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<document id="B9207AF685E05E918E2FAC93CDEF905E" ID-CLB-Dataset="20094" ID-DOI="10.11646/zootaxa.3683.4.1" ID-GBIF-Dataset="4f6d65aa-8c5b-48ca-9d27-ee17d022a1b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="5641557" ID-ZooBank="2A3C7450-C8D8-479C-A295-C2BB49A151EF" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1629866806485" checkinUser="felipe" docAuthor="FISCHER, G. &amp; FISHER, B. L." docDate="2013" docId="A93CCE11FFD1463BFF31E07BFD073730" docLanguage="en" docName="zt03683p356.pdf" docOrigin="Zootaxa 3683 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.3683.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pheidole jonas Forel" docType="treatment" docVersion="15" lastPageNumber="328" masterDocId="5505B669FFC94620FFA6E56CFFB7346C" masterDocTitle="A revision of Pheidole Westwood (Hymenoptera: Formicidae) in the islands of the Southwest Indian Ocean and designation of a neotype for the invasive Pheidole megacephala" masterLastPageNumber="356" masterPageNumber="301" pageNumber="325" updateTime="1699145335802" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="764438177317E9BCE38F382A71FB5A6B">A revision of Pheidole Westwood (Hymenoptera: Formicidae) in the islands of the Southwest Indian Ocean and designation of a neotype for the invasive Pheidole megacephala</mods:title>
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<mods:namePart id="C47EE381D83C2A335CA10B2704FA7DE0">FISCHER, G.</mods:namePart>
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<mods:namePart id="972DEF4F1E946E9ED04610D3261D82C8">FISHER, B. L.</mods:namePart>
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<treatment id="A93CCE11FFD1463BFF31E07BFD073730" ID-DOI="http://doi.org/10.5281/zenodo.5462842" ID-GBIF-Taxon="194286122" ID-Zenodo-Dep="5462842" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730" httpUri="http://treatment.plazi.org/id/A93CCE11FFD1463BFF31E07BFD073730" lastPageId="27" lastPageNumber="328" pageId="24" pageNumber="325">
<subSubSection id="698F2C8CFFD14638FF31E07BFE25315D" box="[151,402,1303,1329]" pageId="24" pageNumber="325" type="nomenclature">
<paragraph id="212A7F07FFD14638FF31E07BFE25315D" blockId="24.[151,402,1303,1362]" box="[151,402,1303,1329]" pageId="24" pageNumber="325">
<heading id="7A62C86BFFD14638FF31E07BFE25315D" bold="true" box="[151,402,1303,1329]" fontSize="11" level="1" pageId="24" pageNumber="325" reason="1">
<taxonomicName id="E6950484FFD14638FF31E07BFE25315D" ID-CoL="6V95C" ID-ENA="615091" authority="Forel" authorityName="Forel" box="[151,402,1303,1329]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD14638FF31E07BFE25315D" bold="true" box="[151,402,1303,1329]" pageId="24" pageNumber="325">
<emphasis id="13E1A315FFD14638FF31E07BFEFE315D" bold="true" box="[151,329,1303,1329]" italics="true" pageId="24" pageNumber="325">Pheidole jonas</emphasis>
Forel
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD14638FF31E056FE82313E" box="[151,309,1338,1362]" pageId="24" pageNumber="325" type="description">
<paragraph id="212A7F07FFD14638FF31E056FE82313E" blockId="24.[151,402,1303,1362]" box="[151,309,1338,1362]" pageId="24" pageNumber="325">
(
<figureCitation id="B9AE6382FFD14638FF39E056FE9A313E" box="[159,301,1338,1362]" captionStart="FIGURE 18" captionStartId="25.[151,250,1739,1762]" captionTargetBox="[151,1436,311,1708]" captionTargetId="figure-162@25.[151,1436,311,1708]" captionTargetPageId="25" captionText="FIGURE 18 Pheidole jonas Forel, full-face view (A), profile (B), and dorsal view (C) of minor worker (CASENT006581) and full-face view (D), profile (E), and dorsal view (F) of major worker (CASENT0060250)." figureDoi="http://doi.org/10.5281/zenodo.5641593" httpUri="https://zenodo.org/record/5641593/files/figure.png" pageId="24" pageNumber="325">Figs. 18AF</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD14638FF31E0EEFE4631F4" box="[151,497,1410,1432]" pageId="24" pageNumber="325" type="reference_group">
<paragraph id="212A7F07FFD14638FF31E0EEFE4631F4" blockId="24.[151,1436,1410,1494]" box="[151,497,1410,1432]" pageId="24" pageNumber="325">
<taxonomicName id="E6950484FFD14638FF31E0EEFE5A31F4" ID-CoL="6V95C" authority="Forel, 1907 c: 82" authorityName="Forel" authorityPageNumber="82" authorityYear="1907" box="[151,493,1410,1432]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD14638FF31E0EEFE8531F4" box="[151,306,1410,1432]" italics="true" pageId="24" pageNumber="325">Pheidole jonas</emphasis>
