDD301CEA1C69410705C7FBCC" box="[609,662,1085,1105]" gridcol="3" gridrow="5" pageId="15" pageNumber="14">0.094</td>
<td id="9C545D8BDD301CEA1C9741070585FBCC" box="[671,724,1085,1105]" gridcol="4" gridrow="5" pageId="15" pageNumber="14">0.074</td>
<td id="9C545D8BDD301CEA1CD541070443FBCC" box="[733,786,1085,1105]" gridcol="5" gridrow="5" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1D1541070403FBCC" box="[797,850,1085,1105]" gridcol="6" gridrow="5" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1D53410704C1FBCC" box="[859,912,1085,1105]" gridcol="7" gridrow="5" pageId="15" pageNumber="14">0.003</td>
<td id="9C545D8BDD301CEA1D924107049EFBCC" box="[922,975,1085,1105]" gridcol="8" gridrow="5" pageId="15" pageNumber="14">0.088</td>
<td id="9C545D8BDD301CEA1DD04107035CFBCC" box="[984,1037,1085,1105]" gridcol="9" gridrow="5" pageId="15" pageNumber="14">0.089</td>
<td id="9C545D8BDD301CEA1A1E4107031AFBCC" box="[1046,1099,1085,1105]" gridcol="10" gridrow="5" pageId="15" pageNumber="14">0.112</td>
<td id="9C545D8BDD301CEA1A5C410703D8FBCC" box="[1108,1161,1085,1105]" gridcol="11" gridrow="5" pageId="15" pageNumber="14">0.085</td>
<td id="9C545D8BDD301CEA1A9A41070396FBCC" box="[1170,1223,1085,1105]" gridcol="12" gridrow="5" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1AD841070254FBCC" box="[1232,1285,1085,1105]" gridcol="13" gridrow="5" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1B0641070212FBCC" box="[1294,1347,1085,1105]" gridcol="14" gridrow="5" pageId="15" pageNumber="14">0.085</td>
<td id="9C545D8BDD301CEA1B44410702D0FBCC" box="[1356,1409,1085,1105]" gridcol="15" gridrow="5" pageId="15" pageNumber="14">0.082</td>
<td id="9C545D8BDD301CEA1B82410702EEFBCC" box="[1418,1471,1085,1105]" gridcol="16" gridrow="5" pageId="15" pageNumber="14">0.092</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D416002EEFBF2" box="[117,1471,1114,1135]" gridrow="6" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D416007DCFBF2" box="[117,141,1114,1135]" gridcol="0" gridrow="6" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88416107DDFBF2" bold="true" box="[128,140,1115,1135]" pageId="15" pageNumber="14">6</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4160055DFBF2" box="[149,524,1114,1135]" gridcol="1" gridrow="6" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D41600652FBF3" box="[149,259,1114,1134]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD30E31A1E9D41600652FBF3" box="[149,259,1114,1134]" italics="true" pageId="15" pageNumber="14">E. solegladi</emphasis>
</taxonomicName>
(VF-0802-1/ 
<accessionNumber id="7EE6AB56DD30E31A1F73416006B6FBF3" box="[379,487,1114,1134]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KM111246" pageId="15" pageNumber="14">KM111246</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B41600509FBF2" box="[547,600,1114,1135]" gridcol="2" gridrow="6" pageId="15" pageNumber="14">0.093</td>
<td id="9C545D8BDD301CEA1C69416005C7FBF2" box="[609,662,1114,1135]" gridcol="3" gridrow="6" pageId="15" pageNumber="14">0.094</td>
<td id="9C545D8BDD301CEA1C9741600585FBF2" box="[671,724,1114,1135]" gridcol="4" gridrow="6" pageId="15" pageNumber="14">0.076</td>
<td id="9C545D8BDD301CEA1CD541600443FBF2" box="[733,786,1114,1135]" gridcol="5" gridrow="6" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1D1541600403FBF2" box="[797,850,1114,1135]" gridcol="6" gridrow="6" pageId="15" pageNumber="14">0.004</td>
<td id="9C545D8BDD301CEA1D53416004C1FBF2" box="[859,912,1114,1135]" gridcol="7" gridrow="6" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1D924160049EFBF2" box="[922,975,1114,1135]" gridcol="8" gridrow="6" pageId="15" pageNumber="14">0.090</td>
<td id="9C545D8BDD301CEA1DD04160035CFBF2" box="[984,1037,1114,1135]" gridcol="9" gridrow="6" pageId="15" pageNumber="14">0.091</td>
<td id="9C545D8BDD301CEA1A1E4160031AFBF2" box="[1046,1099,1114,1135]" gridcol="10" gridrow="6" pageId="15" pageNumber="14">0.115</td>
<td id="9C545D8BDD301CEA1A5C416003D8FBF2" box="[1108,1161,1114,1135]" gridcol="11" gridrow="6" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1A9A41600396FBF2" box="[1170,1223,1114,1135]" gridcol="12" gridrow="6" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1AD841600254FBF2" box="[1232,1285,1114,1135]" gridcol="13" gridrow="6" pageId="15" pageNumber="14">0.085</td>
<td id="9C545D8BDD301CEA1B0641600212FBF2" box="[1294,1347,1114,1135]" gridcol="14" gridrow="6" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1B44416002D0FBF2" box="[1356,1409,1114,1135]" gridcol="15" gridrow="6" pageId="15" pageNumber="14">0.084</td>
<td id="9C545D8BDD301CEA1B82416002EEFBF2" box="[1418,1471,1114,1135]" gridcol="16" gridrow="6" pageId="15" pageNumber="14">0.090</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D414202EEFB10" box="[117,1471,1144,1165]" gridrow="7" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D414207DCFB10" box="[117,141,1144,1165]" gridcol="0" gridrow="7" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88414307DDFB10" bold="true" box="[128,140,1145,1165]" pageId="15" pageNumber="14">7</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4142055DFB10" box="[149,524,1144,1165]" gridcol="1" gridrow="7" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D414207BFFB11" box="[149,238,1144,1164]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD30E31A1E9D414207BFFB11" box="[149,238,1144,1164]" italics="true" pageId="15" pageNumber="14">E. popovi</emphasis>
</taxonomicName>
(FESP13/ 
<accessionNumber id="7EE6AB56DD30E31A1F5A414206EBFB11" box="[338,442,1144,1164]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC215733" pageId="15" pageNumber="14">KC215733</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B41420509FB10" box="[547,600,1144,1165]" gridcol="2" gridrow="7" pageId="15" pageNumber="14">0.061</td>
<td id="9C545D8BDD301CEA1C69414205C7FB10" box="[609,662,1144,1165]" gridcol="3" gridrow="7" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1C9741420585FB10" box="[671,724,1144,1165]" gridcol="4" gridrow="7" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1CD541420443FB10" box="[733,786,1144,1165]" gridcol="5" gridrow="7" pageId="15" pageNumber="14">0.090</td>
<td id="9C545D8BDD301CEA1D1541420403FB10" box="[797,850,1144,1165]" gridcol="6" gridrow="7" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1D53414204C1FB10" box="[859,912,1144,1165]" gridcol="7" gridrow="7" pageId="15" pageNumber="14">0.084</td>
<td id="9C545D8BDD301CEA1D924142049EFB10" box="[922,975,1144,1165]" gridcol="8" gridrow="7" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1DD04142035CFB10" box="[984,1037,1144,1165]" gridcol="9" gridrow="7" pageId="15" pageNumber="14">0.012</td>
<td id="9C545D8BDD301CEA1A1E4142031AFB10" box="[1046,1099,1144,1165]" gridcol="10" gridrow="7" pageId="15" pageNumber="14">0.091</td>
<td id="9C545D8BDD301CEA1A5C414203D8FB10" box="[1108,1161,1144,1165]" gridcol="11" gridrow="7" pageId="15" pageNumber="14">0.064</td>
<td id="9C545D8BDD301CEA1A9A41420396FB10" box="[1170,1223,1144,1165]" gridcol="12" gridrow="7" pageId="15" pageNumber="14">0.059</td>
<td id="9C545D8BDD301CEA1AD841420254FB10" box="[1232,1285,1144,1165]" gridcol="13" gridrow="7" pageId="15" pageNumber="14">0.056</td>
<td id="9C545D8BDD301CEA1B0641420212FB10" box="[1294,1347,1144,1165]" gridcol="14" gridrow="7" pageId="15" pageNumber="14">0.058</td>
<td id="9C545D8BDD301CEA1B44414202D0FB10" box="[1356,1409,1144,1165]" gridcol="15" gridrow="7" pageId="15" pageNumber="14">0.055</td>
<td id="9C545D8BDD301CEA1B82414202EEFB10" box="[1418,1471,1144,1165]" gridcol="16" gridrow="7" pageId="15" pageNumber="14">0.082</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D41AC02EEFB37" box="[117,1471,1174,1194]" gridrow="8" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D41AC07DCFB37" box="[117,141,1174,1194]" gridcol="0" gridrow="8" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E8841AC07DDFB37" bold="true" box="[128,140,1174,1194]" pageId="15" pageNumber="14">8</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D41AC055DFB37" box="[149,524,1174,1194]" gridcol="1" gridrow="8" pageId="15" pageNumber="14">
<taxonomicName id="A6B54D36DD30E31A1E9D41AC07BFFB37" box="[149,238,1174,1194]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD30E31A1E9D41AC07BFFB37" box="[149,238,1174,1194]" italics="true" pageId="15" pageNumber="14">E. popovi</emphasis>
</taxonomicName>
(FESP21/ 
<accessionNumber id="7EE6AB56DD30E31A1F5841AC06EBFB37" box="[336,442,1174,1194]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC215737" pageId="15" pageNumber="14">KC215737</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B41AC0509FB37" box="[547,600,1174,1194]" gridcol="2" gridrow="8" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1C6941AC05C7FB37" box="[609,662,1174,1194]" gridcol="3" gridrow="8" pageId="15" pageNumber="14">0.065</td>
<td id="9C545D8BDD301CEA1C9741AC0585FB37" box="[671,724,1174,1194]" gridcol="4" gridrow="8" pageId="15" pageNumber="14">0.082</td>
<td id="9C545D8BDD301CEA1CD541AC0443FB37" box="[733,786,1174,1194]" gridcol="5" gridrow="8" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1D1541AC0403FB37" box="[797,850,1174,1194]" gridcol="6" gridrow="8" pageId="15" pageNumber="14">0.083</td>
<td id="9C545D8BDD301CEA1D5341AC04C1FB37" box="[859,912,1174,1194]" gridcol="7" gridrow="8" pageId="15" pageNumber="14">0.085</td>
<td id="9C545D8BDD301CEA1D9241AC049EFB37" box="[922,975,1174,1194]" gridcol="8" gridrow="8" pageId="15" pageNumber="14">0.012</td>
<td id="9C545D8BDD301CEA1DD041AC035CFB37" box="[984,1037,1174,1194]" gridcol="9" gridrow="8" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1A1E41AC031AFB37" box="[1046,1099,1174,1194]" gridcol="10" gridrow="8" pageId="15" pageNumber="14">0.086</td>
<td id="9C545D8BDD301CEA1A5C41AC03D8FB37" box="[1108,1161,1174,1194]" gridcol="11" gridrow="8" pageId="15" pageNumber="14">0.064</td>
<td id="9C545D8BDD301CEA1A9A41AC0396FB37" box="[1170,1223,1174,1194]" gridcol="12" gridrow="8" pageId="15" pageNumber="14">0.056</td>
<td id="9C545D8BDD301CEA1AD841AC0254FB37" box="[1232,1285,1174,1194]" gridcol="13" gridrow="8" pageId="15" pageNumber="14">0.056</td>
<td id="9C545D8BDD301CEA1B0641AC0212FB37" box="[1294,1347,1174,1194]" gridcol="14" gridrow="8" pageId="15" pageNumber="14">0.058</td>
