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<document id="F0FA55D959873D8A636C8A4A22EEED86" ID-DOI="10.5281/zenodo.5350190" ID-ISSN="2345-7600" ID-Zenodo-Dep="5350190" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630204682370" checkinUser="tatiana" docAuthor="Murphy, John C., Mumpuni, Lang, Ruud de, Gower, David J. &amp; Sanders, Kate L." docDate="2012" docId="E37587A27E7BFFD9FF21E977FD355D7B" docLanguage="en" docName="60rbz501-514.pdf" docOrigin="Raffles Bulletin of Zoology 60 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Brachyorrhos wallacei Murphy &amp; Mumpuni &amp; Lang &amp; Gower &amp; Sanders 2012, new species" docType="treatment" docVersion="3" lastPageNumber="511" masterDocId="1F4CFFDA7E72FFD3FFCCED2CFFC45E0C" masterDocTitle="The Moluccan Short-Tailed Snakes Of The Genus Brachyorrhos Kuhl (Squamata: Serpentes: Homalopsidae), And The Status Of Calamophis Meyer" masterLastPageNumber="514" masterPageNumber="501" pageNumber="510" updateTime="1699184417687" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1A364B0146F0FADC765C42FE13BCC06F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="35167E66D0808CC01073AF2B4E452359">
<mods:title id="BA95BF3D4BE38A6FDC7400AB5A6DC899">The Moluccan Short-Tailed Snakes Of The Genus Brachyorrhos Kuhl (Squamata: Serpentes: Homalopsidae), And The Status Of Calamophis Meyer</mods:title>
</mods:titleInfo>
<mods:name id="7BE3A306983D7DABD557C24AAFF62D10" type="personal">
<mods:role id="99FBCBA73D393D9183637F86EDF66500">
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</mods:role>
<mods:namePart id="70ED2C0F7B790799970E602A97172D6A">Murphy, John C.</mods:namePart>
</mods:name>
<mods:name id="EBCF5B9731043192A13895354FF8A719" type="personal">
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<mods:roleTerm id="F5253B2B93BABC3E81F03D6F830BB402">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F4AE46C66C90BA604F49B642EABDC79">Mumpuni</mods:namePart>
</mods:name>
<mods:name id="C691C54B2059D196E6CD2BC2D43A38B1" type="personal">
<mods:role id="10AC2CBF1E652CF03DF813BE99956A5A">
<mods:roleTerm id="4B959BF4B04E75FAC5504CEFDDF29D56">Author</mods:roleTerm>
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<mods:namePart id="97FA6EC835AECDE66B58EEF57ADFFCD7">Lang, Ruud de</mods:namePart>
</mods:name>
<mods:name id="5B0236AFA5353EEA7706A88425EE61E8" type="personal">
<mods:role id="10010008B9B0D3381823ECDCC7C583CE">
<mods:roleTerm id="67B905F4FA76CF4C22952B0CC86EDCD8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D037ED4373D18A5B8539FC8EA41D688A">Gower, David J.</mods:namePart>
</mods:name>
<mods:name id="257F7B123B1A35DBC61F990096D85A35" type="personal">
<mods:role id="5879E00EC9A7436F6F3A0528D095ED76">
<mods:roleTerm id="4EEA1DA0D0CCFEB5DFF9637A6EDA7C26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C61BE5B68715EBA694084C26BEA17AE">Sanders, Kate L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="F95E176F61190911F2B757B9CEF7FDD5">text</mods:typeOfResource>
<mods:relatedItem id="0B2B8D101151FED6A9791EA8AC0E6376" type="host">
<mods:titleInfo id="241CDFFB2DDE721761886255ED8839B5">
<mods:title id="21355E9B1B30D071649171DD744CC498">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="6F6DF069ADE425D761EDDF48D736142F">
<mods:date id="900A56937F5C98C514EAD0E1ED53B39E">2012</mods:date>
<mods:detail id="28D5A903D2BEE771FFBC2FB459E22BB6" type="pubDate">