<bibRefCitation id="450402F6FFD14638FE9AE0EEFE5A31F4" author="Forel" box="[316,493,1410,1432]" firstAuthor="Forel" page="82" pageId="24" pageNumber="325" pagination="75 - 92" refId="ref48093" refString="Forel, A. (1907 c). Ameisen von Madagaskar, den Comoren und Ostafrika. Wissenschaftliche Ergebnisse. Reise in Ostafrika, 2, 75 - 92." type="journal article" year="1907" yearSuffix="c">Forel, 1907c: 82</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD14638FE5CE0EFFB9631BA" pageId="24" pageNumber="325" type="materials_examined">
<paragraph id="212A7F07FFD14638FE5CE0EFFB9631BA" blockId="24.[151,1436,1410,1494]" pageId="24" pageNumber="325">
<materialsCitation id="91FD755AFFD14638FE5CE0EFFC9E31DB" ID-GBIF-Occurrence="3736261531" collectionCode="MHNG" collectorName="A. Voeltzkow" country="Comoros" location="Grande Comore" pageId="24" pageNumber="325" specimenCode="CASENT0101758" specimenCount="1" specimenCount-worker="1" stateProvince="Grande Comore" typeStatus="lectotype">
<typeStatus id="FE2EC1A5FFD14638FE5CE0EFFDE831F4" box="[506,607,1411,1432]" pageId="24" pageNumber="325" type="lectotype">Lectotype</typeStatus>
(1 major
<specimenCount id="3793B48EFFD14638FD69E0EEFCAF31F4" box="[719,792,1410,1432]" pageId="24" pageNumber="325" type="worker">worker</specimenCount>
,
<specimenCode id="7133D77CFFD14638FC80E0EEFC5531F4" box="[806,994,1410,1432]" pageId="24" pageNumber="325">CASENT0101758</specimenCode>
) [designated here]:
<collectingCountry id="59823F97FFD14638FB1EE0EEFA2B31F4" box="[1208,1436,1410,1432]" name="Comoros" pageId="24" pageNumber="325">COMORO ISLANDS</collectingCountry>
(
<collectingRegion id="E351B1E5FFD14638FF6BE0CDFEC931DB" box="[205,382,1441,1463]" country="Comoros" name="Grande Comore" pageId="24" pageNumber="325">Grande Comore</collectingRegion>
) (
<emphasis id="13E1A315FFD14638FE38E0CDFD9E31DB" box="[414,553,1441,1463]" italics="true" pageId="24" pageNumber="325">
<collectorName id="8C601AD1FFD14638FE38E0CDFD9E31DB" box="[414,553,1441,1463]" pageId="24" pageNumber="325">A. Voeltzkow</collectorName>
</emphasis>
) (
<collectionCode id="4784E7C2FFD14638FDECE0CDFD2D31DB" box="[586,666,1441,1463]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="24" pageNumber="325">MHNG</collectionCode>
) [examined]
</materialsCitation>
;
<materialsCitation id="91FD755AFFD14638FC9BE0CDFB9631BA" ID-GBIF-Occurrence="3736261584" collectionCode="CASENT" pageId="24" pageNumber="325" specimenCode="CASENT0101758, CASENT0101689" specimenCount="2" specimenCount-worker="2" typeStatus="paralectotype">
<typeStatus id="FE2EC1A5FFD14638FC9BE0CDFC7831DB" box="[829,975,1441,1463]" pageId="24" pageNumber="325" type="paralectotype">paralectotypes</typeStatus>
(1 major
<specimenCount id="3793B48EFFD14638FBF4E0CDFB1331DB" box="[1106,1188,1441,1463]" pageId="24" pageNumber="325" type="worker">workers</specimenCount>
&amp; 3 minor
<specimenCount id="3793B48EFFD14638FAE2E0CDFA2F31DB" box="[1348,1432,1441,1463]" pageId="24" pageNumber="325" type="worker">workers</specimenCount>
,
<specimenCode id="7133D77CFFD14638FF61E0ACFE3231BA" box="[199,389,1472,1494]" pageId="24" pageNumber="325">CASENT0101758</specimenCode>
,
<specimenCode id="7133D77CFFD14638FE28E0ACFDFB31BA" box="[398,588,1472,1494]" pageId="24" pageNumber="325">CASENT0101689</specimenCode>
) same data as lectotype (MHNG) [examined].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD14638FF31E36EFF663332" pageId="24" pageNumber="325" type="diagnosis">
<paragraph id="212A7F07FFD14638FF31E36EFF663332" blockId="24.[151,1436,1538,2030]" pageId="24" pageNumber="325">
<emphasis id="13E1A315FFD14638FF31E36EFEA43277" bold="true" box="[151,275,1538,1563]" pageId="24" pageNumber="325">Diagnosis:</emphasis>
Small species (WL major
<quantity id="E66DD2E2FFD14638FDE0E36EFD5B3276" box="[582,748,1538,1562]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" metricValueMax="9.5" metricValueMin="7.9" pageId="24" pageNumber="325" unit="mm" value="0.87" valueMax="0.95" valueMin="0.79">0.790.95 mm</quantity>
, WL minor
<quantity id="E66DD2E2FFD14638FCDFE36EFB963276" box="[889,1057,1538,1562]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" metricValueMax="6.8999999999999995" metricValueMin="5.3" pageId="24" pageNumber="325" unit="mm" value="0.61" valueMax="0.69" valueMin="0.53">0.530.69 mm</quantity>
), minor workers with moderately short scapes and legs (SI 102122, FI 107122), major workers with moderately short legs (FI 6175), in both subcastes standing hairs comparatively sparse on mesosoma.