<td id="9C545D8BDD301CEA1B4441AC02D0FB37" box="[1356,1409,1174,1194]" gridcol="15" gridrow="8" pageId="15" pageNumber="14">0.054</td>
<td id="9C545D8BDD301CEA1B8241AC02EEFB37" box="[1418,1471,1174,1194]" gridcol="16" gridrow="8" pageId="15" pageNumber="14">0.078</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D418E02EEFB55" box="[117,1471,1204,1224]" gridrow="9" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D418E07DCFB55" box="[117,141,1204,1224]" gridcol="0" gridrow="9" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E88418E07DDFB55" bold="true" box="[128,140,1204,1224]" pageId="15" pageNumber="14">9</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D418E055DFB55" box="[149,524,1204,1224]" gridcol="1" gridrow="9" pageId="15" pageNumber="14">
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<emphasis id="53C1EAA7DD30E31A1E9D418E07A9FB55" box="[149,248,1204,1224]" italics="true" pageId="15" pageNumber="14">E. drenskii</emphasis>
</taxonomicName>
(117F/ 
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)
</td>
<td id="9C545D8BDD301CEA1C2B418E0509FB55" box="[547,600,1204,1224]" gridcol="2" gridrow="9" pageId="15" pageNumber="14">0.073</td>
<td id="9C545D8BDD301CEA1C69418E05C7FB55" box="[609,662,1204,1224]" gridcol="3" gridrow="9" pageId="15" pageNumber="14">0.075</td>
<td id="9C545D8BDD301CEA1C97418E0585FB55" box="[671,724,1204,1224]" gridcol="4" gridrow="9" pageId="15" pageNumber="14">0.094</td>
<td id="9C545D8BDD301CEA1CD5418E0443FB55" box="[733,786,1204,1224]" gridcol="5" gridrow="9" pageId="15" pageNumber="14">0.098</td>
<td id="9C545D8BDD301CEA1D15418E0403FB55" box="[797,850,1204,1224]" gridcol="6" gridrow="9" pageId="15" pageNumber="14">0.104</td>
<td id="9C545D8BDD301CEA1D53418E04C1FB55" box="[859,912,1204,1224]" gridcol="7" gridrow="9" pageId="15" pageNumber="14">0.106</td>
<td id="9C545D8BDD301CEA1D92418E049EFB55" box="[922,975,1204,1224]" gridcol="8" gridrow="9" pageId="15" pageNumber="14">0.084</td>
<td id="9C545D8BDD301CEA1DD0418E035CFB55" box="[984,1037,1204,1224]" gridcol="9" gridrow="9" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1A1E418E031AFB55" box="[1046,1099,1204,1224]" gridcol="10" gridrow="9" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1A5C418E03D8FB55" box="[1108,1161,1204,1224]" gridcol="11" gridrow="9" pageId="15" pageNumber="14">0.067</td>
<td id="9C545D8BDD301CEA1A9A418E0396FB55" box="[1170,1223,1204,1224]" gridcol="12" gridrow="9" pageId="15" pageNumber="14">0.062</td>
<td id="9C545D8BDD301CEA1AD8418E0254FB55" box="[1232,1285,1204,1224]" gridcol="13" gridrow="9" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1B06418E0212FB55" box="[1294,1347,1204,1224]" gridcol="14" gridrow="9" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1B44418E02D0FB55" box="[1356,1409,1204,1224]" gridcol="15" gridrow="9" pageId="15" pageNumber="14">0.064</td>
<td id="9C545D8BDD301CEA1B82418E02EEFB55" box="[1418,1471,1204,1224]" gridcol="16" gridrow="9" pageId="15" pageNumber="14">0.072</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D41EB02EEFB7B" box="[117,1471,1233,1254]" gridrow="10" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D41EB07DCFB7B" box="[117,141,1233,1254]" gridcol="0" gridrow="10" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D41E807DCFB7B" bold="true" box="[117,141,1234,1254]" pageId="15" pageNumber="14">10</emphasis>
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<td id="9C545D8BDD301CEA1E9D41EB055DFB7B" box="[149,524,1233,1254]" gridcol="1" gridrow="10" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D41EB07FBFB78" box="[149,170,1233,1253]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (113F/ 
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)
</td>
<td id="9C545D8BDD301CEA1C2B41EB0509FB7B" box="[547,600,1233,1254]" gridcol="2" gridrow="10" pageId="15" pageNumber="14">0.057</td>
<td id="9C545D8BDD301CEA1C6941EB05C7FB7B" box="[609,662,1233,1254]" gridcol="3" gridrow="10" pageId="15" pageNumber="14">0.059</td>
<td id="9C545D8BDD301CEA1C9741EB0585FB7B" box="[671,724,1233,1254]" gridcol="4" gridrow="10" pageId="15" pageNumber="14">0.084</td>
<td id="9C545D8BDD301CEA1CD541EB0443FB7B" box="[733,786,1233,1254]" gridcol="5" gridrow="10" pageId="15" pageNumber="14">0.078</td>
<td id="9C545D8BDD301CEA1D1541EB0403FB7B" box="[797,850,1233,1254]" gridcol="6" gridrow="10" pageId="15" pageNumber="14">0.080</td>
<td id="9C545D8BDD301CEA1D5341EB04C1FB7B" box="[859,912,1233,1254]" gridcol="7" gridrow="10" pageId="15" pageNumber="14">0.082</td>
<td id="9C545D8BDD301CEA1D9241EB049EFB7B" box="[922,975,1233,1254]" gridcol="8" gridrow="10" pageId="15" pageNumber="14">0.061</td>
<td id="9C545D8BDD301CEA1DD041EB035CFB7B" box="[984,1037,1233,1254]" gridcol="9" gridrow="10" pageId="15" pageNumber="14">0.061</td>
<td id="9C545D8BDD301CEA1A1E41EB031AFB7B" box="[1046,1099,1233,1254]" gridcol="10" gridrow="10" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1A5C41EB03D8FB7B" box="[1108,1161,1233,1254]" gridcol="11" gridrow="10" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1A9A41EB0396FB7B" box="[1170,1223,1233,1254]" gridcol="12" gridrow="10" pageId="15" pageNumber="14">0.040</td>
<td id="9C545D8BDD301CEA1AD841EB0254FB7B" box="[1232,1285,1233,1254]" gridcol="13" gridrow="10" pageId="15" pageNumber="14">0.047</td>
<td id="9C545D8BDD301CEA1B0641EB0212FB7B" box="[1294,1347,1233,1254]" gridcol="14" gridrow="10" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1B4441EB02D0FB7B" box="[1356,1409,1233,1254]" gridcol="15" gridrow="10" pageId="15" pageNumber="14">0.046</td>
<td id="9C545D8BDD301CEA1B8241EB02EEFB7B" box="[1418,1471,1233,1254]" gridcol="16" gridrow="10" pageId="15" pageNumber="14">0.058</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D41D502EEFA9E" box="[117,1471,1263,1283]" gridrow="11" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D41D507DCFA9E" box="[117,141,1263,1283]" gridcol="0" gridrow="11" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D41D507DCFA9E" bold="true" box="[117,141,1263,1283]" pageId="15" pageNumber="14">11</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D41D5055DFA9E" box="[149,524,1263,1283]" gridcol="1" gridrow="11" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D41D507FBFA9E" box="[149,170,1263,1283]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (FESP12/ 
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)
</td>
<td id="9C545D8BDD301CEA1C2B41D50509FA9E" box="[547,600,1263,1283]" gridcol="2" gridrow="11" pageId="15" pageNumber="14">0.048</td>
<td id="9C545D8BDD301CEA1C6941D505C7FA9E" box="[609,662,1263,1283]" gridcol="3" gridrow="11" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1C9741D50585FA9E" box="[671,724,1263,1283]" gridcol="4" gridrow="11" pageId="15" pageNumber="14">0.071</td>
<td id="9C545D8BDD301CEA1CD541D50443FA9E" box="[733,786,1263,1283]" gridcol="5" gridrow="11" pageId="15" pageNumber="14">0.086</td>
<td id="9C545D8BDD301CEA1D1541D50403FA9E" box="[797,850,1263,1283]" gridcol="6" gridrow="11" pageId="15" pageNumber="14">0.076</td>
<td id="9C545D8BDD301CEA1D5341D504C1FA9E" box="[859,912,1263,1283]" gridcol="7" gridrow="11" pageId="15" pageNumber="14">0.078</td>
<td id="9C545D8BDD301CEA1D9241D5049EFA9E" box="[922,975,1263,1283]" gridcol="8" gridrow="11" pageId="15" pageNumber="14">0.057</td>
<td id="9C545D8BDD301CEA1DD041D5035CFA9E" box="[984,1037,1263,1283]" gridcol="9" gridrow="11" pageId="15" pageNumber="14">0.054</td>
<td id="9C545D8BDD301CEA1A1E41D5031AFA9E" box="[1046,1099,1263,1283]" gridcol="10" gridrow="11" pageId="15" pageNumber="14">0.058</td>
<td id="9C545D8BDD301CEA1A5C41D503D8FA9E" box="[1108,1161,1263,1283]" gridcol="11" gridrow="11" pageId="15" pageNumber="14">0.039</td>
<td id="9C545D8BDD301CEA1A9A41D50396FA9E" box="[1170,1223,1263,1283]" gridcol="12" gridrow="11" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1AD841D50254FA9E" box="[1232,1285,1263,1283]" gridcol="13" gridrow="11" pageId="15" pageNumber="14">0.010</td>
<td id="9C545D8BDD301CEA1B0641D50212FA9E" box="[1294,1347,1263,1283]" gridcol="14" gridrow="11" pageId="15" pageNumber="14">0.010</td>
<td id="9C545D8BDD301CEA1B4441D502D0FA9E" box="[1356,1409,1263,1283]" gridcol="15" gridrow="11" pageId="15" pageNumber="14">0.008</td>
<td id="9C545D8BDD301CEA1B8241D502EEFA9E" box="[1418,1471,1263,1283]" gridcol="16" gridrow="11" pageId="15" pageNumber="14">0.041</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D403702EEFABC" box="[117,1471,1293,1313]" gridrow="12" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D403707DCFABC" box="[117,141,1293,1313]" gridcol="0" gridrow="12" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D403707DCFABC" bold="true" box="[117,141,1293,1313]" pageId="15" pageNumber="14">12</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4037055DFABC" box="[149,524,1293,1313]" gridcol="1" gridrow="12" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D403707FBFABC" box="[149,170,1293,1313]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (FESP11/ 
<accessionNumber id="7EE6AB56DD30E31A1F27403706C8FABC" box="[303,409,1293,1313]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC215731" pageId="15" pageNumber="14">KC215731</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B40370509FABC" box="[547,600,1293,1313]" gridcol="2" gridrow="12" pageId="15" pageNumber="14">0.048</td>
<td id="9C545D8BDD301CEA1C69403705C7FABC" box="[609,662,1293,1313]" gridcol="3" gridrow="12" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1C9740370585FABC" box="[671,724,1293,1313]" gridcol="4" gridrow="12" pageId="15" pageNumber="14">0.074</td>
<td id="9C545D8BDD301CEA1CD540370443FABC" box="[733,786,1293,1313]" gridcol="5" gridrow="12" pageId="15" pageNumber="14">0.090</td>
<td id="9C545D8BDD301CEA1D1540370403FABC" box="[797,850,1293,1313]" gridcol="6" gridrow="12" pageId="15" pageNumber="14">0.078</td>
<td id="9C545D8BDD301CEA1D53403704C1FABC" box="[859,912,1293,1313]" gridcol="7" gridrow="12" pageId="15" pageNumber="14">0.080</td>
<td id="9C545D8BDD301CEA1D924037049EFABC" box="[922,975,1293,1313]" gridcol="8" gridrow="12" pageId="15" pageNumber="14">0.054</td>
<td id="9C545D8BDD301CEA1DD04037035CFABC" box="[984,1037,1293,1313]" gridcol="9" gridrow="12" pageId="15" pageNumber="14">0.053</td>