<mods:number id="4A0B0B5085DA7BA44AB0B3B5843C48F9">2012-08-31</mods:number>
</mods:detail>
<mods:detail id="1968D137B0CEE154707B8E6BC10C6F36" type="volume">
<mods:number id="FEA1CE9666662A30E1BDFD2CF589A707">60</mods:number>
</mods:detail>
<mods:detail id="C3265B227E870EA733125C40A89B7BF4" type="issue">
<mods:number id="D40EA63973316B33BA526C7A490A33FE">2</mods:number>
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<mods:start id="6BC9FDC6A248AE2251A4D406E3C464B9">501</mods:start>
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</mods:extent>
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<mods:classification id="FE765A398AB899B9D73CEE8714C71970">journal article</mods:classification>
<mods:identifier id="C4A2A8BBD2D1DF0D6E16028EE3371B12" type="DOI">10.5281/zenodo.5350190</mods:identifier>
<mods:identifier id="9C6898AB2A5FC7479C2CEB5FC1F1CD4B" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="4E7B782B85BB752633DCA2B18C4756B4" type="Zenodo-Dep">5350190</mods:identifier>
</mods:mods>
<treatment id="E37587A27E7BFFD9FF21E977FD355D7B" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B" httpUri="http://treatment.plazi.org/id/E37587A27E7BFFD9FF21E977FD355D7B" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510">
<subSubSection id="23C6653F7E7BFFDAFF21E977FDB15A78" box="[237,629,1115,1141]" pageId="9" pageNumber="510" type="nomenclature">
<paragraph id="6B6336B47E7BFFDAFF21E977FDB15A78" blockId="9.[237,629,1115,1172]" box="[237,629,1115,1141]" pageId="9" pageNumber="510">
<heading id="302B81D87E7BFFDAFF21E977FDB15A78" bold="true" box="[237,629,1115,1141]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="510" reason="2">
<emphasis id="59A8EAA67E7BFFDAFF21E977FDB15A78" bold="true" box="[237,629,1115,1141]" pageId="9" pageNumber="510">
<taxonomicName id="ACDC4D377E7BFFDAFF21E977FE235A78" authority="Murphy &amp; Mumpuni &amp; Lang &amp; Gower &amp; Sanders, 2012" authorityName="Murphy &amp; Mumpuni &amp; Lang &amp; Gower &amp; Sanders" authorityYear="2012" box="[237,487,1115,1140]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="wallacei" status="sp. nov.">
<emphasis id="59A8EAA67E7BFFDAFF21E977FE235A78" bold="true" box="[237,487,1115,1140]" italics="true" pageId="9" pageNumber="510">Brachyorrhos wallacei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="429B57DD7E7BFFDAFE3CE971FDB15A78" box="[496,629,1116,1141]" pageId="9" pageNumber="510" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFDAFE4DE950FE245A98" box="[385,480,1148,1172]" pageId="9" pageNumber="510" type="description">
<paragraph id="6B6336B47E7BFFDAFE4DE950FE245A98" blockId="9.[237,629,1115,1172]" box="[385,480,1148,1172]" pageId="9" pageNumber="510">
(
<figureCitation id="F3E72A317E7BFFDAFE45E950FE1C5A98" box="[393,472,1148,1172]" captionStart="Fig" captionStartId="7.[115,150,2033,2055]" captionTargetBox="[113,1451,1076,2011]" captionTargetId="figure-66@7.[113,1452,1076,2011]" captionTargetPageId="7" captionText="Fig. 4. A comparison of the labial architecture for four species of Brachyorrhos: (A) B. albus (FMNH 134323) Ambon; (B) B. gastrotaenius (BMNH 6312425) Buru holotype; (C) B. raffrayi (MNHN 5174) Ternate, holotype; (D) B. wallacei (MZB 3463) Halmahera holotype. Labials are numbered, T denotes tallest labial; P = preocular; PFL = prefrontal-loreal shield; PLP = preocular-loreal-prefrontal shield." figureDoi="http://doi.org/10.5281/zenodo.5350198" httpUri="https://zenodo.org/record/5350198/files/figure.png" pageId="9" pageNumber="510">Fig. 4d</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFDAFFBDE990FE455B07" pageId="9" pageNumber="510" type="reference_group">