<emphasis id="13E1A315FFD14638FCEAE326FC2F320F" bold="true" box="[844,920,1610,1635]" pageId="24" pageNumber="325">Major</emphasis>
head posterior half irregularly rugoreticulate, submedian hypostomal teeth large, median process absent, promesonotal process short with shallowly concave groove and almost vertical posterior declivity, spines short-spinose to almost triangular (PSLI 1014), postpetiole higher than long (LPpI 8395), without ventral process, but with prominent lateral process and thus on average 2.5 times as wide as petiole.
<emphasis id="13E1A315FFD14638FE6BE3B6FDAE329F" bold="true" box="[461,537,1754,1779]" pageId="24" pageNumber="325">Minor</emphasis>
workers head subrectangular and punctate, clypeus smooth to superficially punctate, median and lateral carinae absent, mesosoma regularly punctate, often with weak irregular rugulae on pronotum, spines short-subtriangular (PSLI 1217), postpetiole on average 1.4 times longer than high (LPpI 122 163).
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD1463AFF61E206FACA318B" lastPageId="26" lastPageNumber="327" pageId="24" pageNumber="325" type="description">
<paragraph id="212A7F07FFD14639FF61E206FEDD349B" blockId="24.[151,1436,1538,2030]" lastBlockId="25.[151,1436,151,247]" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
<emphasis id="13E1A315FFD14638FF61E206FD8533EF" bold="true" box="[199,562,1898,1923]" pageId="24" pageNumber="325">Description of major workers:</emphasis>
Measurements (n=16): HW 0.961.18 (1.06), HL 0.981.18 (1.07), SL 0.47 0.60 (0.56), MDL 0.450.59 (0.52), EL 0.110.16 (0.13), WL 0.790.95 (0.87), PNH 0.390.47 (0.43), PNW 0.45 0.55 (0.50), MNH 0.590.73 (0.65), PDH 0.270.35 (0.30), PTL 0.280.42 (0.35), PPL 0.180.22 (0.20), PTH 0.170.22 (0.21), PPH 0.160.21 (0.19), PTW 0.130.17 (0.15), PPW 0.320.46 (0.38), PSL 0.110.14 (0.12), MFL 0.650.81 (0.74), MTL 0.450.60 (0.54), CI 95102 (99), SI 4756 (52), MDI 4551 (49), EI 1114 (12), FI 6175 (70), PSLI 1014 (12), LPpI 90126 (109), DPpI 152242 (188), PpWI 218300 (256), PpLI 5064 (57), PpHI 83106 (90).
</paragraph>
<caption id="75EA2F8FFFD04639FF31E3A7FBAC336C" ID-DOI="http://doi.org/10.5281/zenodo.5641593" ID-Zenodo-Dep="5641593" httpUri="https://zenodo.org/record/5641593/files/figure.png" pageId="25" pageNumber="326" startId="25.[151,250,1739,1762]" targetBox="[151,1436,311,1708]" targetPageId="25">
<paragraph id="212A7F07FFD04639FF31E3A7FBAC336C" blockId="25.[151,1436,1739,1792]" pageId="25" pageNumber="326">
<emphasis id="13E1A315FFD04639FF31E3A7FEAF328E" bold="true" box="[151,280,1739,1762]" pageId="25" pageNumber="326">FIGURE 18</emphasis>
<taxonomicName id="E6950484FFD04639FEB9E3A7FE43328D" authority="Forel" authorityName="Forel" box="[287,500,1739,1761]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD04639FEB9E3A7FE01328D" box="[287,438,1739,1761]" italics="true" pageId="25" pageNumber="326">Pheidole jonas</emphasis>
Forel
</taxonomicName>
, full-face view (A), profile (B), and dorsal view (C) of minor worker (CASENT006581) and full-face view (D), profile (E), and dorsal view (F) of major worker (CASENT0060250).
