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<mods:title id="C71AD24C0A40EEC972056F7B97370D96">Contribution to the systematics of the genus Eurythenes S. I. Smith in Scudder, 1882 (Crustacea: Amphipoda: Lysianassoidea: Eurytheneidae)</mods:title>
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<emphasis id="3FF6EAB4FFA1FFA76CE3FA2FFD3621D6" bold="true" box="[151,648,1443,1469]" pageId="4" pageNumber="5">
Superfamily
<taxonomicName id="CA824D25FFA1FFA76D4DFA2FFD3621D6" ID-CoL="7NGQH" authority="Dana, 1849" authorityName="Dana" authorityYear="1849" box="[313,648,1443,1469]" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="superFamily" superFamily="Lysianassoidea">
Lysianassoidea
<bibRefCitation id="69134B57FFA1FFA76D8FFA2FFD3621D6" author="Dana" box="[507,648,1443,1469]" pageId="4" pageNumber="5" refString="Dana, J. D. (1849) Synopsis of the genera of Gammaracea. American Journal of Science and Arts, Series 2, 8, 135 - 140." type="journal article" year="1849">Dana, 1849</bibRefCitation>
</taxonomicName>
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<emphasis id="3FF6EAB4FFA1FFA76CE3FA65FE9A2269" bold="true" box="[151,292,1513,1538]" pageId="4" pageNumber="5">
<typeStatus id="D2398804FFA1FFA76CE3FA65FF6C2269" box="[151,210,1513,1538]" pageId="4" pageNumber="5">Type</typeStatus>
genus.
</emphasis>
<taxonomicName id="CA824D25FFA1FFA76D5FFA65FD9F2269" authority="S.I. Smith" authorityName="S.I. Smith" box="[299,545,1513,1538]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA1FFA76D5FFA65FE17226A" box="[299,425,1513,1537]" italics="true" pageId="4" pageNumber="5">Eurythenes</emphasis>
S.I. Smith
</taxonomicName>
in Scudder, 1882.
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<paragraph id="0D3D36A6FFA1FFA76CB3F981FDF0224D" blockId="4.[151,1437,1513,2006]" box="[199,590,1549,1574]" pageId="4" pageNumber="5">
<emphasis id="3FF6EAB4FFA1FFA76CB3F981FE7B224D" bold="true" box="[199,453,1549,1574]" pageId="4" pageNumber="5">Generic composition.</emphasis>
Monotypic.
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<paragraph id="0D3D36A6FFA1FFA76CB3F9BDFD0A2221" blockId="4.[151,1437,1513,2006]" box="[199,692,1585,1610]" pageId="4" pageNumber="5">
<emphasis id="3FF6EAB4FFA1FFA76CB3F9BDFEE72221" bold="true" box="[199,345,1585,1610]" pageId="4" pageNumber="5">Description.</emphasis>
See
<bibRefCitation id="69134B57FFA1FFA76DFAF9BDFD0E2221" author="Stoddart" box="[398,688,1585,1610]" pageId="4" pageNumber="5" refString="Stoddart, H. E. &amp; Lowry, J. K. (2004) The deep-sea lysianassoid genus Eurythenes (Crustacea, Amphipoda, Eurytheneidae n. fam.). Zoosystema, 26 (3), 425 - 468." type="journal article" year="2004">Stoddart &amp; Lowry (2004)</bibRefCitation>
.