<paragraph id="6B6336B47E7BFFDAFFBDE990FDCE5ADD" blockId="9.[113,752,1211,1291]" box="[113,522,1211,1233]" pageId="9" pageNumber="510">
<treatmentCitationGroup id="4BCC119A7E7BFFDAFFBDE990FDCE5ADD" box="[113,522,1211,1233]" pageId="9" pageNumber="510">
<treatmentCitation id="EA7D10A57E7BFFDAFFBDE990FDCE5ADD" author="Boulenger, G" box="[113,522,1211,1233]" page="305" pageId="9" pageNumber="510" year="1893">
<taxonomicName id="ACDC4D377E7BFFDAFFBDE990FDCE5ADD" authority="Boulenger, 1893: 305" authorityName="Boulenger" authorityPageNumber="305" authorityYear="1893" box="[113,522,1211,1233]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="albus">
<emphasis id="59A8EAA67E7BFFDAFFBDE990FEF55ADC" box="[113,305,1212,1232]" italics="true" pageId="9" pageNumber="510">Brachyorrhos albus</emphasis>
<bibRefCitation id="0F4D4B457E7BFFDAFEF4E997FDCE5ADD" author="Boulenger, G" box="[312,522,1211,1233]" pageId="9" pageNumber="510" refId="ref9837" refString="Boulenger, G. A., 1893. Catalogue of the Snakes in the British Museum. Volume 2. Trustees of the British Museum, London. 448 pp." type="book" year="1893">Boulenger, 1893: 305</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B6336B47E7BFFDAFFBDE9F5FE455B07" blockId="9.[113,752,1211,1291]" pageId="9" pageNumber="510">
<taxonomicName id="ACDC4D377E7BFFDAFFBDE9F5FDF45AE2" authority="Setiadi &amp; Hamidy, 2006" authorityName="Setiadi &amp; Hamidy" authorityYear="2006" box="[113,560,1240,1262]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="albus">
<emphasis id="59A8EAA67E7BFFDAFFBDE9F5FEF05AE1" box="[113,308,1241,1261]" italics="true" pageId="9" pageNumber="510">Brachyorrhos albus</emphasis>
Setiadi &amp; Hamidy, 2006
</taxonomicName>
(two un-numbered photographs, no pagination)
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFDAFFBDE801FF6B5BD8" pageId="9" pageNumber="510" type="materials_examined">
<paragraph id="6B6336B47E7BFFDAFFBDE801FF6B5BD8" blockId="9.[113,752,1325,1492]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDE801FEA65B48" bold="true" box="[113,354,1325,1348]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDE801FEFE5B4F" bold="true" box="[113,314,1325,1347]" italics="true" pageId="9" pageNumber="510">Material Examined</emphasis>
. —
</emphasis>
<materialsCitation id="DBB43CE97E7BFFDAFEA0E801FF6B5BD8" collectionCode="MZB, USNM" county="Weda Bay" location="Halmahera" municipality="Weda Bay Mines" pageId="9" pageNumber="510" specimenCode="MZB 3463, MZB 3464, MZB 3462, USNM 237145-150, MZB 2111, MZB 4487" specimenCount="10" specimenCount-adult="10" typeStatus="holotype">
<emphasis id="59A8EAA67E7BFFDAFEA0E801FE0A5B4F" bold="true" box="[364,462,1325,1347]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFEA0E801FE0C5B4F" bold="true" box="[364,456,1325,1347]" italics="true" pageId="9" pageNumber="510">
<typeStatus id="B46788167E7BFFDAFEA0E801FE0C5B4F" box="[364,456,1325,1347]" pageId="9" pageNumber="510" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="3B7A9ECF7E7BFFDAFE14E803FD835B48" box="[472,583,1326,1348]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="510" type="Museum">MZB 3463</specimenCode>
an adult female with a total length of