</paragraph>
</caption>
<paragraph id="212A7F07FFD0463AFF61E241FADD3683" blockId="25.[151,1437,1837,2005]" lastBlockId="26.[151,1436,151,2015]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">Head about as wide as long (CI 95102), sides weakly convex, posterior emargination comparatively shallow, weakly V-shaped. Mandibles short (MDI 4551), smooth, and shiny, with very few short rugulae laterobasally. Clypeus smooth, posteromedially often with weakly developed, sometimes anteriorly converging carinae, median carina absent, 2 pairs of lateral carinae present. Frontal carinae very short or inconspicuous, antennal scrobe absent. Face weakly punctate, anteriorly with longitudinal rugae, sides of head irregularly longitudinally rugose to weakly rugoreticulate, head posterior 1/3 irregularly rugoreticulate, ventrally weakly to superficially punctate. Submedian hypostomal teeth large and very conspicuous in profile view, median process absent. Scapes relatively short (SI 4756) with decumbent pilosity and few longer erect to suberect standing hairs on outer edge. Promesonotal dome high, strongly convex, rounded to angulate anteriorly, promesonotal process in profile short with shallow, concave groove and situated low at mesonotum, posteriorly bluntly angulate with vertical declivity toward metanotal groove, in dorsal view with humeri not produced laterally. Dorsal surface of promesonotum punctate to very weakly punctate, additionally weakly rugoreticulate or with irregular transverse rugulae, rest of mesosoma punctate, often with slightly reduced sculpture basally on katepisternum and propodeum. Metanotal groove not impressed, cross-ribs mostly short, weakly developed, with superficially punctate interspaces. Spines short-spinose to subtriangular (PSLI 1014), distinctly shorter than distance between their bases. Metafemur moderately short (FI 6175), metatibia pilosity decumbent, on outer edge also with few longer, suberect hairs. Petiole in dorsal view anteriorly wider with prominent lateral process, postpetiole in profile usually longer than high (LPpI mean 109), ventral process absent, in dorsal view with short to relatively long lateral processes, between two and three times wider than petiole (PpWI 218300). Petiole and postpetiole punctate everywhere, postpetiolar dorsum sometimes superficially sculptured. First gastral tergite anteriorly weakly punctate to micropunctate. Standing hairs on mesosoma and metasoma acute, long, and slender, only moderately abundant, shorter on head, and with interspersed shorter decumbent to subdecumbent pilosity. Whole body orange with yellow to dark brown legs.</paragraph>
<paragraph id="212A7F07FFD3463AFF61E797FE2F3780" blockId="26.[151,1436,151,2015]" pageId="26" pageNumber="327">
<emphasis id="13E1A315FFD3463AFF61E797FD843778" bold="true" box="[199,563,763,788]" pageId="26" pageNumber="327">Description of minor workers:</emphasis>
Measurements (n=16): HW 0.370.50 (0.46), HL 0.450.57 (0.51), SL 0.44 0.55 (0.50), MDL 0.260.33 (0.30), EL 0.080.11 (0.09), WL 0.530.69 (0.60), PNH 0.210.28 (0.23), PNW 0.27 0.34 (0.30), MNH 0.340.48 (0.41), PDH 0.170.22 (0.20), PTL 0.180.26 (0.22), PPL 0.110.16 (0.13), PTH 0.110.13 (0.12), PPH 0.080.11 (0.09), PTW 0.070.10 (0.08), PPW 0.110.15 (0.13), PSL 0.060.09 (0.07), MFL 0.450.61 (0.53), MTL 0.340.49 (0.40), CI 8394 (90), SI 102122 (109), MDI 6073 (66), EI 1923 (21), FI 107122 (116), PSLI 1217 (15), LPpI 122163 (144), DPpI 88109 (97), PpWI 133175 (155), PpLI 5274 (61), PpHI 7383 (77).