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<subSubSection id="4598652DFFA1FFA66CB3F9D9FBEB26EF" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="0D3D36A6FFA1FFA66CB3F9D9FBEB26EF" blockId="4.[151,1437,1513,2006]" lastBlockId="5.[151,1437,151,645]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="3FF6EAB4FFA1FFA76CB3F9D9FE852205" bold="true" box="[199,315,1621,1646]" pageId="4" pageNumber="5">Remarks.</emphasis>
During the past two decades, Lowry and his collaborators have split the old family Lysianassidaeelevated to the rank of superfamily Lysianassoidea—into a large number of new families. In their studies, a largely phenetic approach was applied, in the sense that each group of taxa sharing distinctive characters was automatically assigned to a separate family or genus, without attempting to unravel the deep interrelationships between taxa. In one case, they even rejected taxa definitions based on phylogenetic analyses (
<bibRefCitation id="69134B57FFA1FFA76874F96AFEC0234A" author="Lowry" pageId="4" pageNumber="5" refString="Lowry, J. K. &amp; Kilgallen, N. M. (2014) A generic review of the lysianassoid family Uristidae and descriptions of new taxa from Australian waters (Crustacea, Amphipoda, Uristidae). Zootaxa, 3867 (1), 1 - 92. http: // dx. doi. org / 10.11646 / zootaxa. 3867.1.1" type="journal article" year="2014">
Lowry &amp; Kilgallen 2014: discussion on
<taxonomicName id="CA824D25FFA1FFA76CCAF885FEC0234A" box="[190,382,1801,1825]" class="Malacostraca" family="Lysianassidae" genus="Pseudorchomene" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA1FFA76CCAF885FEC0234A" box="[190,382,1801,1825]" italics="true" pageId="4" pageNumber="5">Pseudorchomene</emphasis>
</taxonomicName>
</bibRefCitation>
). In the course of this division process,
<bibRefCitation id="69134B57FFA1FFA76F16F885FB312349" author="Stoddart" box="[866,1167,1801,1826]" pageId="4" pageNumber="5" refString="Stoddart, H. E. &amp; Lowry, J. K. (2004) The deep-sea lysianassoid genus Eurythenes (Crustacea, Amphipoda, Eurytheneidae n. fam.). Zoosystema, 26 (3), 425 - 468." type="journal article" year="2004">Stoddart &amp; Lowry (2004)</bibRefCitation>
erected the monotypic family
<taxonomicName id="CA824D25FFA1FFA76C9DF8A1FE35232D" box="[233,395,1837,1862]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Eurytheneidae</taxonomicName>
for the genus
<taxonomicName id="CA824D25FFA1FFA76E45F8A1FD11232E" box="[561,687,1837,1861]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA1FFA76E45F8A1FD11232E" box="[561,687,1837,1861]" italics="true" pageId="4" pageNumber="5">Eurythenes</emphasis>
</taxonomicName>
. Recent works such as
<bibRefCitation id="69134B57FFA1FFA76FCAF8A2FB73232D" author="Havermans" box="[958,1229,1837,1862]" pageId="4" pageNumber="5" refString="Havermans, C., Nagy, Z. T., Sonet, G., De Broyer, C. &amp; Martin, P. (2010) Incongruence between molecular phylogeny and morphological classification in amphipod crustaceans: A case study of Antarctic lysianassoids. Molecular Phylogenetics and Evolution, 55, 202 - 209. http: // dx. doi. org / 10.1016 / j. ympev. 2009.10.025" type="journal article" year="2010">
Havermans
<emphasis id="3FF6EAB4FFA1FFA76833F8A2FB3F232E" box="[1095,1153,1837,1861]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(2010
</bibRefCitation>
,
<bibRefCitation id="69134B57FFA1FFA768AEF8A2FAAB232D" author="Havermans" box="[1242,1301,1838,1862]" pageId="4" pageNumber="5" refString="Havermans, C., Nagy, Z. T., Sonet, G., De Broyer, C. &amp; Martin, P. (2011) DNA barcoding reveals new insights into the diversity of Antarctic species of Orchomene sensu lato (Crustacea: Amphipoda: Lysianassoidea). Deep - Sea Research Part II: Topical Studies in Oceanography, 58 (1 - 2), 230 - 241. http: // dx. doi. org / doi: 10.1016 / j. dsr 2.2010.09.028" type="journal article" year="2011">2011</bibRefCitation>