<quantity id="AC249B517E7BFFDAFE89E867FE515B6D" box="[325,405,1355,1377]" metricMagnitude="-1" metricUnit="m" metricValue="7.22" pageId="9" pageNumber="510" unit="mm" value="722.0">722 mm</quantity>
from Jailolo, Halmahera. This new species is based on
<specimenCount id="7DDAFD3D7E7BFFDAFEE0E844FE475B72" box="[300,387,1384,1406]" pageId="9" pageNumber="510" type="adult">10 adults</specimenCount>
from the island of
<location id="6E03606F7E7BFFDAFDF5E844FD665B72" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B:6E03606F7E7BFFDAFDF5E844FD665B72" box="[569,674,1384,1406]" county="Weda Bay" municipality="Weda Bay Mines" name="Halmahera" pageId="9" pageNumber="510">Halmahera</location>
(
<location id="6E03606F7E7BFFDAFD62E844FD2B5B72" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B:6E03606F7E7BFFDAFD62E844FD2B5B72" box="[686,751,1384,1406]" county="Weda Bay" municipality="Weda Bay Mines" name="Jailolo" pageId="9" pageNumber="510">Jailolo</location>
–
<specimenCode id="3B7A9ECF7E7BFFDAFF49E8A9FF375B97" box="[133,243,1413,1435]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="510" type="Museum">MZB 3464</specimenCode>
,
<specimenCode id="3B7A9ECF7E7BFFDAFF33E8A9FEA95B97" box="[255,365,1413,1435]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="510" type="Museum">MZB 3462</specimenCode>
;
<location id="6E03606F7E7BFFDAFEB5E8A9FE7F5B97" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B:6E03606F7E7BFFDAFEB5E8A9FE7F5B97" box="[377,443,1413,1435]" county="Weda Bay" municipality="Weda Bay Mines" name="Jailolo" pageId="9" pageNumber="510">Jailolo</location>
,
<location id="6E03606F7E7BFFDAFE0BE8A9FD4A5B97" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B:6E03606F7E7BFFDAFE0BE8A9FD4A5B97" box="[455,654,1413,1435]" county="Weda Bay" municipality="Weda Bay Mines" name="Kampung PasirPatih" pageId="9" pageNumber="510">Kampung PasirPatih</location>
–
<specimenCode id="3B7A9ECF7E7BFFDAFD66E8A9FF295BBB" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="510" type="Museum">USNM 237145–150</specimenCode>
;
<location id="6E03606F7E7BFFDAFF36E88DFE935BBB" LSID="urn:lsid:plazi:treatment:E37587A27E7BFFD9FF21E977FD355D7B:6E03606F7E7BFFDAFF36E88DFE935BBB" box="[250,343,1441,1463]" county="Weda Bay" municipality="Weda Bay Mines" name="Sidangoli" pageId="9" pageNumber="510">Sidangoli</location>
–
<specimenCode id="3B7A9ECF7E7BFFDAFEB9E88EFE215BBB" box="[373,485,1442,1463]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="510" type="Museum">MZB 2111</specimenCode>
;
<collectingMunicipality id="8B07ACCE7E7BFFDAFE3DE88DFD595BBB" box="[497,669,1441,1463]" pageId="9" pageNumber="510">
<collectingCounty id="82024E387E7BFFDAFE3DE88DFD935BBB" box="[497,599,1441,1463]" pageId="9" pageNumber="510">Weda Bay</collectingCounty>
Mines
</collectingMunicipality>
–
<specimenCode id="3B7A9ECF7E7BFFDAFD77E88EFF615BD8" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="510" type="Museum">MZB 4487</specimenCode>
.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFDAFFBDE8D0FDFC5918" pageId="9" pageNumber="510" type="diagnosis">
<paragraph id="6B6336B47E7BFFDAFFBDE8D0FDFC5918" blockId="9.[113,752,1532,1812]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDE8D0FF225819" bold="true" box="[113,230,1532,1557]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDE8D0FF1B5818" bold="true" box="[113,223,1532,1556]" italics="true" pageId="9" pageNumber="510">Diagnosis</emphasis>
.