</paragraph>
<paragraph id="212A7F07FFD3463AFF61E69BFACA318B" blockId="26.[151,1436,151,2015]" pageId="26" pageNumber="327">
Head subrectangular, slightly longer than wide (CI 8394), sides convex, posterior head margin weakly concave medially, occipital carina invisible in full-face view. Mandibles moderately short (MDI 6073), smooth and shiny. Clypeus smooth to superficially punctate, carinae absent. Face completely punctate, malar carinae faintly developed toward posterior eye level. Scapes short (SI 102122), when laid back surpassing posterior head margin by length of 10
<superScript id="D6E0D24FFFD3463AFE02E1EBFE0730F9" attach="left" box="[420,432,1159,1173]" fontSize="6" pageId="26" pageNumber="327">th</superScript>
funicular segment, pilosity mostly subdecumbent to suberect. Promesonotal outline in profile convex, weakly elongated, often subangulate anteriorly and posteriorly, posterior process absent to inconspicuous. Metanotal groove barely to shallowly impressed, cross-ribs conspicuous to superficial with smooth interspaces. Propodeum in profile about slightly higher than long, highest point immediately after metanotal groove, propodeal spines short subtriangular, significantly shorter than distance between their bases (PSLI 1217). Mesosoma punctate to weakly punctate. Metafemur short (FI 107122), metatibia with decumbent to subdecumbent pilosity. Postpetiole significantly longer than high (LPpI 122163), about as long as wide (DpPI 88 109), lower than petiole (PpHI 7383), and without ventral process. Both waist segments superficially to weakly punctate. Gaster smooth and shiny. Standing hairs moderately short, comparatively scarce on mesosoma and metasoma, stiff and acute, shorter pilosity more abundant, decumbent to subdecumbent. Color orange or brown.
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD3463BFF61E09EFEEF349B" lastPageId="27" lastPageNumber="328" pageId="26" pageNumber="327" type="discussion">
<paragraph id="212A7F07FFD3463BFF61E09EFEEF349B" blockId="26.[151,1436,151,2015]" lastBlockId="27.[151,1436,151,860]" lastPageId="27" lastPageNumber="328" pageId="26" pageNumber="327">
<emphasis id="13E1A315FFD3463AFF61E09EFEF93267" bold="true" box="[199,334,1522,1547]" pageId="26" pageNumber="327">Discussion:</emphasis>
<taxonomicName id="E6950484FFD3463AFEFEE09FFDB63266" box="[344,513,1523,1547]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD3463AFEFEE09FFDB63266" box="[344,513,1523,1547]" italics="true" pageId="26" pageNumber="327">Pheidole jonas</emphasis>
</taxonomicName>
is known only from the
<collectingCountry id="59823F97FFD3463AFC80E09EFC3A3266" box="[806,909,1522,1546]" name="Comoros" pageId="26" pageNumber="327">Comoros</collectingCountry>
Islands and
<collectingCountry id="59823F97FFD3463AFB84E09FFB343267" box="[1058,1155,1523,1547]" name="Mayotte" pageId="26" pageNumber="327">Mayotte</collectingCountry>
. It was collected mostly from rainforest leaf litter, but also from rotten logs, under moss, and above ground in rainforest and montane rainforest in elevations between 20 and
<quantity id="E66DD2E2FFD3463AFDF3E356FD1A323E" box="[597,685,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.2349999999999999" pageId="26" pageNumber="327" unit="m" value="1235.0">1235 m</quantity>
. The populations on Anjouan, Grand Comore, and
<collectingCountry id="59823F97FFD3463AFB4CE357FAF0323F" box="[1258,1351,1595,1619]" name="Mayotte" pageId="26" pageNumber="327">Mayotte</collectingCountry>
islands are morphologically distinct, but share enough characteristics with each other and with the original
<typeStatus id="FE2EC1A5FFD3463AFB4BE333FAAA321B" box="[1261,1309,1631,1655]" pageId="26" pageNumber="327">type</typeStatus>
material to be identified as a single species, until further research proves otherwise. The specimens from Anjouan are all orange in color, like the
<typeStatus id="FE2EC1A5FFD3463AFE18E3CBFE4B32D3" box="[446,508,1703,1727]" pageId="26" pageNumber="327">types</typeStatus>
, whereas those from the other two islands are brown. The Grand Comore and
<collectingCountry id="59823F97FFD3463AFF31E3A7FF43328F" box="[151,244,1739,1763]" name="Mayotte" pageId="26" pageNumber="327">Mayotte</collectingCountry>
populations can be distinguished further by minor allometric differences in the head, pronotum and postpetiole. The majors from Grand Comore tend to have a slightly wider head (CI 99102 versus CI 9598) and a wider postpetiole (PpWI 218267 versus PpWI 279300) than those from
<collectingCountry id="59823F97FFD3463AFC54E27FFBE43347" box="[1010,1107,1811,1835]" name="Mayotte" pageId="26" pageNumber="327">Mayotte</collectingCountry>
.