<td id="9C545D8BDD301CEA1A1E4037031AFABC" box="[1046,1099,1293,1313]" gridcol="10" gridrow="12" pageId="15" pageNumber="14">0.060</td>
<td id="9C545D8BDD301CEA1A5C403703D8FABC" box="[1108,1161,1293,1313]" gridcol="11" gridrow="12" pageId="15" pageNumber="14">0.046</td>
<td id="9C545D8BDD301CEA1A9A40370396FABC" box="[1170,1223,1293,1313]" gridcol="12" gridrow="12" pageId="15" pageNumber="14">0.010</td>
<td id="9C545D8BDD301CEA1AD840370254FABC" box="[1232,1285,1293,1313]" gridcol="13" gridrow="12" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1B0640370212FABC" box="[1294,1347,1293,1313]" gridcol="14" gridrow="12" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B44403702D0FABC" box="[1356,1409,1293,1313]" gridcol="15" gridrow="12" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B82403702EEFABC" box="[1418,1471,1293,1313]" gridcol="16" gridrow="12" pageId="15" pageNumber="14">0.048</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D401002EEFAA2" box="[117,1471,1322,1343]" gridrow="13" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D401007DCFAA2" box="[117,141,1322,1343]" gridcol="0" gridrow="13" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D401107DCFAA2" bold="true" box="[117,141,1323,1343]" pageId="15" pageNumber="14">13</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4010055DFAA2" box="[149,524,1322,1343]" gridcol="1" gridrow="13" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D401107FBFAA2" box="[149,170,1323,1343]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (FESP09/ 
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)
</td>
<td id="9C545D8BDD301CEA1C2B40100509FAA2" box="[547,600,1322,1343]" gridcol="2" gridrow="13" pageId="15" pageNumber="14">0.050</td>
<td id="9C545D8BDD301CEA1C69401005C7FAA2" box="[609,662,1322,1343]" gridcol="3" gridrow="13" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1C9740100585FAA2" box="[671,724,1322,1343]" gridcol="4" gridrow="13" pageId="15" pageNumber="14">0.076</td>
<td id="9C545D8BDD301CEA1CD540100443FAA2" box="[733,786,1322,1343]" gridcol="5" gridrow="13" pageId="15" pageNumber="14">0.092</td>
<td id="9C545D8BDD301CEA1D1540100403FAA2" box="[797,850,1322,1343]" gridcol="6" gridrow="13" pageId="15" pageNumber="14">0.080</td>
<td id="9C545D8BDD301CEA1D53401004C1FAA2" box="[859,912,1322,1343]" gridcol="7" gridrow="13" pageId="15" pageNumber="14">0.082</td>
<td id="9C545D8BDD301CEA1D924010049EFAA2" box="[922,975,1322,1343]" gridcol="8" gridrow="13" pageId="15" pageNumber="14">0.056</td>
<td id="9C545D8BDD301CEA1DD04010035CFAA2" box="[984,1037,1322,1343]" gridcol="9" gridrow="13" pageId="15" pageNumber="14">0.055</td>
<td id="9C545D8BDD301CEA1A1E4010031AFAA2" box="[1046,1099,1322,1343]" gridcol="10" gridrow="13" pageId="15" pageNumber="14">0.060</td>
<td id="9C545D8BDD301CEA1A5C401003D8FAA2" box="[1108,1161,1322,1343]" gridcol="11" gridrow="13" pageId="15" pageNumber="14">0.048</td>
<td id="9C545D8BDD301CEA1A9A40100396FAA2" box="[1170,1223,1322,1343]" gridcol="12" gridrow="13" pageId="15" pageNumber="14">0.010</td>
<td id="9C545D8BDD301CEA1AD840100254FAA2" box="[1232,1285,1322,1343]" gridcol="13" gridrow="13" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B0640100212FAA2" box="[1294,1347,1322,1343]" gridcol="14" gridrow="13" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1B44401002D0FAA2" box="[1356,1409,1322,1343]" gridcol="15" gridrow="13" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B82401002EEFAA2" box="[1418,1471,1322,1343]" gridcol="16" gridrow="13" pageId="15" pageNumber="14">0.048</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D407202EEFAC0" box="[117,1471,1352,1373]" gridrow="14" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D407207DCFAC0" box="[117,141,1352,1373]" gridcol="0" gridrow="14" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D407307DCFAC0" bold="true" box="[117,141,1353,1373]" pageId="15" pageNumber="14">14</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D4072055DFAC0" box="[149,524,1352,1373]" gridcol="1" gridrow="14" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D407207FBFAC1" box="[149,170,1352,1372]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (FESP10/ 
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)
</td>
<td id="9C545D8BDD301CEA1C2B40720509FAC0" box="[547,600,1352,1373]" gridcol="2" gridrow="14" pageId="15" pageNumber="14">0.046</td>
<td id="9C545D8BDD301CEA1C69407205C7FAC0" box="[609,662,1352,1373]" gridcol="3" gridrow="14" pageId="15" pageNumber="14">0.048</td>
<td id="9C545D8BDD301CEA1C9740720585FAC0" box="[671,724,1352,1373]" gridcol="4" gridrow="14" pageId="15" pageNumber="14">0.071</td>
<td id="9C545D8BDD301CEA1CD540720443FAC0" box="[733,786,1352,1373]" gridcol="5" gridrow="14" pageId="15" pageNumber="14">0.087</td>
<td id="9C545D8BDD301CEA1D1540720403FAC0" box="[797,850,1352,1373]" gridcol="6" gridrow="14" pageId="15" pageNumber="14">0.077</td>
<td id="9C545D8BDD301CEA1D53407204C1FAC0" box="[859,912,1352,1373]" gridcol="7" gridrow="14" pageId="15" pageNumber="14">0.079</td>
<td id="9C545D8BDD301CEA1D924072049EFAC0" box="[922,975,1352,1373]" gridcol="8" gridrow="14" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1DD04072035CFAC0" box="[984,1037,1352,1373]" gridcol="9" gridrow="14" pageId="15" pageNumber="14">0.052</td>
<td id="9C545D8BDD301CEA1A1E4072031AFAC0" box="[1046,1099,1352,1373]" gridcol="10" gridrow="14" pageId="15" pageNumber="14">0.060</td>
<td id="9C545D8BDD301CEA1A5C407203D8FAC0" box="[1108,1161,1352,1373]" gridcol="11" gridrow="14" pageId="15" pageNumber="14">0.044</td>
<td id="9C545D8BDD301CEA1A9A40720396FAC0" box="[1170,1223,1352,1373]" gridcol="12" gridrow="14" pageId="15" pageNumber="14">0.008</td>
<td id="9C545D8BDD301CEA1AD840720254FAC0" box="[1232,1285,1352,1373]" gridcol="13" gridrow="14" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B0640720212FAC0" box="[1294,1347,1352,1373]" gridcol="14" gridrow="14" pageId="15" pageNumber="14">0.000</td>
<td id="9C545D8BDD301CEA1B44407202D0FAC0" box="[1356,1409,1352,1373]" gridcol="15" gridrow="14" pageId="15" pageNumber="14">-</td>
<td id="9C545D8BDD301CEA1B82407202EEFAC0" box="[1418,1471,1352,1373]" gridcol="16" gridrow="14" pageId="15" pageNumber="14">0.047</td>
</tr>
<tr id="DF8534F7DD301CEA1E7D405C02EEFAE7" box="[117,1471,1382,1402]" gridrow="15" pageId="15" pageNumber="14">
<th id="9C545D8BDD301CEA1E7D405C07DCFAE7" box="[117,141,1382,1402]" gridcol="0" gridrow="15" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7D405C07DCFAE7" bold="true" box="[117,141,1382,1402]" pageId="15" pageNumber="14">15</emphasis>
</th>
<td id="9C545D8BDD301CEA1E9D405C055DFAE7" box="[149,524,1382,1402]" gridcol="1" gridrow="15" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E9D405C07FBFAE7" box="[149,170,1382,1402]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. (120F/ 
<accessionNumber id="7EE6AB56DD30E31A1F18405C062BFAE7" box="[272,378,1382,1402]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC215663" pageId="15" pageNumber="14">KC215663</accessionNumber>
)
</td>
<td id="9C545D8BDD301CEA1C2B405C0509FAE7" box="[547,600,1382,1402]" gridcol="2" gridrow="15" pageId="15" pageNumber="14">0.063</td>
<td id="9C545D8BDD301CEA1C69405C05C7FAE7" box="[609,662,1382,1402]" gridcol="3" gridrow="15" pageId="15" pageNumber="14">0.065</td>
<td id="9C545D8BDD301CEA1C97405C0585FAE7" box="[671,724,1382,1402]" gridcol="4" gridrow="15" pageId="15" pageNumber="14">0.079</td>
<td id="9C545D8BDD301CEA1CD5405C0443FAE7" box="[733,786,1382,1402]" gridcol="5" gridrow="15" pageId="15" pageNumber="14">0.081</td>
<td id="9C545D8BDD301CEA1D15405C0403FAE7" box="[797,850,1382,1402]" gridcol="6" gridrow="15" pageId="15" pageNumber="14">0.086</td>
<td id="9C545D8BDD301CEA1D53405C04C1FAE7" box="[859,912,1382,1402]" gridcol="7" gridrow="15" pageId="15" pageNumber="14">0.084</td>
<td id="9C545D8BDD301CEA1D92405C049EFAE7" box="[922,975,1382,1402]" gridcol="8" gridrow="15" pageId="15" pageNumber="14">0.077</td>
<td id="9C545D8BDD301CEA1DD0405C035CFAE7" box="[984,1037,1382,1402]" gridcol="9" gridrow="15" pageId="15" pageNumber="14">0.073</td>
<td id="9C545D8BDD301CEA1A1E405C031AFAE7" box="[1046,1099,1382,1402]" gridcol="10" gridrow="15" pageId="15" pageNumber="14">0.067</td>
<td id="9C545D8BDD301CEA1A5C405C03D8FAE7" box="[1108,1161,1382,1402]" gridcol="11" gridrow="15" pageId="15" pageNumber="14">0.056</td>
<td id="9C545D8BDD301CEA1A9A405C0396FAE7" box="[1170,1223,1382,1402]" gridcol="12" gridrow="15" pageId="15" pageNumber="14">0.039</td>
<td id="9C545D8BDD301CEA1AD8405C0254FAE7" box="[1232,1285,1382,1402]" gridcol="13" gridrow="15" pageId="15" pageNumber="14">0.046</td>
<td id="9C545D8BDD301CEA1B06405C0212FAE7" box="[1294,1347,1382,1402]" gridcol="14" gridrow="15" pageId="15" pageNumber="14">0.046</td>
<td id="9C545D8BDD301CEA1B44405C02D0FAE7" box="[1356,1409,1382,1402]" gridcol="15" gridrow="15" pageId="15" pageNumber="14">0.045</td>
<td id="9C545D8BDD301CEA1B82405C02EEFAE7" box="[1418,1471,1382,1402]" gridcol="16" gridrow="15" pageId="15" pageNumber="14">-</td>
</tr>
</table>
</paragraph>
<caption id="35CA663DDD30E31A1E7840AC0607FA7B" ID-Table-UUID="35CA663DDD30E31A1E7840AC0607FA7B" httpUri="http://table.plazi.org/id/35CA663DDD30E31A1E7840AC0607FA7B" pageId="15" pageNumber="14" startId="15.[112,168,1430,1451]" targetBox="[117,1471,936,1402]" targetIsTable="true" targetPageId="15">
<paragraph id="610A36B5DD30E31A1E7840AC0607FA7B" blockId="15.[112,1474,1430,2033]" pageId="15" pageNumber="14">
<emphasis id="53C1EAA7DD30E31A1E7840AC07EDFA36" bold="true" box="[112,188,1430,1451]" pageId="15" pageNumber="14">Table 2</emphasis>
. A matrix of genetic distances among the COI genes of the 
<taxonomicName id="A6B54D36DD30E31A1D0F40AD0422FA31" authorityName="Thorell" authorityYear="1876" box="[775,883,1431,1452]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD30E31A1D0F40AD0422FA31" box="[775,883,1431,1452]" italics="true" pageId="15" pageNumber="14">Euscorpius</emphasis>
</taxonomicName>
species from Bulgaria, eastern Serbia, and northern Greece. p-distance (below the diagonal); Kimura 2-parameter distance (above the diagonal). In the brackets are mentioned specimen codes/accession number from GenBank.