</emphasis>
— A
<taxonomicName id="ACDC4D377E7BFFDAFEFCE8D2FE015819" box="[304,453,1534,1557]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="59A8EAA67E7BFFDAFEFCE8D2FE015819" box="[304,453,1534,1557]" italics="true" pageId="9" pageNumber="510">Brachyorrhos</emphasis>
</taxonomicName>
with no preocular, usually seven (rarely six) upper labials, the tallest of which is six (rarely five); ventrals 177–182, subcaudals 38–42. This snake is very similar to
<taxonomicName id="ACDC4D377E7BFFDAFEB0EB72FE295879" authorityName="Sauvage" authorityYear="1879" box="[380,493,1630,1653]" class="Reptilia" family="Homalopsidae" genus="Atractocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="raffrayi">
<emphasis id="59A8EAA67E7BFFDAFEB0EB72FE295879" box="[380,493,1630,1653]" italics="true" pageId="9" pageNumber="510">B. raffrayi</emphasis>
</taxonomicName>
which has upper labial three in the orbit and the third is the tallest. Specimens of
<taxonomicName id="ACDC4D377E7BFFDAFFBDEBB2FF0F58B9" authorityName="Murphy &amp; Mumpuni &amp; Lang &amp; Gower &amp; Sanders" authorityYear="2012" box="[113,203,1694,1717]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="wallacei">
<emphasis id="59A8EAA67E7BFFDAFFBDEBB2FF0F58B9" box="[113,203,1694,1717]" italics="true" pageId="9" pageNumber="510">wallacei</emphasis>
</taxonomicName>
usually have the fourth upper labial (rarely 3 + 4) in the orbit, and the sixth is the tallest. Our mitochondrial data show that
<taxonomicName id="ACDC4D377E7BFFDAFEDAEBF2FE4258F9" authorityName="Sauvage" authorityYear="1879" box="[278,390,1758,1781]" class="Reptilia" family="Homalopsidae" genus="Atractocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="raffrayi">
<emphasis id="59A8EAA67E7BFFDAFEDAEBF2FE4258F9" box="[278,390,1758,1781]" italics="true" pageId="9" pageNumber="510">B. raffrayi</emphasis>
</taxonomicName>
and
<taxonomicName id="ACDC4D377E7BFFDAFE72EBF2FDF358F9" authorityName="Murphy &amp; Mumpuni &amp; Lang &amp; Gower &amp; Sanders" authorityYear="2012" box="[446,567,1758,1781]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="wallacei">
<emphasis id="59A8EAA67E7BFFDAFE72EBF2FDF358F9" box="[446,567,1758,1781]" italics="true" pageId="9" pageNumber="510">B. wallacei</emphasis>
</taxonomicName>
are separated by a deep molecular divergence at this locus.