<taxonomicName id="E6950484FFD3463AFBC6E27FFB0D3346" box="[1120,1210,1811,1834]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD3463AFBC6E27FFB0D3346" box="[1120,1210,1811,1834]" italics="true" pageId="26" pageNumber="327">P. jonas</emphasis>
</taxonomicName>
is probably closely related to
<taxonomicName id="E6950484FFD3463AFEA1E254FED93323" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[263,366,1847,1871]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="vulcan">
<emphasis id="13E1A315FFD3463AFEA1E254FED93323" box="[263,366,1847,1871]" italics="true" pageId="26" pageNumber="327">P. vulcan</emphasis>
</taxonomicName>
and
<taxonomicName id="E6950484FFD3463AFE03E254FE5C3323" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[421,491,1847,1871]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="loki">
<emphasis id="13E1A315FFD3463AFE03E254FE5C3323" box="[421,491,1847,1871]" italics="true" pageId="26" pageNumber="327">P. loki</emphasis>
</taxonomicName>
. The minor workers of
<taxonomicName id="E6950484FFD3463AFD51E254FCE73323" box="[759,848,1848,1871]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD3463AFD51E254FCE73323" box="[759,848,1848,1871]" italics="true" pageId="26" pageNumber="327">P. jonas</emphasis>
</taxonomicName>
can be separated from those of
<taxonomicName id="E6950484FFD3463AFB11E254FAAA3323" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[1207,1309,1847,1871]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="vulcan">
<emphasis id="13E1A315FFD3463AFB11E254FAAA3323" box="[1207,1309,1847,1871]" italics="true" pageId="26" pageNumber="327">P. vulcan</emphasis>
</taxonomicName>
by slightly longer mandibles, scapes, and postpetiole (MDI 6073, SI 102122, LPpI 122163 versus MDI 5463, SI 96106, LPpI
<quantity id="E66DD2E2FFD3463AFF7EE213FEEF33FB" box="[216,344,1919,1943]" metricMagnitude="0" metricUnit="m" metricValue="2.9718" metricValueMax="3.048" metricValueMin="2.8956" pageId="26" pageNumber="327" unit="in" value="117.0" valueMax="120.0" valueMin="114.0">114120 in</quantity>
<taxonomicName id="E6950484FFD3463AFEC7E2ECFE7E33FB" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[353,457,1919,1943]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="vulcan">
<emphasis id="13E1A315FFD3463AFEC7E2ECFE7E33FB" box="[353,457,1919,1943]" italics="true" pageId="26" pageNumber="327">P. vulcan</emphasis>
</taxonomicName>
), while the majors have a lower promesonotal process with a conspicuously concave transverse groove in profile view. Standing hairs are significantly less abundant and less flexuous in both worker castes than in those of
<taxonomicName id="E6950484FFD3463AFE3BE2A4FDB233B3" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[413,517,1991,2015]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="vulcan">
<emphasis id="13E1A315FFD3463AFE3BE2A4FDB233B3" box="[413,517,1991,2015]" italics="true" pageId="26" pageNumber="327">P. vulcan</emphasis>
</taxonomicName>
. The minor workers of
<taxonomicName id="E6950484FFD3463AFCB1E2A4FCE933B3" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[791,862,1991,2015]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="loki">
<emphasis id="13E1A315FFD3463AFCB1E2A4FCE933B3" box="[791,862,1991,2015]" italics="true" pageId="26" pageNumber="327">P. loki</emphasis>
</taxonomicName>
(majors are unknown) can be easily distinguished from the two other species, as the scapes are on average longer (SI 117122) and the sculpture on dorsal head and mesosoma is largely reduced, whereas the heads and mesosomas of
<taxonomicName id="E6950484FFD2463BFC3BE5D1FC4034B8" box="[925,1015,189,212]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="jonas">
<emphasis id="13E1A315FFD2463BFC3BE5D1FC4034B8" box="[925,1015,189,212]" italics="true" pageId="27" pageNumber="328">P. jonas</emphasis>
</taxonomicName>
and
<taxonomicName id="E6950484FFD2463BFB96E5D1FB2034B8" authorityName="Fischer &amp; Fisher" authorityYear="2013" box="[1072,1175,188,212]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="vulcan">
<emphasis id="13E1A315FFD2463BFB96E5D1FB2034B8" box="[1072,1175,188,212]" italics="true" pageId="27" pageNumber="328">P. vulcan</emphasis>
</taxonomicName>
possess comparatively strong punctures.