</paragraph>
</caption>
<paragraph id="610A36B5DD30E3051EA8438206F2FE7E" blockId="15.[112,1474,1430,2033]" lastBlockId="16.[113,772,234,1635]" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
Of 33 other species of related euscorpiids with described hemispermatophores, the 
<emphasis id="53C1EAA7DD30E31A1F8E43E206CBF96D" box="[390,410,1752,1776]" italics="true" pageId="15" pageNumber="14">bl</emphasis>
was absent in only one species, 
<taxonomicName id="A6B54D36DD30E31A1E7843C205DDF88C" authority="(De Geer, 1778)" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[112,652,1784,1809]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD30E31A1E7843C206ECF88D" box="[112,445,1784,1808]" italics="true" pageId="15" pageNumber="14">Tetratrichobothrius flavicaudis</emphasis>
(De Geer, 1778)
</taxonomicName>
(
<figureCitation id="F98E2A30DD30E31A1CAD43C205ACF88D" box="[677,765,1784,1808]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="15" pageNumber="14">Fig. 50</figureCitation>
; Fet &amp; Soleglad, 2002; 
<bibRefCitation id="05244B44DD30E31A1F6242220568F8AD" author="JACOB" box="[362,569,1816,1840]" pageId="15" pageNumber="14" pagination="59 - 72" refId="ref7240" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 b. Complex male genitalia (hemispermatophores) are not diagnostic for cryptic species in the genus Euscorpius (Scorpiones: Euscorpiidae). Organisms Diversity &amp; Evolution, 4: 59 - 72." type="journal article" year="2004">Jacob et al., 2004b</bibRefCitation>
; Vachon, 1948). It has been reported to be present in 6 species of 
<emphasis id="53C1EAA7DD30E31A1C7642020453F8CD" box="[638,770,1848,1872]" italics="true" pageId="15" pageNumber="14">Alpiscorpius</emphasis>
<bibRefCitation id="05244B44DD30E31A1E784262063EF8EC" author="GANTENBEIN" box="[112,367,1880,1905]" pageId="15" pageNumber="14" pagination="49 - 65" refId="ref6987" refString="GANTENBEIN, B., V. FET, C. R. LARGIADER &amp; A. SCHOLL. 1999. First DNA phylogeny of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) and its bearing on taxonomy and biogeography of this genus. Biogeographica (Paris), 75 (2): 49 - 65." type="journal article" year="1999">Gantenbein et al., 1999</bibRefCitation>
(
<taxonomicName id="A6B54D36DD30E31A1F894263058CF8ED" authority="(Di Caporiacco, 1950)" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[385,733,1880,1904]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="alpha">
<emphasis id="53C1EAA7DD30E31A1F89426306C0F8ED" box="[385,401,1881,1904]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1FAA4262068FF8ED" box="[418,478,1880,1904]" italics="true" pageId="15" pageNumber="14">alpha</emphasis>
(Di Caporiacco, 1950)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1CE3426305AAF8ED" box="[747,763,1881,1904]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<taxonomicName id="A6B54D36DD30E31A1E784243069DF80D" authority="(Di Caporiacco, 1950)" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[112,460,1912,1936]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gamma">
<emphasis id="53C1EAA7DD30E31A1E78424307ECF80D" box="[112,189,1913,1936]" italics="true" pageId="15" pageNumber="14">gamma</emphasis>
(Di Caporiacco, 1950)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1FD7424306BEF80D" box="[479,495,1913,1936]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1C0D4243053DF80D" box="[517,620,1913,1936]" italics="true" pageId="15" pageNumber="14">germanus</emphasis>
(C.L. Koch, 1837); 
<taxonomicName id="A6B54D36DD30E31A1EB442A30553F82C" authority="(Kessler, 1874)" baseAuthorityName="Kessler" baseAuthorityYear="1874" box="[188,514,1944,1969]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mingrelicus">
<emphasis id="53C1EAA7DD30E31A1EB442A3079DF82D" box="[188,204,1945,1968]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1ED342A20604F82D" box="[219,341,1944,1968]" italics="true" pageId="15" pageNumber="14">mingrelicus</emphasis>
(Kessler, 1874)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1C0642A3054FF82D" box="[526,542,1945,1968]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1C2542A205D9F82D" box="[557,648,1944,1968]" italics="true" pageId="15" pageNumber="14">phrygius</emphasis>
(Bonacina, 1980); 
<taxonomicName id="A6B54D36DD30E31A1EC84283054AF84D" authority="(Lacroix, 1995))" authorityName="" baseAuthorityName="Lacroix" baseAuthorityYear="1995" box="[192,539,1976,2000]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="uludagensis">
<emphasis id="53C1EAA7DD30E31A1EC842830781F84D" box="[192,208,1977,2000]" italics="true" pageId="15" pageNumber="14">A</emphasis>
. 
<emphasis id="53C1EAA7DD30E31A1EEA4282060EF84D" box="[226,351,1976,2000]" italics="true" pageId="15" pageNumber="14">uludagensis</emphasis>
(Lacroix, 1995))
</taxonomicName>
(
<bibRefCitation id="05244B44DD30E31A1C26428205AFF84D" author="JACOB" box="[558,766,1976,2000]" pageId="15" pageNumber="14" pagination="59 - 72" refId="ref7240" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 b. Complex male genitalia (hemispermatophores) are not diagnostic for cryptic species in the genus Euscorpius (Scorpiones: Euscorpiidae). Organisms Diversity &amp; Evolution, 4: 59 - 72." type="journal article" year="2004">Jacob et al. 2004b</bibRefCitation>
; Molteni et al., 1983; 
<bibRefCitation id="05244B44DD30E31A1F5F42E2055CF86C" author="SCHERABON" box="[343,525,2008,2033]" pageId="15" pageNumber="14" pagination="77 - 154" refId="ref7825" refString="SCHERABON, B. 1987. Die Skorpione Osterreiches in vergleichender Sicht unter besonderer Berucksichtigung Karntens. Carinthia II, 45: 77 - 154." type="journal article" year="1987">Scherabon, 1987</bibRefCitation>
; 
<bibRefCitation id="05244B44DD30E31A1C1242E305BAF86D" author="TROPEA" box="[538,747,2008,2032]" pageId="15" pageNumber="14" pagination="1 - 16" refId="ref7987" refString="TROPEA, G. &amp; V. FET. 2015. Two new Euscorpius species from central-western Greece (Scorpiones: Euscorpiidae). Euscorpius, 199: 1 - 16." type="journal article" year="2015">Tropea et al., 2015</bibRefCitation>
c), and in 26 species of 
<taxonomicName id="A6B54D36DD30E31A1A1840C303D7F98D" authorityName="Thorell" authorityYear="1876" box="[1040,1158,1529,1552]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD30E31A1A1840C303D7F98D" box="[1040,1158,1529,1552]" italics="true" pageId="15" pageNumber="14">Euscorpius</emphasis>
</taxonomicName>
(
<taxonomicName id="A6B54D36DD30E31A1A9E40C302ECF98C" authority="Tropea et al., 2012" authorityName="Tropea" authorityYear="2012" box="[1174,1469,1528,1553]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="avcii">
<emphasis id="53C1EAA7DD30E31A1A9E40C303B9F98D" box="[1174,1256,1529,1552]" italics="true" pageId="15" pageNumber="14">E. avcii</emphasis>
Tropea et al., 2012
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1D27432303F3F9AD" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" box="[815,1186,1560,1584]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="balearicus">
<emphasis id="53C1EAA7DD30E31A1D27432304EDF9AD" box="[815,956,1560,1584]" italics="true" pageId="15" pageNumber="14">E. balearicus</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1AA5432302ECF9AC" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[1197,1469,1560,1585]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="birulai">
<emphasis id="53C1EAA7DD30E31A1AA543230242F9AD" box="[1197,1299,1560,1584]" italics="true" pageId="15" pageNumber="14">E. birulai</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1D2743030369F9CC" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" box="[815,1080,1592,1617]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="candiota">
<emphasis id="53C1EAA7DD30E31A1D27430304F8F9CD" box="[815,937,1592,1616]" italics="true" pageId="15" pageNumber="14">E. candiota</emphasis>
Birula, 1903
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1A4D430302CFF9CD" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[1093,1438,1592,1616]" class="Arachnida" family="Scorpionidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="53C1EAA7DD30E31A1A4D4303038EF9CD" box="[1093,1247,1592,1616]" italics="true" pageId="15" pageNumber="14">E. carpathicus</emphasis>
(Linnaeus, 1767)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1BA2430304C6F9ED" italics="true" pageId="15" pageNumber="14">E. ciliciensis</emphasis>
Birula, 1898; 
<taxonomicName id="A6B54D36DD30E31A1A3F436302CCF9EC" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[1079,1437,1624,1649]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="53C1EAA7DD30E31A1A3F43630390F9ED" box="[1079,1217,1625,1648]" italics="true" pageId="15" pageNumber="14">E. concinnus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1BA243630293F9ED" box="[1450,1474,1625,1648]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
<taxonomicName id="A6B54D36DD30E31A1D2743420311F90C" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[815,1088,1656,1681]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD30E31A1D27434204DDF90D" box="[815,908,1656,1680]" italics="true" pageId="15" pageNumber="14">deltshevi</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1A46434302CCF90D" authority="Tropea et al., 2015" authorityName="Tropea" authorityYear="2015" box="[1102,1437,1656,1680]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="drenskii">
<emphasis id="53C1EAA7DD30E31A1A4643430390F90D" box="[1102,1217,1656,1680]" italics="true" pageId="15" pageNumber="14">E. drenskii</emphasis>
<bibRefCitation id="05244B44DD30E31A1AC3434302CCF90D" author="TROPEA" box="[1227,1437,1656,1680]" pageId="15" pageNumber="14" pagination="1 - 16" refId="ref7987" refString="TROPEA, G. &amp; V. FET. 2015. Two new Euscorpius species from central-western Greece (Scorpiones: Euscorpiidae). Euscorpius, 199: 1 - 16." type="journal article" year="2015">Tropea et al., 2015</bibRefCitation>
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1BA243430401F92C" italics="true" pageId="15" pageNumber="14">E. feti</emphasis>
Tropea, 2013; 
<taxonomicName id="A6B54D36DD30E31A1DE543A30213F92D" authority="Tropea &amp; Fet, 2015" authorityName="Tropea &amp; Fet" authorityYear="2015" box="[1005,1346,1688,1713]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="giachinoi">
<emphasis id="53C1EAA7DD30E31A1DE543A3033CF92D" box="[1005,1133,1688,1712]" italics="true" pageId="15" pageNumber="14">E. giachinoi</emphasis>
Tropea &amp; Fet, 2015
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1B4443A30290F92D" authority="Of" authorityName="Of" box="[1356,1473,1689,1712]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gocmeni">
<emphasis id="53C1EAA7DD30E31A1B4443A30290F92D" box="[1356,1473,1689,1712]" italics="true" pageId="15" pageNumber="14">E. gocmeni</emphasis>
</taxonomicName>
Tropea et al., 2014; 
<taxonomicName id="A6B54D36DD30E31A1A0E4383021AF94D" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" box="[1030,1355,1720,1744]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="hadzii">
<emphasis id="53C1EAA7DD30E31A1A0E43830335F94D" box="[1030,1124,1720,1744]" italics="true" pageId="15" pageNumber="14">E. hadzii</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1B5E4383049AF96C" authority="(Herbst, 1800)" baseAuthorityName="Herbst" baseAuthorityYear="1800" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="53C1EAA7DD30E31A1B5E43830290F94D" box="[1366,1473,1720,1744]" italics="true" pageId="15" pageNumber="14">E. italicus</emphasis>
(Herbst, 1800)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1DDD43E30267F96C" authority="Tropea et al., 2014" authorityName="Tropea" authorityYear="2014" box="[981,1334,1752,1777]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="kinzelbachi">
<emphasis id="53C1EAA7DD30E31A1DDD43E30338F96D" box="[981,1129,1752,1776]" italics="true" pageId="15" pageNumber="14">E. kinzelbachi</emphasis>
Tropea et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1B4843E30290F96D" box="[1344,1473,1752,1776]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="kritscheri">
<emphasis id="53C1EAA7DD30E31A1B4843E30290F96D" box="[1344,1473,1752,1776]" italics="true" pageId="15" pageNumber="14">E. kritscheri</emphasis>
</taxonomicName>
<bibRefCitation id="05244B44DD30E31A1D2743C30480F88C" author="FET" box="[815,977,1784,1809]" pageId="15" pageNumber="14" pagination="3 - 10" refId="ref6804" refString="FET, V., M. E. SOLEGLAD, A. PARMAKELIS, P. KOTSAKIOZI &amp; I. STATHI. 2013. A new species of Euscorpius from Tinos Island, Greece (Scorpiones: Euscorpiidae). Revista Iberica de Aracnologia, 23: 3 - 10." type="journal article" year="2013">Fet et al., 2013</bibRefCitation>
; 
<taxonomicName id="A6B54D36DD30E31A1DD443C30240F88C" authority="Yagmur et al., 2013" authorityName="Yagmur" authorityYear="2013" box="[988,1297,1784,1809]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="lycius">
<emphasis id="53C1EAA7DD30E31A1DD443C30366F88D" box="[988,1079,1784,1808]" italics="true" pageId="15" pageNumber="14">E. lycius</emphasis>
Yağmur et al., 2013
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1B1443C30290F88D" authority="Fet Fet &amp; Soleglad, 2002" authorityName="Fet Fet &amp; Soleglad" authorityYear="2002" box="[1308,1473,1784,1808]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mylonasi">
<emphasis id="53C1EAA7DD30E31A1B1443C302C9F88D" box="[1308,1432,1784,1808]" italics="true" pageId="15" pageNumber="14">E. mylonasi</emphasis>
Fet
</taxonomicName>
et al., 2014; 
<taxonomicName id="A6B54D36DD30E31A1DBA42230254F8AD" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" box="[946,1285,1816,1840]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="oglasae">
<emphasis id="53C1EAA7DD30E31A1DBA42230370F8AD" box="[946,1057,1816,1840]" italics="true" pageId="15" pageNumber="14">E. oglasae</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1B18422304E5F8CC" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="53C1EAA7DD30E31A1B184223022AF8AD" box="[1296,1403,1817,1840]" italics="true" pageId="15" pageNumber="14">E. sicanus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1DB54203038BF8CC" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[957,1242,1848,1873]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD30E31A1DB542030368F8CD" box="[957,1081,1848,1872]" italics="true" pageId="15" pageNumber="14">E. solegladi</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1AEB42030293F8CD" authority="Tropea Gantenbein et al., 1999" authorityName="Tropea Gantenbein" authorityYear="1999" box="[1251,1474,1848,1872]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="stahlavskyi">
<emphasis id="53C1EAA7DD30E31A1AEB42030222F8CD" box="[1251,1395,1848,1872]" italics="true" pageId="15" pageNumber="14">E. stahlavskyi</emphasis>
Tropea
</taxonomicName>
et al., 2014; 
<taxonomicName id="A6B54D36DD30E31A1DA6426303BCF8EC" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[942,1261,1880,1905]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="tauricus">
<emphasis id="53C1EAA7DD30E31A1DA642630370F8ED" box="[942,1057,1881,1904]" italics="true" pageId="15" pageNumber="14">E. tauricus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1AFF426304E8F80C" authority="(C.L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="53C1EAA7DD30E31A1AFF426302D5F8ED" box="[1271,1412,1881,1904]" italics="true" pageId="15" pageNumber="14">E. tergestinus</emphasis>
(C.L. Koch, 1837)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD30E31A1DCE4243048FF80D" box="[966,990,1913,1936]" italics="true" pageId="15" pageNumber="14">E.</emphasis>
sp. 