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFDAFFBDEA10FAA35B98" pageId="9" pageNumber="510" type="description">
<paragraph id="6B6336B47E7BFFDAFFBDEA10FBAD5F98" blockId="9.[113,752,1851,2100]" lastBlockId="9.[812,1452,188,404]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDEA10FEBA5959" bold="true" box="[113,382,1851,1877]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFFBDEA10FEB35958" bold="true" box="[113,375,1851,1876]" italics="true" pageId="9" pageNumber="510">
Description of
<typeStatus id="B46788167E7BFFDAFED8EA17FEB35958" box="[276,375,1851,1876]" pageId="9" pageNumber="510" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
— An adult female
<quantity id="AC249B517E7BFFDAFD94EA10FD6B5958" box="[600,687,1852,1876]" metricMagnitude="-1" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="510" unit="mm" value="660.0">660 mm</quantity>
SVL,
<quantity id="AC249B517E7BFFDAFFBDEA71FF7A5978" box="[113,190,1885,1909]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="510" unit="mm" value="82.0">82 mm</quantity>
tail. Rostral visible from above, internasals paired, prefrontal fused with loreal and preocular; nasal scales bilobed with posterior lobe dorsal to the first labial; frontal longer than seam of parietals; parietals extend laterally to make contact with upper postocular; two postoculars; one primary temporal scale contacts the lower postocular, a secondary temporal on each side contacts upper labial six; seven upper labials, 2–4 make contact with prefrontal, the fourth enters the orbit, the fifth is the tallest; seven lower labials, the first four make contact with the chin shields; dorsal scales rows are 19 at mid-body; ventrals 180; subcaudals 42. Dorsum grey, anterior body has scale rows one and two with red-brown pigment, replaced by grey posteriorly, ventral surface a uniform red brown.
</paragraph>
<paragraph id="6B6336B47E7BFFDAFCE0EC91FAF25DD8" blockId="9.[812,1453,444,981]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0EC91FC5A5FD9" bold="true" box="[812,926,445,469]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0EC91FC525FD8" bold="true" box="[812,918,445,468]" italics="true" pageId="9" pageNumber="510">
<typeStatus id="B46788167E7BFFDAFCE0EC91FC525FD8" box="[812,918,445,468]" pageId="9" pageNumber="510" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
— The nasals are divided or semi-divided with the nostril in the anterior portion of the scale. Primary temporal scale(s) can be one or two and plate-like. Upper labials number six (or seven), 2-3 (or 2-4) make contact with the fused loreal-prefrontal shield; when the third upper labial enters the orbit the fifth upper labial is the tallest; when the fourth upper labial enters the orbit the sixth is the tallest. One specimen (MZB 3464) has the postoculars fused on the left side. Lower labials seven, the first pair of lower labials make contact on the midline of the chin posterior to the mental. The first four contact the only pair of chin shields. Dorsal scales on the body are smooth and in 19 rows, the number of rows at the 10
<superScript id="9CA99BFC7E7BFFDAFB8CEE17FB885D46" attach="left" box="[1088,1100,827,842]" fontSize="6" pageId="9" pageNumber="510">th</superScript>
ventral may be 19 or 20. Males have 169–182 (N = 7, = 177.14, SD = 4.25) ventrals and 33–42 (N = 7, = 38.5, SD = 3.35) subcaudals. Females have 170–174 (N = 3, = 171.3, SD = 1.88) ventrals and 26–31 (N = 3, = 28.6, SD = 2.04) subcaudals.
</paragraph>
<paragraph id="6B6336B47E7BFFDAFCE0EED1FCA25AB8" blockId="9.[812,1451,1020,1204]" pageId="9" pageNumber="510">
In alcohol, grey-black above with some yellow under the chin; Setiadi and Hamidy (2006) contains two photographs of a live specimen of this species, showing a uniform dark brown-purple snake with a white venter. The specimen in their photo has 7 upper labials, with 3
<superScript id="9CA99BFC7E7BFFDAFB02E957FB1F5A86" attach="left" box="[1230,1243,1147,1162]" fontSize="6" pageId="9" pageNumber="510">rd</superScript>
upper labial in the orbit.
</paragraph>
<paragraph id="6B6336B47E7BFFDAFCE0E9F0FAA35B98" blockId="9.[812,1453,1244,1429]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0E9F0FCA45AF9" bold="true" box="[812,864,1244,1269]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0E9F0FC9C5AF8" bold="true" box="[812,856,1244,1268]" italics="true" pageId="9" pageNumber="510">Size</emphasis>
.