),
<bibRefCitation id="69134B57FFA1FFA7695EF8A1FE602301" pageId="4" pageNumber="5" refString="d'Udekem d'Acoz, C. &amp; Havermans, C. (2012) Two new Pseudorchomene species from the Southern Ocean, with phylogenetic remarks on the genus and related species (Crustacea: Amphipoda: Lysianassoidea: Lysianassidae: Tryphosinae). Zootaxa, 3310, 1 - 50." type="journal article">d'Udekem d'Acoz &amp; Havermans (2012)</bibRefCitation>
and
<bibRefCitation id="69134B57FFA1FFA76E62F8DDFCB12301" box="[534,783,1873,1898]" pageId="4" pageNumber="5" refString="Corrigan, L. J., Horton, T., Fotherby, H., White, T. A. &amp; Hoelzel, A. R. (2014) Adaptative evolution of the deep-sea amphipods from the superfamily Lysiassanoidea [sic] in the North Atlantic. Evolutionary Biology, 41 (1), 154 - 165. http: // dx. doi. org / 10.1007 / s 11692 - 013 - 9255 - 2" type="journal article">
Corrigan
<emphasis id="3FF6EAB4FFA1FFA76EF6F8DEFD0B2302" box="[642,693,1873,1897]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. (2014)
</bibRefCitation>
have shown that an important part of traditional or semiphenetic classifications of lysianassoid amphipods are not supported by molecular phylogenies. Therefore, the validity (and the composition) of the family
<taxonomicName id="CA824D25FFA1FFA76EEEF815FC8223D9" box="[666,828,1945,1970]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Eurytheneidae</taxonomicName>
remains an open question, as indeed for many other lysianassoid families. Future molecular phylogenies involving a larger number of lysianassoid taxa and a higher number of genetic markers will probably help to solve such issues, but for the time being, with limited data at hand, the family
<taxonomicName id="CA824D25FFA0FFA66D62FF30FE0624BE" box="[278,440,188,213]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Eurytheneidae</taxonomicName>
is accepted. The question of its delimitation is another issue to be solved. Should the
<taxonomicName id="CA824D25FFA0FFA66CE3FF53FE872493" box="[151,313,223,248]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Eurytheneidae</taxonomicName>
remain restricted to
<taxonomicName id="CA824D25FFA0FFA66E5CFF6CFD182493" box="[552,678,224,248]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66E5CFF6CFD182493" box="[552,678,224,248]" italics="true" pageId="5" pageNumber="6">Eurythenes</emphasis>
</taxonomicName>
as proposed by
<bibRefCitation id="69134B57FFA0FFA66F10FF53FB362493" author="Stoddart" box="[868,1160,223,248]" pageId="5" pageNumber="6" refString="Stoddart, H. E. &amp; Lowry, J. K. (2004) The deep-sea lysianassoid genus Eurythenes (Crustacea, Amphipoda, Eurytheneidae n. fam.). Zoosystema, 26 (3), 425 - 468." type="journal article" year="2004">Stoddart &amp; Lowry (2004)</bibRefCitation>
or should the family be expanded to include other genera? In a recent multigene phylogenetic study including 15 scavenger lysianassoid taxa,
<bibRefCitation id="69134B57FFA0FFA66CADFEABFE63252B" box="[217,477,295,320]" pageId="5" pageNumber="6" refString="Corrigan, L. J., Horton, T., Fotherby, H., White, T. A. &amp; Hoelzel, A. R. (2014) Adaptative evolution of the deep-sea amphipods from the superfamily Lysiassanoidea [sic] in the North Atlantic. Evolutionary Biology, 41 (1), 154 - 165. http: // dx. doi. org / 10.1007 / s 11692 - 013 - 9255 - 2" type="journal article">
Corrigan
<emphasis id="3FF6EAB4FFA0FFA66D3DFEA5FE38252B" box="[329,390,296,320]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2014)
</bibRefCitation>
have shown that
<taxonomicName id="CA824D25FFA0FFA66EC3FEA4FC8B252B" box="[695,821,296,320]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66EC3FEA4FC8B252B" box="[695,821,296,320]" italics="true" pageId="5" pageNumber="6">Eurythenes</emphasis>
</taxonomicName>