<taxonomicName id="A6B54D36DD30E31A1A06424203F5F80D" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[1038,1188,1912,1936]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="tergestinus">
cf. 
<emphasis id="53C1EAA7DD30E31A1A3B424303F5F80D" box="[1075,1188,1913,1936]" italics="true" pageId="15" pageNumber="14">tergestinus</emphasis>
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD30E31A1ABD42430434F82D" authority="Tropea &amp; Fet, 2015" authorityName="Tropea &amp; Fet" authorityYear="2015" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="vailatii">
<emphasis id="53C1EAA7DD30E31A1ABD4243024EF80D" box="[1205,1311,1912,1936]" italics="true" pageId="15" pageNumber="14">E. vailatii</emphasis>
Tropea &amp; Fet, 2015
</taxonomicName>
) (
<figureCitation id="F98E2A30DD30E31A1D7342A204AAF82D" box="[891,1019,1944,1968]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="15" pageNumber="14">Figs. 51–57</figureCitation>
; Fet &amp; Soleglad, 2002; 
<bibRefCitation id="05244B44DD30E31A1AFC42A302C4F82C" author="FET" box="[1268,1429,1944,1969]" pageId="15" pageNumber="14" pagination="3 - 10" refId="ref6804" refString="FET, V., M. E. SOLEGLAD, A. PARMAKELIS, P. KOTSAKIOZI &amp; I. STATHI. 2013. A new species of Euscorpius from Tinos Island, Greece (Scorpiones: Euscorpiidae). Revista Iberica de Aracnologia, 23: 3 - 10." type="journal article" year="2013">Fet et al., 2013</bibRefCitation>
; Fet et al., 2014; 
<bibRefCitation id="05244B44DD30E31A1DB9428203F6F84C" author="GANTENBEIN" box="[945,1191,1976,2001]" pageId="15" pageNumber="14" pagination="13 - 43" refId="ref7109" refString="GANTENBEIN, B., M. E. SOLEGLAD, V. FET, P. CRUCITTI &amp; E. V. FET. 2002. Euscorpius naupliensis (C. L. Koch, 1837) (Scorpiones: Euscorpiidae) from Greece: elevation to the species level justified by molecular and morphological data. Revista Iberica de Aracnologia, 6: 13 - 43." type="journal article" year="2002">Gantenbein et al., 2002</bibRefCitation>
; 
<bibRefCitation id="05244B44DD30E31A1AB94282023FF84C" author="JACOB" box="[1201,1390,1976,2001]" pageId="15" pageNumber="14" pagination="72 - 84" refId="ref7178" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 a. Morphology and function of male genitalia (spermatophores) in Euscorpius italicus (Euscorpiidae, Scorpiones): complex spermatophore structures enable safe sperm transfer. Journal of Morphology, 260: 72 - 84." type="journal article" year="2004">Jacob et al, 2004a</bibRefCitation>
, 
<bibRefCitation id="05244B44DD30E31A1B70428202ECF84D" author="JACOB" box="[1400,1469,1976,2000]" pageId="15" pageNumber="14" pagination="59 - 72" refId="ref7240" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 b. Complex male genitalia (hemispermatophores) are not diagnostic for cryptic species in the genus Euscorpius (Scorpiones: Euscorpiidae). Organisms Diversity &amp; Evolution, 4: 59 - 72." type="journal article" year="2004">2004b</bibRefCitation>
; 
<bibRefCitation id="05244B44DD30E31A1D2742E204B5F86C" author="SCHERABON" box="[815,996,2008,2033]" pageId="15" pageNumber="14" pagination="77 - 154" refId="ref7825" refString="SCHERABON, B. 1987. Die Skorpione Osterreiches in vergleichender Sicht unter besonderer Berucksichtigung Karntens. Carinthia II, 45: 77 - 154." type="journal article" year="1987">Scherabon, 1987</bibRefCitation>
; Tropea et al., 2014a, 2014b, 2015b, 2015c; Tropea &amp; Fet, 2015; Tropea &amp; Ozimec, 2019; Vignoli et al., 2007; Yağmur et al., 2013; Yağmur &amp; Tropea, 2013). The 
<emphasis id="53C1EAA7DD2FE3051CE744310452FEBE" box="[751,771,267,291]" italics="true" pageId="16" pageNumber="15">bl</emphasis>
was not directly reported in two other species. 
<bibRefCitation id="05244B44DD2FE3051C6744110788FEFE" author="GANTENBEIN" pageId="16" pageNumber="15" pagination="13 - 43" refId="ref7109" refString="GANTENBEIN, B., M. E. SOLEGLAD, V. FET, P. CRUCITTI &amp; E. V. FET. 2002. Euscorpius naupliensis (C. L. Koch, 1837) (Scorpiones: Euscorpiidae) from Greece: elevation to the species level justified by molecular and morphological data. Revista Iberica de Aracnologia, 6: 13 - 43." type="journal article" year="2002">Gantenbein et al. (2002)</bibRefCitation>
described and illustrated the hemispermatophore of 
<taxonomicName id="A6B54D36DD2FE3051E7944510683FE1E" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[113,466,363,387]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="naupliensis">
<emphasis id="53C1EAA7DD2FE3051E79445107D0FE1F" box="[113,129,363,386]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051E8644510654FE1E" box="[142,261,363,387]" italics="true" pageId="16" pageNumber="15">naupliensis</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
, which seemed to lack the 
<emphasis id="53C1EAA7DD2FE3051CE0445105ADFE1E" box="[744,764,363,387]" italics="true" pageId="16" pageNumber="15">bl</emphasis>
, but their text and figure were inconclusive. Tropea et al. (2013) omitted mention of the 
<emphasis id="53C1EAA7DD2FE3051F6A44910627FE5E" box="[354,374,427,451]" italics="true" pageId="16" pageNumber="15">bl</emphasis>
in 
<taxonomicName id="A6B54D36DD2FE3051F9D44910563FE5E" box="[405,562,427,451]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="erymanthius">
<emphasis id="53C1EAA7DD2FE3051F9D449106F4FE5F" box="[405,421,427,450]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051FB944910563FE5E" box="[433,562,427,451]" italics="true" pageId="16" pageNumber="15">erymanthius</emphasis>
</taxonomicName>
, but did not exclude it, and its presence is unclear.