</emphasis>
— Males have SVLs of
<quantity id="AC249B517E7BFFDAFBA8E9F0FB2E5AF8" box="[1124,1258,1244,1268]" metricMagnitude="-1" metricUnit="m" metricValue="5.5649999999999995" metricValueMax="6.6" metricValueMin="4.529999999999999" pageId="9" pageNumber="510" unit="mm" value="556.5" valueMax="660.0" valueMin="453.0">453–660mm</quantity>
(N = 7, = 561.8, SD = 67.7); tails that are
<quantity id="AC249B517E7BFFDAFB8BE9D0FB785B18" box="[1095,1212,1276,1300]" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="8.2" metricValueMin="5.6" pageId="9" pageNumber="510" unit="mm" value="69.0" valueMax="82.0" valueMin="56.0">56–82 mm</quantity>
(n = 7, = 71, SD = 10.03), and t/svl ratios 12.0–13.4 (N = 7, = 12.67, SD = 0.47). Females have SVLs of
<quantity id="AC249B517E7BFFDAFBAEE810FB2A5B58" box="[1122,1262,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="4.279999999999999" metricValueMax="4.62" metricValueMin="3.94" pageId="9" pageNumber="510" unit="mm" value="428.0" valueMax="462.0" valueMin="394.0">394–462 mm</quantity>
(N = 3, = 417.0, SD = 31.8) and tails that are
<quantity id="AC249B517E7BFFDAFB9FE870FB075B78" box="[1107,1219,1372,1396]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.1" metricValueMin="2.9" pageId="9" pageNumber="510" unit="mm" value="40.0" valueMax="51.0" valueMin="29.0">29–51 mm</quantity>
(= 38.0, SD = 9.41); tail/SVL ratios 7–11% (N = 7, = 9%, SD = 0.015).
</paragraph>
</subSubSection>
<subSubSection id="23C6653F7E7BFFD9FCE0E897FD355D7B" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510" type="etymology">
<paragraph id="6B6336B47E7BFFDAFCE0E897FA575BF8" blockId="9.[812,1451,1467,1524]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0E897FC6C5BD9" bold="true" box="[812,936,1467,1493]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0E897FC655BD8" bold="true" box="[812,929,1467,1492]" italics="true" pageId="9" pageNumber="510">Etymology</emphasis>
.
</emphasis>
— Named in honour of Alfred Russell Wallace for his contributions to the natural history of Halmahera.
</paragraph>
<paragraph id="6B6336B47E7BFFDAFCE0EB37FA6F5638" blockId="9.[812,1452,1563,2100]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0EB37FB965839" bold="true" box="[812,1106,1563,1589]" pageId="9" pageNumber="510">
<emphasis id="59A8EAA67E7BFFDAFCE0EB37FC6C5838" bold="true" box="[812,936,1563,1588]" italics="true" pageId="9" pageNumber="510">Unresolved</emphasis>
<taxonomicName id="ACDC4D377E7BFFDAFC61EB31FB895839" box="[941,1101,1565,1589]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">Brachyorrhos</taxonomicName>
.