forms a clade together with the genera
<taxonomicName id="CA824D25FFA0FFA66968FEA4FA2B252B" box="[1308,1429,296,320]" class="Malacostraca" family="Lysianassidae" genus="Cyclocaris" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66968FEA4FA2B252B" box="[1308,1429,296,320]" italics="true" pageId="5" pageNumber="6">Cyclocaris</emphasis>
</taxonomicName>
,
<taxonomicName id="CA824D25FFA0FFA66CE3FEC0FEAB250F" box="[151,277,332,356]" class="Malacostraca" family="Lysianassidae" genus="Paralicella" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66CE3FEC0FEAB250F" box="[151,277,332,356]" italics="true" pageId="5" pageNumber="6">Paralicella</emphasis>
</taxonomicName>
and
<taxonomicName id="CA824D25FFA0FFA66D23FEC0FE77250F" box="[343,457,332,356]" class="Malacostraca" family="Uristidae" genus="Stephonyx" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66D23FEC0FE77250F" box="[343,457,332,356]" italics="true" pageId="5" pageNumber="6">Stephonyx</emphasis>
</taxonomicName>
. While these genera exhibit significant morphological differences, they also share similarities, such as a reduction of coxa 1, which might be synapomorphies. To visualise the differences between these genera and
<taxonomicName id="CA824D25FFA0FFA66D28FE18FE6425C7" box="[348,474,404,428]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66D28FE18FE6425C7" box="[348,474,404,428]" italics="true" pageId="5" pageNumber="6">Eurythenes</emphasis>
</taxonomicName>
, see for instance the illustrations of
<taxonomicName id="CA824D25FFA0FFA66F0DFE18FC4D25C7" box="[889,1011,404,428]" class="Malacostraca" family="Lysianassidae" genus="Cyclocaris" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66F0DFE18FC4D25C7" box="[889,1011,404,428]" italics="true" pageId="5" pageNumber="6">Cyclocaris</emphasis>
</taxonomicName>
given by
<bibRefCitation id="69134B57FFA0FFA66811FE18FA9825C7" author="Sars" box="[1125,1318,404,428]" pageId="5" pageNumber="6" refString="Sars, G. O. (1900) Crustacea. In: Nansen, F. (Ed.), The Norwegian North Pole Expedition 1893 - 1896. Scientific Results, 1 (5), pp. 1 - 141, pls. 1 - 36." type="journal article" year="1900">G.O. Sars (1900)</bibRefCitation>
,
<bibRefCitation id="69134B57FFA0FFA66947FE19FEF625BB" author="Lowry" pageId="5" pageNumber="6" refString="Lowry, J. K. &amp; Stoddart, H. E. (2011) The new deep-sea families Cebocaridae fam. nov., Cyclocaridae fam. nov. and Thoriellidae fam. nov. (Crustacea: Amphipoda: Lysianassoidea). Zootaxa, 27, 53 - 68." type="journal article" year="2011">Lowry &amp; Stoddart (2011)</bibRefCitation>
and
<bibRefCitation id="69134B57FFA0FFA66DF7FE34FD0925BB" author="Horton" box="[387,695,439,464]" pageId="5" pageNumber="6" refString="Horton, T. &amp; Thurston, M. H. (2014) A revision of the bathyal and abyssal necrophage genus Cyclocaris Stebbing, 1888 (Crustacea: Amphipoda: Cyclocarididae) with the addition of two new species from the Atlantic Ocean. Zootaxa, 3796 (3), 507 - 527. http: // dx. doi. org / 10.11646 / zootaxa. 3796.3.6" type="journal article" year="2014">Horton &amp; Thurston (2014)</bibRefCitation>
, those of
<taxonomicName id="CA824D25FFA0FFA66F5EFE34FC1625BB" box="[810,936,440,464]" class="Malacostraca" family="Lysianassidae" genus="Paralicella" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66F5EFE34FC1625BB" box="[810,936,440,464]" italics="true" pageId="5" pageNumber="6">Paralicella</emphasis>
</taxonomicName>
by Barnard &amp; Ingram (1990), and those of
<taxonomicName id="CA824D25FFA0FFA66CE3FE50FEB7259F" box="[151,265,476,500]" class="Malacostraca" family="Uristidae" genus="Stephonyx" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66CE3FE50FEB7259F" box="[151,265,476,500]" italics="true" pageId="5" pageNumber="6">Stephonyx</emphasis>
</taxonomicName>
by