</paragraph>
<paragraph id="610A36B5DD2FE3051EA944D105F9FC7E" blockId="16.[113,772,234,1635]" pageId="16" pageNumber="15">
Absence of the 
<emphasis id="53C1EAA7DD2FE3051F4F44D10632FD9E" box="[327,355,491,515]" italics="true" pageId="16" pageNumber="15">ldi</emphasis>
is another character shared between 
<taxonomicName id="A6B54D36DD2FE3051CE444D10785FDBE" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD2FE3051CE444D105ADFD9F" box="[748,764,491,514]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051E7947310785FDBE" box="[113,212,523,547]" italics="true" pageId="16" pageNumber="15">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD2FE3051ED54731064AFDBF" box="[221,283,523,547]" pageId="16" pageNumber="15" rank="species">
<emphasis id="53C1EAA7DD2FE3051ED5473107A7FDBE" bold="true" box="[221,246,523,547]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051F0D47310645FDBE" bold="true" box="[261,276,523,547]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
and 
<taxonomicName id="A6B54D36DD2FE3051F59473106B0FDBE" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[337,481,523,547]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD2FE3051F59473106B0FDBE" box="[337,481,523,547]" italics="true" pageId="16" pageNumber="15">T. flavicaudis</emphasis>
</taxonomicName>
. When this lobe is weakly developed, it can be intraspecifically variable, being absent among some individuals of a species (e.g., 
<taxonomicName id="A6B54D36DD2FE3051C3B477105DDFDFE" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[563,652,587,611]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="alpha">
<emphasis id="53C1EAA7DD2FE3051C3B47710512FDFF" box="[563,579,587,610]" italics="true" pageId="16" pageNumber="15">A</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051C47477105DDFDFE" box="[591,652,587,611]" italics="true" pageId="16" pageNumber="15">alpha</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B54D36DD2FE3051C9047710455FDFF" baseAuthorityName="Di Caporiacco" baseAuthorityYear="1950" box="[664,772,587,610]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gamma">
<emphasis id="53C1EAA7DD2FE3051C90477105F9FDFF" box="[664,680,587,610]" italics="true" pageId="16" pageNumber="15">A</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051CBD47710455FDFF" box="[693,772,587,610]" italics="true" pageId="16" pageNumber="15">gamma</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B54D36DD2FE3051EAA4751067CFD1F" baseAuthorityName="C. L. Koch, Koch" baseAuthorityYear="1837" box="[162,301,619,642]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="germanus">
<emphasis id="53C1EAA7DD2FE3051EAA475107E3FD1F" box="[162,178,619,642]" italics="true" pageId="16" pageNumber="15">A</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051ECC4751067CFD1F" box="[196,301,619,642]" italics="true" pageId="16" pageNumber="15">germanus</emphasis>
</taxonomicName>
; 
<bibRefCitation id="05244B44DD2FE3051F3747510541FD1E" author="JACOB" box="[319,528,619,643]" pageId="16" pageNumber="15" pagination="59 - 72" refId="ref7240" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 b. Complex male genitalia (hemispermatophores) are not diagnostic for cryptic species in the genus Euscorpius (Scorpiones: Euscorpiidae). Organisms Diversity &amp; Evolution, 4: 59 - 72." type="journal article" year="2004">Jacob et al, 2004b</bibRefCitation>
). It has been reported to be present in 15 species of 
<taxonomicName id="A6B54D36DD2FE3051FB247B10563FD3F" authorityName="Thorell" authorityYear="1876" box="[442,562,651,674]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD2FE3051FB247B10563FD3F" box="[442,562,651,674]" italics="true" pageId="16" pageNumber="15">Euscorpius</emphasis>
</taxonomicName>
(i.e., 
<taxonomicName id="A6B54D36DD2FE3051C7B47B10607FD5F" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="balearicus">
<emphasis id="53C1EAA7DD2FE3051C7B47B10452FD3E" box="[627,771,651,675]" italics="true" pageId="16" pageNumber="15">E. balearicus</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F6A4791052BFD5E" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[354,634,683,707]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="birulai">
<emphasis id="53C1EAA7DD2FE3051F6A4791069BFD5E" box="[354,458,683,707]" italics="true" pageId="16" pageNumber="15">E. birulai</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051C8E479107AAFD7E" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="candiota">
<emphasis id="53C1EAA7DD2FE3051C8E47910452FD5E" box="[646,771,683,707]" italics="true" pageId="16" pageNumber="15">E. candiota</emphasis>
Birula, 1903
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F0147F1053BFD7E" authority="(Linnaeus, 1767)" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[265,618,715,739]" class="Arachnida" family="Scorpionidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="carpathicus">
<emphasis id="53C1EAA7DD2FE3051F0147F106F7FD7E" box="[265,422,715,739]" italics="true" pageId="16" pageNumber="15">E. carpathicus</emphasis>
(Linnaeus, 1767)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051C7F47F1061AFC9E" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="53C1EAA7DD2FE3051C7F47F10452FD7F" box="[631,771,715,738]" italics="true" pageId="16" pageNumber="15">E. concinnus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F5247D105C3FC9E" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[346,658,747,771]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD2FE3051F5247D1068BFC9E" box="[346,474,747,771]" italics="true" pageId="16" pageNumber="15">E. deltshevi</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051CA847D10605FCBF" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="hadzii">
<emphasis id="53C1EAA7DD2FE3051CA847D10452FC9E" box="[672,771,747,771]" italics="true" pageId="16" pageNumber="15">E. hadzii</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F5746310522FCBE" authority="(Herbst, 1800)" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[351,627,779,803]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="53C1EAA7DD2FE3051F574631069CFCBE" box="[351,461,779,803]" italics="true" pageId="16" pageNumber="15">E. italicus</emphasis>
(Herbst, 1800)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051C774631064DFCDE" authority="Fet et al., 2013" authorityName="Fet" authorityYear="2013" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="kritscheri">
<emphasis id="53C1EAA7DD2FE3051C7746310452FCBE" box="[639,771,779,803]" italics="true" pageId="16" pageNumber="15">E. kritscheri</emphasis>
<bibRefCitation id="05244B44DD2FE3051E794611064DFCDE" author="FET" box="[113,284,811,835]" pageId="16" pageNumber="15" pagination="3 - 10" refId="ref6804" refString="FET, V., M. E. SOLEGLAD, A. PARMAKELIS, P. KOTSAKIOZI &amp; I. STATHI. 2013. A new species of Euscorpius from Tinos Island, Greece (Scorpiones: Euscorpiidae). Revista Iberica de Aracnologia, 23: 3 - 10." type="journal article" year="2013">Fet et al., 2013</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F224611050EFCDE" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[298,607,811,835]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="mylonasi">
<emphasis id="53C1EAA7DD2FE3051F22461106FBFCDE" box="[298,426,811,835]" italics="true" pageId="16" pageNumber="15">E. mylonasi</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051C6446110660FCFF" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="oglasae">
<emphasis id="53C1EAA7DD2FE3051C644611058EFCDE" box="[620,735,811,835]" italics="true" pageId="16" pageNumber="15">E. oglasae</emphasis>
Di Caporiacco, 1950
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F324671052BFCFE" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[314,634,843,867]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="53C1EAA7DD2FE3051F32467106F9FCFF" box="[314,424,843,866]" italics="true" pageId="16" pageNumber="15">E. sicanus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051C8C4671064DFC1E" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD2FE3051C8C46710452FCFE" box="[644,771,843,867]" italics="true" pageId="16" pageNumber="15">E. solegladi</emphasis>
Fet et al., 2014
</taxonomicName>
; 
<taxonomicName id="A6B54D36DD2FE3051F21465105C2FC1E" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[297,659,875,899]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="tergestinus">
<emphasis id="53C1EAA7DD2FE3051F21465106ECFC1F" box="[297,445,875,898]" italics="true" pageId="16" pageNumber="15">E. tergestinus</emphasis>
(C.L. Koch, 1837)
</taxonomicName>
; 
<emphasis id="53C1EAA7DD2FE3051CA8465105E9FC1F" box="[672,696,875,898]" italics="true" pageId="16" pageNumber="15">E.</emphasis>
sp. 
<taxonomicName id="A6B54D36DD2FE3051CE0465107B4FC3F" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="tergestinus">
cf. 
<emphasis id="53C1EAA7DD2FE3051E7946B107B4FC3F" box="[113,229,907,930]" italics="true" pageId="16" pageNumber="15">tergestinus</emphasis>
</taxonomicName>
) (
<figureCitation id="F98E2A30DD2FE3051F0946B106D6FC3F" box="[257,391,906,931]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="16" pageNumber="15">Figs. 51–57</figureCitation>
; Fet &amp; Soleglad, 2002; 
<bibRefCitation id="05244B44DD2FE3051C9046B107FBFC5E" author="FET" pageId="16" pageNumber="15" pagination="3 - 10" refId="ref6804" refString="FET, V., M. E. SOLEGLAD, A. PARMAKELIS, P. KOTSAKIOZI &amp; I. STATHI. 2013. A new species of Euscorpius from Tinos Island, Greece (Scorpiones: Euscorpiidae). Revista Iberica de Aracnologia, 23: 3 - 10." type="journal article" year="2013">Fet et al., 2013</bibRefCitation>
; Fet et al., 2014; 
<bibRefCitation id="05244B44DD2FE3051F7A46910529FC5E" author="GANTENBEIN" box="[370,632,939,963]" pageId="16" pageNumber="15" pagination="13 - 43" refId="ref7109" refString="GANTENBEIN, B., M. E. SOLEGLAD, V. FET, P. CRUCITTI &amp; E. V. FET. 2002. Euscorpius naupliensis (C. L. Koch, 1837) (Scorpiones: Euscorpiidae) from Greece: elevation to the species level justified by molecular and morphological data. Revista Iberica de Aracnologia, 6: 13 - 43." type="journal article" year="2002">Gantenbein et al., 2002</bibRefCitation>
; 
<bibRefCitation id="05244B44DD2FE3051C8D469107E7FC7E" author="JACOB" pageId="16" pageNumber="15" pagination="72 - 84" refId="ref7178" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 a. Morphology and function of male genitalia (spermatophores) in Euscorpius italicus (Euscorpiidae, Scorpiones): complex spermatophore structures enable safe sperm transfer. Journal of Morphology, 260: 72 - 84." type="journal article" year="2004">Jacob et al, 2004a</bibRefCitation>
, 
<bibRefCitation id="05244B44DD2FE3051EC846F10657FC7E" author="JACOB" box="[192,262,971,995]" pageId="16" pageNumber="15" pagination="59 - 72" refId="ref7240" refString="JACOB, A., B. GANTENBEIN, M. E. BRAUNWALDER, W. NENTWIG &amp; C. KROPF. 2004 b. Complex male genitalia (hemispermatophores) are not diagnostic for cryptic species in the genus Euscorpius (Scorpiones: Euscorpiidae). Organisms Diversity &amp; Evolution, 4: 59 - 72." type="journal article" year="2004">2004b</bibRefCitation>
; 
<bibRefCitation id="05244B44DD2FE3051F1946F10696FC7E" author="SCHERABON" box="[273,455,971,995]" pageId="16" pageNumber="15" pagination="77 - 154" refId="ref7825" refString="SCHERABON, B. 1987. Die Skorpione Osterreiches in vergleichender Sicht unter besonderer Berucksichtigung Karntens. Carinthia II, 45: 77 - 154." type="journal article" year="1987">Scherabon, 1987</bibRefCitation>
; Vignoli et al, 2007).
</paragraph>
<paragraph id="610A36B5DD2FE3051EA946D105FBFAFF" blockId="16.[113,772,234,1635]" pageId="16" pageNumber="15">
The 
<emphasis id="53C1EAA7DD2FE3051ED346D107AAFB9E" box="[219,251,1003,1027]" italics="true" pageId="16" pageNumber="15">lde</emphasis>
appears to be almost universally present in hemispermatophores of these genera that have been described to date. However, none have been reported to exhibit apparent division into two sub-lobes. Perhaps the most proximate condition is in 
<taxonomicName id="A6B54D36DD2FE3051F04415106C4FB1E" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[268,405,1131,1155]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD2FE3051F04415106C4FB1E" box="[268,405,1131,1155]" italics="true" pageId="16" pageNumber="15">T. flavicaudis</emphasis>
</taxonomicName>
, whose 
<emphasis id="53C1EAA7DD2FE3051FE34151055AFB1E" box="[491,523,1131,1155]" italics="true" pageId="16" pageNumber="15">lde</emphasis>
is bilobate or bidentate, but undivided (
<figureCitation id="F98E2A30DD2FE3051F1241B1063CFB3F" box="[282,365,1162,1187]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="16" pageNumber="15">Fig. 50</figureCitation>
). In molecular studies, 
<taxonomicName id="A6B54D36DD2FE3051C7C41B10452FB3E" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[628,771,1163,1187]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD2FE3051C7C41B10452FB3E" box="[628,771,1163,1187]" italics="true" pageId="16" pageNumber="15">T. flavicaudis</emphasis>
</taxonomicName>
resolved as a basal lineage and outgroup that was not closely related to 
<taxonomicName id="A6B54D36DD2FE3051EDC41F10664FB7F" box="[212,309,1227,1250]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="popovi">
<emphasis id="53C1EAA7DD2FE3051EDC41F107B5FB7F" box="[212,228,1227,1250]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051EE741F10664FB7F" box="[239,309,1227,1250]" italics="true" pageId="16" pageNumber="15">popovi</emphasis>
</taxonomicName>
(
<bibRefCitation id="05244B44DD2FE3051F4841F10551FB7F" author="TROPEA" box="[320,512,1226,1251]" pageId="16" pageNumber="15" pagination="1 - 16" refId="ref7987" refString="TROPEA, G. &amp; V. FET. 2015. Two new Euscorpius species from central-western Greece (Scorpiones: Euscorpiidae). Euscorpius, 199: 1 - 16." type="journal article" year="2015">Tropea et al., 2015</bibRefCitation>
a: 11, fig. 20), which is the species most similar morphologically to 
<taxonomicName id="A6B54D36DD2FE3051C0441D105D6FA9E" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[524,647,1259,1283]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD2FE3051C0441D1054DFA9F" box="[524,540,1259,1282]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051C2F41D105D6FA9E" box="[551,647,1259,1283]" italics="true" pageId="16" pageNumber="15">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD2FE3051C8441D10594FA9F" box="[652,709,1259,1283]" pageId="16" pageNumber="15" rank="species">
<emphasis id="53C1EAA7DD2FE3051C8441D105F5FA9E" bold="true" box="[652,676,1259,1283]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051CB841D105EFFA9E" bold="true" box="[688,702,1259,1283]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
Thus, in 
<taxonomicName id="A6B54D36DD2FE3051E864031065DFABE" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" box="[142,268,1291,1315]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD2FE3051E86403107CFFABF" box="[142,158,1291,1314]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051EA44031065DFABE" box="[172,268,1291,1315]" italics="true" pageId="16" pageNumber="15">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD2FE3051F1C4031061EFABF" box="[276,335,1291,1315]" pageId="16" pageNumber="15" rank="species">
<emphasis id="53C1EAA7DD2FE3051F1C4031067DFABE" bold="true" box="[276,300,1291,1315]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051F3340310618FABE" bold="true" box="[315,329,1291,1315]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, 
<emphasis id="53C1EAA7DD2FE3051F5640310628FABE" box="[350,377,1291,1315]" italics="true" pageId="16" pageNumber="15">ldi</emphasis>
and 
<emphasis id="53C1EAA7DD2FE3051FA640310693FABE" box="[430,450,1291,1315]" italics="true" pageId="16" pageNumber="15">bl</emphasis>
may have been independently lost much later in the evolutionary history of the group. Their absence is probably the primitive state in 
<taxonomicName id="A6B54D36DD2FE3051C13407105F2FAFE" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[539,675,1355,1379]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD2FE3051C13407105F2FAFE" box="[539,675,1355,1379]" italics="true" pageId="16" pageNumber="15">T. flavicaudis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="610A36B5DD2FE3051EAA4051067DF9FE" blockId="16.[113,772,234,1635]" pageId="16" pageNumber="15">
The unusual morphology of the distal carina of 
<taxonomicName id="A6B54D36DD2FE3051CE440510784FA3E" authority="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Byronová &amp; Šťáhlavský" authorityYear="2020" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="thracicus" status="sp. nov.">
<emphasis id="53C1EAA7DD2FE3051CE4405105ADFA1F" box="[748,764,1387,1410]" italics="true" pageId="16" pageNumber="15">E</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051E7A40B10784FA3E" box="[114,213,1419,1443]" italics="true" pageId="16" pageNumber="15">thracicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="48F257DCDD2FE3051ED440B10646FA3F" box="[220,279,1419,1443]" pageId="16" pageNumber="15" rank="species">
<emphasis id="53C1EAA7DD2FE3051ED440B107A4FA3E" bold="true" box="[220,245,1419,1443]" pageId="16" pageNumber="15">sp</emphasis>
. 