</emphasis>
— (1) The Aru Islands specimen clearly belongs with this genus despite the speculation that it is
<taxonomicName id="ACDC4D377E7BFFDAFC5BEB72FAEA5879" authority="(Iskandar &amp; Colijn, 2001)" baseAuthorityName="Iskandar &amp; Colijn" baseAuthorityYear="2001" box="[919,1326,1628,1653]" class="Reptilia" family="Homalopsidae" genus="Calamophis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="jobiensis">
<emphasis id="59A8EAA67E7BFFDAFC5BEB72FC3D5879" box="[919,1017,1630,1653]" italics="true" pageId="9" pageNumber="510">jobiensis</emphasis>
(Iskandar &amp; Colijn, 2001)
</taxonomicName>
. It clusters with the Buru island
<taxonomicName id="ACDC4D377E7BFFDAFBD9EB52FB615899" box="[1045,1189,1662,1685]" class="Reptilia" family="Colubridae" genus="Rhabdion" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="gastrotaenius">
<emphasis id="59A8EAA67E7BFFDAFBD9EB52FB615899" box="[1045,1189,1662,1685]" italics="true" pageId="9" pageNumber="510">gastrotaenius</emphasis>
</taxonomicName>
in our PCA and cluster analysis. Its locality data may be in error. (2) The island of Buru appears to have a second species with distinct labial architecture and a long tail. However, this specimen is poorly preserved and more sampling is needed to inform its status. (3) Seram also supports a second species of
<taxonomicName id="ACDC4D377E7BFFDAFADBEA32FA6F5939" box="[1303,1451,1822,1845]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="59A8EAA67E7BFFDAFADBEA32FA6F5939" box="[1303,1451,1822,1845]" italics="true" pageId="9" pageNumber="510">Brachyorrhos</emphasis>
</taxonomicName>
that is exceptionally small (smaller than late term embryos of
<taxonomicName id="ACDC4D377E7BFFDAFC83EA72FC6B5979" authorityName="Setiadi &amp; Hamidy" authorityYear="2006" box="[847,943,1886,1909]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="albus">
<emphasis id="59A8EAA67E7BFFDAFC83EA72FC6B5979" box="[847,943,1886,1909]" italics="true" pageId="9" pageNumber="510">B. albus</emphasis>
</taxonomicName>
) and sexually mature with a distinct loreal scale, preocular, and low ventral count. These specimens will be discussed elsewhere pending further investigation. (4) Morotai also supports a population of
<taxonomicName id="ACDC4D377E7BFFDAFADFEA92FA6F59D9" box="[1299,1451,1982,2005]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="59A8EAA67E7BFFDAFADFEA92FA6F59D9" box="[1299,1451,1982,2005]" italics="true" pageId="9" pageNumber="510">Brachyorrhos</emphasis>
</taxonomicName>
(USNM 159973) that is superficially similar to
<taxonomicName id="ACDC4D377E7BFFDAFAFFEAF2FA6059F9" authorityName="Sauvage" authorityYear="1879" box="[1331,1444,2014,2037]" class="Reptilia" family="Homalopsidae" genus="Atractocephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="raffrayi">
<emphasis id="59A8EAA67E7BFFDAFAFFEAF2FA6059F9" box="[1331,1444,2014,2037]" italics="true" pageId="9" pageNumber="510">B. raffrayi</emphasis>
</taxonomicName>
, with a slightly different labial architecture, and it may require molecular analysis to confirm its relationships. (5)
</paragraph>
<paragraph id="6B6336B47E78FFD9FF44EE2CFCC35D1B" blockId="10.[136,775,768,887]" box="[136,775,767,791]" pageId="10" pageNumber="511">
<taxonomicName id="ACDC4D377E78FFD9FF44EE2CFED85D1B" box="[136,284,768,791]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="511" phylum="Chordata" rank="genus">
<emphasis id="59A8EAA67E78FFD9FF44EE2CFED85D1B" box="[136,284,768,791]" italics="true" pageId="10" pageNumber="511">Brachyorrhos</emphasis>
</taxonomicName>
has also been reported from: Banda, Batjan,
</paragraph>
<paragraph id="6B6336B47E78FFD9FF44EE33FD355D7B" blockId="10.[136,775,768,887]" pageId="10" pageNumber="511">
Haruku, Nusalaut, and Saparua; all of these islands seem likely to support
<taxonomicName id="ACDC4D377E78FFD9FEF7EE6CFE0B5D5B" box="[315,463,832,855]" class="Reptilia" family="Homalopsidae" genus="Brachyorrhos" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="511" phylum="Chordata" rank="genus">
<emphasis id="59A8EAA67E78FFD9FEF7EE6CFE0B5D5B" box="[315,463,832,855]" italics="true" pageId="10" pageNumber="511">Brachyorrhos</emphasis>
</taxonomicName>
populations, but we have not located specimens to confirm their presence or identities.
</paragraph>
</subSubSection>
</treatment>
</document>
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