<bibRefCitation id="69134B57FFA0FFA66D4AFE50FD37259F" author="Diffenthal" box="[318,649,476,501]" pageId="5" pageNumber="6" refString="Diffenthal, M. &amp; Horton, T. (2007) Stephonyx arabiensis (Crustacea: Amphipoda: Lysianassoidea: Uristidae), a new deepwater scavenger species from the Indian Ocean, with a key to the genus Stephonyx. Zootaxa, 1665, 31 - 41." type="journal article" year="2007">Diffenthal &amp; Horton (2007)</bibRefCitation>
,
<bibRefCitation id="69134B57FFA0FFA66EEEFE50FC1C259F" box="[666,930,476,501]" pageId="5" pageNumber="6" refString="Narahara, Y., Tomikawa, K. &amp; Torigoe, K. (2012) Four species of the genus Stephonyx (Crustacea: Amphipoda: Uristidae) from Japan, with description of a new species. Journal of Natural History, 46 (23 - 24), 1477 - 1507. http: // dx. doi. org / 10.1080 / 00222933.2012.675598" type="journal article">
Narahara
<emphasis id="3FF6EAB4FFA0FFA66F79FE51FCF5259F" box="[781,843,476,500]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2012)
</bibRefCitation>
and
<bibRefCitation id="69134B57FFA0FFA66F97FE51FA9E259F" author="Lowry" box="[995,1312,476,501]" pageId="5" pageNumber="6" refString="Lowry, J. K. &amp; Kilgallen, N. M. (2014) A generic review of the lysianassoid family Uristidae and descriptions of new taxa from Australian waters (Crustacea, Amphipoda, Uristidae). Zootaxa, 3867 (1), 1 - 92. http: // dx. doi. org / 10.11646 / zootaxa. 3867.1.1" type="journal article" year="2014">Lowry &amp; Kilgallen (2014)</bibRefCitation>
. If further molecular studies based on a larger number of taxa confirm the existence of a cluster of genera surrounding
<taxonomicName id="CA824D25FFA0FFA66CE3FDA8FEAB2657" box="[151,277,548,572]" class="Malacostraca" family="Lysianassidae" genus="Eurythenes" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66CE3FDA8FEAB2657" box="[151,277,548,572]" italics="true" pageId="5" pageNumber="6">Eurythenes</emphasis>
</taxonomicName>
, it would be logical to expand the definition of
<taxonomicName id="CA824D25FFA0FFA66F41FDA8FC692656" box="[821,983,548,573]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Eurytheneidae</taxonomicName>
to include these additional genera. This would imply the relegation of the monotypic family Cyclocarididae
<bibRefCitation id="69134B57FFA0FFA66FA1FDC4FB43260B" author="Lowry" box="[981,1277,583,608]" pageId="5" pageNumber="6" refString="Lowry, J. K. &amp; Stoddart, H. E. (2011) The new deep-sea families Cebocaridae fam. nov., Cyclocaridae fam. nov. and Thoriellidae fam. nov. (Crustacea: Amphipoda: Lysianassoidea). Zootaxa, 27, 53 - 68." type="journal article" year="2011">Lowry &amp; Stoddart, 2011</bibRefCitation>
(
<typeStatus id="D2398804FFA0FFA66967FDC4FAFB260B" box="[1299,1349,584,608]" pageId="5" pageNumber="6">type</typeStatus>
genus:
<taxonomicName id="CA824D25FFA0FFA66CE3FDE0FEAE26EF" box="[151,272,620,644]" class="Malacostraca" family="Lysianassidae" genus="Cyclocaris" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="3FF6EAB4FFA0FFA66CE3FDE0FEAE26EF" box="[151,272,620,644]" italics="true" pageId="5" pageNumber="6">Cyclocaris</emphasis>
</taxonomicName>
) to the rank of junior synonym of
<taxonomicName id="CA824D25FFA0FFA66EE7FDE0FBEF26EF" authority="Stoddart &amp; Lowry, 2004" authorityName="Stoddart &amp; Lowry" authorityYear="2004" box="[659,1105,620,645]" class="Malacostraca" family="Eurytheneidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">
Eurytheneidae
<bibRefCitation id="69134B57FFA0FFA66F48FDE0FBEF26EF" author="Stoddart" box="[828,1105,620,645]" pageId="5" pageNumber="6" refString="Stoddart, H. E. &amp; Lowry, J. K. (2004) The deep-sea lysianassoid genus Eurythenes (Crustacea, Amphipoda, Eurytheneidae n. fam.). Zoosystema, 26 (3), 425 - 468." type="journal article" year="2004">Stoddart &amp; Lowry, 2004</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>