<emphasis id="53C1EAA7DD2FE3051F0A40B10640FA3E" bold="true" box="[258,273,1419,1443]" pageId="16" pageNumber="15">n</emphasis>
.
</taxonomicNameLabel>
, was consistent across 6 hemispermatophores from 
<specimenCount id="77B3FD3CDD2FE3051EBA4091065CFA5E" box="[178,269,1451,1475]" pageId="16" pageNumber="15" type="male">3 males</specimenCount>
, indicating a taxonomically stable character. This underlines the importance of accurately characterizing and illustrating these structures for comparative analysis. Hemispermatophore anatomy has yet to be reported in the majority of other related euscorpiids (9 
<taxonomicName id="A6B54D36DD2FE3051C28431105F6F9DE" authorityName="Koch" authorityYear="1837" box="[544,679,1579,1603]" class="Arachnida" family="Euscorpiidae" genus="Alpiscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD2FE3051C28431105F6F9DE" box="[544,679,1579,1603]" italics="true" pageId="16" pageNumber="15">Alpiscorpius</emphasis>
</taxonomicName>
spp., 36 
<taxonomicName id="A6B54D36DD2FE3051E7A437107BBF9FF" authorityName="Thorell" authorityYear="1876" box="[114,234,1611,1634]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD2FE3051E7A437107BBF9FF" box="[114,234,1611,1634]" italics="true" pageId="16" pageNumber="15">Euscorpius</emphasis>
</taxonomicName>
spp.).
</paragraph>
</subSubSection>
<subSubSection id="29AF653EDD2FE3051E7A43B106D6F8DE" pageId="16" pageNumber="15" type="distribution">
<paragraph id="610A36B5DD2FE3051E7A43B10598F93E" blockId="16.[114,713,1675,1699]" box="[114,713,1675,1699]" pageId="16" pageNumber="15">
DISTRIBUTION. 
<collectingCountry id="19A27625DD2FE3051F1A43B1063FF93E" box="[274,366,1675,1699]" name="Bulgaria" pageId="16" pageNumber="15">Bulgaria</collectingCountry>
(eastern Rodopi Mts.) (
<figureCitation id="F98E2A30DD2FE3051C6543B105ECF93E" box="[621,701,1675,1699]" captionStart="Figure 61" captionStartId="15.[116,184,886,908]" captionTargetBox="[113,1474,234,877]" captionTargetId="figure-360@15.[113,1474,234,880]" captionTargetPageId="15" captionText="Figure 61. Map of distribution of Euscorpius populations analysed by DNA (see Table 2)." figureDoi="http://doi.org/10.5281/zenodo.4648946" httpUri="https://zenodo.org/record/4648946/files/figure.png" pageId="16" pageNumber="15">Fig. 61</figureCitation>
).
</paragraph>
<paragraph id="610A36B5DD2FE3051E7A43F106D6F8DE" blockId="16.[114,772,1738,2051]" pageId="16" pageNumber="15">
<materialsCitation id="D1DD3CE8DD2FE3051E7A43F106D6F8DE" ID-GBIF-Occurrence="3114128308" collectionCode="MATERIAL" collectorName="M. Stockmann." country="France" latitude="43.459763" location="Var Department" longLatPrecision="1" longitude="6.281816" municipality="Le Thoronet" pageId="16" pageNumber="15" specimenCount="1" specimenCount-male="1">
COMPARATIVE HEMISPERMATOPHORE 
<collectionCode id="07A4AE70DD2FE3051FEC43F10517F97E" box="[484,582,1739,1763]" pageId="16" pageNumber="15">MATERIAL</collectionCode>
(
<specimenCount id="77B3FD3CDD2FE3051C5D43F1053AF97F" box="[597,619,1739,1762]" pageId="16" pageNumber="15" type="male">♂</specimenCount>
, 
<figureCitation id="F98E2A30DD2FE3051C7E43F105A3F97F" box="[630,754,1738,1763]" captionStart="Figures50–58" captionStartId="13.[115,192,1982,2004]" captionTargetBox="[115,1472,214,1975]" captionTargetId="figure-136@13.[113,1474,211,1977]" captionTargetPageId="13" captionText="Figures50–58. Comparison of the capsule lobesof the hemispermatophoresof nine euscorpiids (anterior-convexviews).50. Tetratrichobothrius flavicaudis. 51. Euscorpius sp., nr tergestinus. 52. E. concinnus. 53. E. solegladi. 54. E. italicus. 55. E. deltshevi. 56. E. sicanus. 57. E. candiota. 58. E. thracicus sp. n., paratype No. 1864 Abbreviations: lde: distal external lobe; ldi: distal internal lobe; bl: basal lobe; cls: crownlike structure. Capsules of left hemispermatophores shown except for Fig. 57, which shows the right capsule (in mirror image for comparison). Scale bar: 500 μm (in Fig. 56) applies to Figs. 50–58." figureDoi="http://doi.org/10.5281/zenodo.4648942" httpUri="https://zenodo.org/record/4648942/files/figure.png" pageId="16" pageNumber="15">Figs. 50–57</figureCitation>
). 
<taxonomicName id="A6B54D36DD2FE3051E7A43D10533F89E" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[114,610,1771,1795]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="53C1EAA7DD2FE3051E7A43D10637F89F" box="[114,358,1771,1794]" italics="true" pageId="16" pageNumber="15">Euscorpius concinnus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051C7F43D10596F89E" bold="true" box="[631,711,1771,1795]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051C7F43D10596F89E" box="[631,711,1771,1795]" name="France" pageId="16" pageNumber="15">France</collectingCountry>
</emphasis>
, 
<location id="646A606EDD2FE3051CD643D107A2F8BF" LSID="urn:lsid:plazi:treatment:E91C87A3DD3BE3051F1C437C0352FDFE:646A606EDD2FE3051CD643D107A2F8BF" country="France" latitude="43.459763" longLatPrecision="1" longitude="6.281816" municipality="Le Thoronet" name="Var Department" pageId="16" pageNumber="15">Var Department</location>
, 
<collectorName id="CC405363DD2FE3051F0D423106C7F8BE" box="[261,406,1803,1827]" pageId="16" pageNumber="15">
<collectingMunicipality id="816EACCFDD2FE3051F0D423106C7F8BE" box="[261,406,1803,1827]" pageId="16" pageNumber="15">Le Thoronet</collectingMunicipality>
</collectorName>
, 
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<geoCoordinate id="04815072DD2FE3051C4F42310598F8BE" box="[583,713,1803,1827]" degrees="6.281816" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="6.281816">6.281816°E</geoCoordinate>
, 
<collectorName id="CC405363DD2FE3051CD442310455F8BF" box="[732,772,1803,1826]" pageId="16" pageNumber="15">No.</collectorName>
1642, leg. 
<collectorName id="CC405363DD2FE3051EEA421106D6F8DE" box="[226,391,1835,1859]" pageId="16" pageNumber="15">M. Stockmann.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="29AF653EDD2FE3051E7A42710352FDFE" pageId="16" pageNumber="15" type="description">
<paragraph id="610A36B5DD2FE3051E7A427107A0F83E" blockId="16.[114,772,1738,2051]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051E7A427106BFF8FE" authority="Birula, 1903" authorityName="Birula" authorityYear="1903" box="[114,494,1867,1891]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="candiota">
<emphasis id="53C1EAA7DD2FE3051E7A42710602F8FE" box="[114,339,1867,1891]" italics="true" pageId="16" pageNumber="15">Euscorpius candiota</emphasis>
<materialsCitation id="D1DD3CE8DD2FE3051F68427106BFF8FE" ID-GBIF-Occurrence="3114128306" box="[352,494,1867,1891]" collectorName="No. &amp; M. Stockmann." country="Greece" latitude="35.149605" location="Lassithi Plateau" longLatPrecision="1" longitude="25.50466" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Kriti">Birula, 1903</materialsCitation>
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051FF74270051FF8FF" bold="true" box="[511,590,1866,1890]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051FF74270051FF8FF" box="[511,590,1866,1890]" name="Greece" pageId="16" pageNumber="15">Greece</collectingCountry>
</emphasis>
, 
<collectingRegion id="A371F857DD2FE3051C6A427105CFF8FE" box="[610,670,1867,1891]" country="Greece" name="Kriti" pageId="16" pageNumber="15">Crete</collectingRegion>
, 
<location id="646A606EDD2FE3051CA742710792F81E" LSID="urn:lsid:plazi:treatment:E91C87A3DD3BE3051F1C437C0352FDFE:646A606EDD2FE3051CA742710792F81E" country="Greece" latitude="35.149605" longLatPrecision="1" longitude="25.50466" name="Lassithi Plateau" pageId="16" pageNumber="15" stateProvince="Kriti">Lassithi Plateau</location>
, 
<geoCoordinate id="04815072DD2FE3051ED242500639F81F" box="[218,360,1898,1922]" degrees="35.149605" direction="north" orientation="latitude" pageId="16" pageNumber="15" precision="1" value="35.149605">35.149605°N</geoCoordinate>
<geoCoordinate id="04815072DD2FE3051F744250055BF81F" box="[380,522,1898,1922]" degrees="25.504660" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="25.50466">25.504660°E</geoCoordinate>
, 
<collectorName id="CC405363DD2FE3051C2A42510518F81F" box="[546,585,1899,1922]" pageId="16" pageNumber="15">No.</collectorName>
1728, leg. 
<collectorName id="CC405363DD2FE3051CED425107A0F83E" pageId="16" pageNumber="15">M. Stockmann.</collectorName>
</paragraph>
<paragraph id="610A36B5DD2FE3051E7A42910584F79E" blockId="16.[114,772,1738,2051]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051E7A429106BDF85E" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[114,492,1963,1987]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="deltshevi">
<emphasis id="53C1EAA7DD2FE3051E7A4291061CF85E" box="[114,333,1963,1987]" italics="true" pageId="16" pageNumber="15">Euscorpius deltshevi</emphasis>
<materialsCitation id="D1DD3CE8DD2FE3051F59429106BDF85E" ID-GBIF-Occurrence="3114128304" box="[337,492,1963,1987]" collectingDate="2020-10-02" collectorName="No. &amp; O. Vanek." country="Bulgaria" latitude="42.940334" location="Dragoman Municipality" longLatPrecision="1" longitude="22.966797" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Sofiya">Fet et al., 2014</materialsCitation>
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051FFC42900506F85F" bold="true" box="[500,599,1962,1986]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051FFC42900506F85F" box="[500,599,1962,1986]" name="Bulgaria" pageId="16" pageNumber="15">Bulgaria</collectingCountry>
</emphasis>
, 
<collectingRegion id="A371F857DD2FE3051C6B42910451F85E" box="[611,768,1963,1987]" country="Bulgaria" name="Sofiya" pageId="16" pageNumber="15">Sofia Province</collectingRegion>
, 
<location id="646A606EDD2FE3051E7A42F10622F87E" LSID="urn:lsid:plazi:treatment:E91C87A3DD3BE3051F1C437C0352FDFE:646A606EDD2FE3051E7A42F10622F87E" box="[114,371,1995,2019]" country="Bulgaria" latitude="42.940334" longLatPrecision="1" longitude="22.966797" name="Dragoman Municipality" pageId="16" pageNumber="15" stateProvince="Sofiya">Dragoman Municipality</location>
, Petrovski Krast Mtn., 
<geoCoordinate id="04815072DD2FE3051C6F42F10452F87E" box="[615,771,1995,2019]" degrees="42.9403326" direction="north" orientation="latitude" pageId="16" pageNumber="15" precision="1" value="42.940334">42.9403326°N</geoCoordinate>
<geoCoordinate id="04815072DD2FE3051E7A42D1065FF79E" box="[114,270,2027,2051]" degrees="22.9667976" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="22.966797">22.9667976°E</geoCoordinate>
, 
<date id="150B1075DD2FE3051F1142D10690F79E" box="[281,449,2027,2051]" pageId="16" pageNumber="15" value="2020-10-02">
<collectingDate id="054FE99DDD2FE3051F1142D10690F79E" box="[281,449,2027,2051]" pageId="16" pageNumber="15" value="2020-10-02">2 October 2020</collectingDate>
</date>
, 
<collectorName id="CC405363DD2FE3051FC442D106A5F79F" box="[460,500,2027,2050]" pageId="16" pageNumber="15">No.</collectorName>
1867, leg. 
<collectorName id="CC405363DD2FE3051C6242D10584F79E" box="[618,725,2027,2051]" pageId="16" pageNumber="15">O. Vaněk.</collectorName>
</paragraph>
<paragraph id="610A36B5DD2FE3051D3945D10277FEBE" blockId="16.[817,1475,234,611]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051D3945D103F8FE9E" authority="(Herbst, 1800)" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[817,1193,235,259]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="italicus">
<emphasis id="53C1EAA7DD2FE3051D3945D10351FE9E" box="[817,1024,235,259]" italics="true" pageId="16" pageNumber="15">Euscorpius italicus</emphasis>
(Herbst, 1800)
</taxonomicName>
, 
<materialsCitation id="D1DD3CE8DD2FE3051ABE45D00272FEBE" ID-GBIF-Occurrence="3114128303" country="Croatia" latitude="45.06642" location="Presika" longLatPrecision="1" longitude="14.138763" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Istarska">
<emphasis id="53C1EAA7DD2FE3051ABE45D0025CFE9F" bold="true" box="[1206,1293,234,258]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051ABE45D0025CFE9F" box="[1206,1293,234,258]" name="Croatia" pageId="16" pageNumber="15">Croatia</collectingCountry>
</emphasis>
, 
<collectingRegion id="A371F857DD2FE3051B1445D10204FE9E" box="[1308,1365,235,259]" country="Croatia" name="Istarska" pageId="16" pageNumber="15">Istria</collectingRegion>
, 
<quantity id="A64D9B50DD2FE3051B6A45D102CBFE9E" box="[1378,1434,235,259]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="15" unit="km" value="2.0">2 km</quantity>
SE 
<location id="646A606EDD2FE3051D39443104D3FEBE" LSID="urn:lsid:plazi:treatment:E91C87A3DD3BE3051F1C437C0352FDFE:646A606EDD2FE3051D39443104D3FEBE" box="[817,898,267,291]" country="Croatia" latitude="45.06642" longLatPrecision="1" longitude="14.138763" name="Presika" pageId="16" pageNumber="15" stateProvince="Istarska">Presika</location>
, 
<geoCoordinate id="04815072DD2FE3051D844431034AFEBE" box="[908,1051,267,291]" degrees="45.066421" direction="north" orientation="latitude" pageId="16" pageNumber="15" precision="1" value="45.06642">45.066421°N</geoCoordinate>
<geoCoordinate id="04815072DD2FE3051A2A443103E0FEBE" box="[1058,1201,267,291]" degrees="14.138763" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="14.138763">14.138763°E</geoCoordinate>
, No. 1646
</materialsCitation>
.
</paragraph>
<paragraph id="610A36B5DD2FE3051D39441102E9FE1E" blockId="16.[817,1475,234,611]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051D39441104F8FEDF" authorityName="Thorell" authorityYear="1876" box="[817,937,299,322]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="53C1EAA7DD2FE3051D39441104F8FEDF" box="[817,937,299,322]" italics="true" pageId="16" pageNumber="15">Euscorpius</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="A6B54D36DD2FE3051DD14411033FFEDF" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[985,1134,299,323]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="tergestinus">
cf. 
<emphasis id="53C1EAA7DD2FE3051DF24411033FFEDF" box="[1018,1134,299,322]" italics="true" pageId="16" pageNumber="15">tergestinus</emphasis>
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051A7244100380FEDF" bold="true" box="[1146,1233,298,322]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051A7244100380FEDF" box="[1146,1233,298,322]" name="Croatia" pageId="16" pageNumber="15">Croatia</collectingCountry>
</emphasis>
, Krk Island, No. 1481. 
<taxonomicName id="A6B54D36DD2FE3051D394471039FFEFE" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[817,1230,331,355]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="sicanus">
<emphasis id="53C1EAA7DD2FE3051D39447104AFFEFF" box="[817,1022,331,354]" italics="true" pageId="16" pageNumber="15">Euscorpius sicanus</emphasis>
(C. L. Koch, 1837)
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051AD04471025CFEFE" bold="true" box="[1240,1293,331,355]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051AD04471025CFEFE" box="[1240,1293,331,355]" name="Italy" pageId="16" pageNumber="15">Italy</collectingCountry>
</emphasis>
, 
<collectingRegion id="A371F857DD2FE3051B1144710209FEFE" box="[1305,1368,331,355]" country="Italy" name="Sicilia" pageId="16" pageNumber="15">Sicily</collectingRegion>
, Syracuse Province, Noto Antica, 
<date id="150B1075DD2FE3051A22445103E7FE1E" box="[1066,1206,363,387]" pageId="16" pageNumber="15" value="2014-08">August 2014</date>
, No. 881, leg. C. Turiel.
</paragraph>
<paragraph id="610A36B5DD2FE3051D3944B1042BFD9E" blockId="16.[817,1475,234,611]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051D3944B103ECFE3E" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" box="[817,1213,395,419]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="solegladi">
<emphasis id="53C1EAA7DD2FE3051D3944B10343FE3E" box="[817,1042,395,419]" italics="true" pageId="16" pageNumber="15">Euscorpius solegladi</emphasis>
<materialsCitation id="D1DD3CE8DD2FE3051A1144B103ECFE3E" ID-GBIF-Occurrence="3114128302" box="[1049,1213,395,419]" collectingDate="2020-10" collectionCode="T" country="Bulgaria" county="Petrich Municipality" latitude="41.64618" location="Kozhuch Mt." longLatPrecision="1" longitude="23.361132" municipality="Rupite" pageId="16" pageNumber="15" specimenCount="1" stateProvince="Blagoevgrad">Fet et al., 2014</materialsCitation>
</taxonomicName>
, 
<emphasis id="53C1EAA7DD2FE3051AC144B0027DFE3F" bold="true" box="[1225,1324,394,418]" pageId="16" pageNumber="15">
<collectingCountry id="19A27625DD2FE3051AC144B0027DFE3F" box="[1225,1324,394,418]" name="Bulgaria" pageId="16" pageNumber="15">Bulgaria</collectingCountry>
</emphasis>
, 
<collectingRegion id="A371F857DD2FE3051B3244B104C2FE5E" country="Bulgaria" name="Blagoevgrad" pageId="16" pageNumber="15">Blagoevgrad Province</collectingRegion>
, 
<collectingCounty id="886B4E39DD2FE3051DA6449103CDFE5E" box="[942,1180,427,451]" pageId="16" pageNumber="15">Petrich Municipality</collectingCounty>
, 
<collectingMunicipality id="816EACCFDD2FE3051AB044910253FE5E" box="[1208,1282,427,451]" pageId="16" pageNumber="15">Rupite</collectingMunicipality>
, 
<location id="646A606EDD2FE3051B16449102EFFE5F" LSID="urn:lsid:plazi:treatment:E91C87A3DD3BE3051F1C437C0352FDFE:646A606EDD2FE3051B16449102EFFE5F" box="[1310,1470,427,451]" country="Bulgaria" county="Petrich Municipality" latitude="41.64618" longLatPrecision="1" longitude="23.361132" municipality="Rupite" name="Kozhuch Mt." pageId="16" pageNumber="15" stateProvince="Blagoevgrad">Kozhuch Mt.</location>
, 
<geoCoordinate id="04815072DD2FE3051D3944F10491FE7E" box="[817,960,459,483]" degrees="41.646179" direction="north" orientation="latitude" pageId="16" pageNumber="15" precision="1" value="41.64618">41.646179°N</geoCoordinate>
<geoCoordinate id="04815072DD2FE3051DC144F10306FE7E" box="[969,1111,459,483]" degrees="23.361132" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="23.361132">23.361132°E</geoCoordinate>
, 
<date id="150B1075DD2FE3051A6C44F103AAFE7E" box="[1124,1275,459,483]" pageId="16" pageNumber="15" value="2020-10">
<collectingDate id="054FE99DDD2FE3051A6C44F103AAFE7E" box="[1124,1275,459,483]" pageId="16" pageNumber="15" value="2020-10">October 2020</collectingDate>
</date>
, No. 1861, leg. T. Ryšan.
</paragraph>
<paragraph id="610A36B5DD2FE3051D3947310352FDFE" blockId="16.[817,1475,234,611]" pageId="16" pageNumber="15">
<taxonomicName id="A6B54D36DD2FE3051D3947310265FDBE" authority="(De Geer, 1778)" baseAuthorityName="De Geer" baseAuthorityYear="1778" box="[817,1332,523,547]" class="Arachnida" family="Euscorpiidae" genus="Tetratrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="flavicaudis">
<emphasis id="53C1EAA7DD2FE3051D394731032DFDBE" box="[817,1148,523,547]" italics="true" pageId="16" pageNumber="15">Tetratrichobothrius flavicaudis</emphasis>
(De Geer, 1778)
</taxonomicName>
, 
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<collectingCountry id="19A27625DD2FE3051B48473102C1FDBE" box="[1344,1424,523,547]" name="France" pageId="16" pageNumber="15">France</collectingCountry>
</emphasis>
, Var Department, Gonfaron, 
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<geoCoordinate id="04815072DD2FE3051AC14711021AFDDE" box="[1225,1355,555,579]" degrees="6.294433" direction="east" orientation="longitude" pageId="16" pageNumber="15" precision="1" value="6.294433">6.294433°E</geoCoordinate>
, No. 1350, leg. M. Stockmann.
</paragraph>
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</treatment>
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