</paragraph>
</subSubSection>
<subSubSection id="698F2C8CFFD2463BFF61E46FFD073730" pageId="27" pageNumber="328" type="materials_examined">
<paragraph id="212A7F07FFD2463BFF61E46FFD073730" blockId="27.[151,1436,151,860]" pageId="27" pageNumber="328">
<emphasis id="13E1A315FFD2463BFF61E46FFD883570" bold="true" box="[199,575,259,284]" pageId="27" pageNumber="328">Additional material examined:</emphasis>
<materialsCitation id="91FD755AFFD2463BFDEDE46FFCE43553" ID-GBIF-Occurrence="3736261458" collectingDate="2009-01-25" collectorName="B. L. Fisher" country="Comoros" elevation="900" latitude="-12.2247" location="Lac Dzialandee" longLatPrecision="7" longitude="44.4312" pageId="27" pageNumber="328" specimenCount="1" stateProvince="Anjouan">
<emphasis id="13E1A315FFD2463BFDEDE46FFD683570" bold="true" box="[587,735,259,284]" pageId="27" pageNumber="328">
<collectingCountry id="59823F97FFD2463BFDEDE46FFD683570" box="[587,735,259,284]" name="Comoros" pageId="27" pageNumber="328">COMOROS</collectingCountry>
</emphasis>
:
<emphasis id="13E1A315FFD2463BFD55E46FFCEC3570" bold="true" box="[755,859,259,284]" pageId="27" pageNumber="328">
<collectingRegion id="E351B1E5FFD2463BFD55E46FFCEC3570" box="[755,859,259,284]" country="Comoros" name="Anjouan" pageId="27" pageNumber="328">Anjouan</collectingRegion>
</emphasis>
:
<location id="244A29DCFFD2463BFCC9E468FB923570" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFCC9E468FB923570" box="[879,1061,260,284]" country="Comoros" latitude="-12.2247" longLatPrecision="7" longitude="44.4312" name="Lac Dzialandee" pageId="27" pageNumber="328" stateProvince="Anjouan">Lac Dzialandee</location>
,
<geoCoordinate id="44A119C0FFD2463BFB93E468FB1A3570" box="[1077,1197,260,284]" degrees="12.2247" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.2247">12.2247 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFB18E468FA8F3577" box="[1214,1336,260,284]" degrees="44.4312" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="44.4312">44.4312 E</geoCoordinate>
, 900 m, disturbed montane rainforest,
<date id="552B59C7FFD2463BFE53E44BFDE83553" box="[501,607,295,319]" pageId="27" pageNumber="328" value="2009-01-25">
<collectingDate id="456FA02FFFD2463BFE53E44BFDE83553" box="[501,607,295,319]" pageId="27" pageNumber="328" value="2009-01-25">25.i.2009</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFDD2E445FC8C352C" box="[628,827,296,320]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFDD2E445FD4F352C" box="[628,760,296,320]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFCF8E444FD223508" ID-GBIF-Occurrence="3736261454" collectingDate="2009-01-24" collectorName="B. L. Fisher" elevation="1225" latitude="-12.2204" location="Mount Ntringui" longLatPrecision="7" longitude="44.4292" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFCF8E444FBA03553" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFCF8E444FBA03553" box="[862,1047,295,319]" latitude="-12.2204" longLatPrecision="7" longitude="44.4292" name="Mount Ntringui" pageId="27" pageNumber="328">Mount Ntringui</location>
,
<geoCoordinate id="44A119C0FFD2463BFB81E44BFB283553" box="[1063,1183,295,319]" degrees="12.2204" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.2204">12.2204 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFB16E44BFA9D3553" box="[1200,1322,295,319]" degrees="44.4292" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="44.4292">44.4292 E</geoCoordinate>
, 1225 m, montane forest,
<date id="552B59C7FFD2463BFEEAE420FE023508" box="[332,437,332,356]" pageId="27" pageNumber="328" value="2009-01-24">
<collectingDate id="456FA02FFFD2463BFEEAE420FE023508" box="[332,437,332,356]" pageId="27" pageNumber="328" value="2009-01-24">24.i.2009</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFE60E421FDCB3508" box="[454,636,332,356]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFE60E421FDF33508" box="[454,580,332,356]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFD3DE427FDD935EB" ID-GBIF-Occurrence="3736261560" collectingDate="2008-03-13" collectorName="B. L. Fisher" elevation="1125" latitude="-11.8134" location="Karthala" longLatPrecision="7" longitude="43.4194" pageId="27" pageNumber="328" specimenCount="1" stateProvince="Grande Comore">
<emphasis id="13E1A315FFD2463BFD3DE427FCE93508" bold="true" box="[667,862,331,356]" pageId="27" pageNumber="328">
<collectingRegion id="E351B1E5FFD2463BFD3DE427FCE93508" box="[667,862,331,356]" country="Comoros" name="Grande Comore" pageId="27" pageNumber="328">Grande Comore</collectingRegion>
</emphasis>
:
<location id="244A29DCFFD2463BFCCAE420FC783508" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFCCAE420FC783508" box="[876,975,332,356]" latitude="-11.8134" longLatPrecision="7" longitude="43.4194" name="Karthala" pageId="27" pageNumber="328" stateProvince="Grande Comore">Karthala</location>
,
<geoCoordinate id="44A119C0FFD2463BFC7DE420FBFA3508" box="[987,1101,332,356]" degrees="11.8134" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-11.8134">11.8134 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFBFFE420FB79350F" box="[1113,1230,332,356]" degrees="43.4194" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="43.4194">43.4194 E</geoCoordinate>
, 1125 m, montane rainforest,
<date id="552B59C7FFD2463BFEB7E403FE3D35EB" box="[273,394,367,391]" pageId="27" pageNumber="328" value="2008-03-13">
<collectingDate id="456FA02FFFD2463BFEB7E403FE3D35EB" box="[273,394,367,391]" pageId="27" pageNumber="328" value="2008-03-13">13.iii.2008</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFE3AE41DFDE135E4" box="[412,598,368,392]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFE3AE41DFDAC35E4" box="[412,539,368,392]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFDD3E403FEC335C0" ID-GBIF-Occurrence="3736261520" collectingDate="2008-03-14" collectorName="B. L. Fisher" elevation="1000" latitude="-11.827" location="Karthala" longLatPrecision="55" longitude="43.4295" municipality="Karthala" pageId="27" pageNumber="328" specimenCount="1">
<collectingMunicipality id="C14EE57DFFD2463BFDD3E403FD6F35EB" box="[629,728,367,391]" pageId="27" pageNumber="328">Karthala</collectingMunicipality>
,
<geoCoordinate id="44A119C0FFD2463BFD43E403FCFD35EB" box="[741,842,367,391]" degrees="11.827" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="55" value="-11.827">11.827 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFCF1E403FC7935EB" box="[855,974,367,391]" degrees="43.4295" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="43.4295">43.4295 E</geoCoordinate>
, 1000 m, montane rainforest,
<date id="552B59C7FFD2463BFA84E403FA2C35EB" box="[1314,1435,367,391]" pageId="27" pageNumber="328" value="2008-03-14">
<collectingDate id="456FA02FFFD2463BFA84E403FA2C35EB" box="[1314,1435,367,391]" pageId="27" pageNumber="328" value="2008-03-14">14.iii.2008</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFF06E4F9FEEB35C0" box="[160,348,404,428]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFF06E4F9FE9735C0" box="[160,288,404,428]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFEDBE4F8FA2B35C0" ID-GBIF-Occurrence="3736261541" box="[381,1436,404,428]" collectingDate="2008-03-17" collectorName="B. L. Fisher" elevation="1235" latitude="-11.7533" location="Karthala" longLatPrecision="7" longitude="43.3079" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFEDBE4F8FE5735C0" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFEDBE4F8FE5735C0" box="[381,480,404,428]" latitude="-11.7533" longLatPrecision="7" longitude="43.3079" name="Karthala" pageId="27" pageNumber="328">Karthala</location>
,
<geoCoordinate id="44A119C0FFD2463BFE4BE4F8FDD735C0" box="[493,608,404,428]" degrees="11.7533" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-11.7533">11.7533 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFDC8E4F8FD5235C7" box="[622,741,404,428]" degrees="43.3079" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="43.3079">43.3079 E</geoCoordinate>
, 1235 m, montane rainforest,
<date id="552B59C7FFD2463BFB9BE4F8FB0135C0" box="[1085,1206,404,428]" pageId="27" pageNumber="328" value="2008-03-17">
<collectingDate id="456FA02FFFD2463BFB9BE4F8FB0135C0" box="[1085,1206,404,428]" pageId="27" pageNumber="328" value="2008-03-17">17.iii.2008</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFB6EE4F9FA3335C0" box="[1224,1412,404,428]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFB6EE4F9FAFF35C0" box="[1224,1352,404,428]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFF31E4DBFB2C35A3" ID-GBIF-Occurrence="3736261373" box="[151,1179,439,464]" collectingDate="2009-01-21" collectorName="B. L. Fisher" elevation="275" latitude="-12.3067" location="Ouallah" longLatPrecision="7" longitude="43.6641" pageId="27" pageNumber="328" specimenCount="1" stateProvince="Moheli">
<emphasis id="13E1A315FFD2463BFF31E4DBFF5C35BC" bold="true" box="[151,235,439,464]" pageId="27" pageNumber="328">
<collectingRegion id="E351B1E5FFD2463BFF31E4DBFF5C35BC" box="[151,235,439,464]" country="Comoros" name="Moheli" pageId="27" pageNumber="328">Mohéli</collectingRegion>
</emphasis>
:
<location id="244A29DCFFD2463BFF5AE4DBFEE035A3" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFF5AE4DBFEE035A3" box="[252,343,439,463]" latitude="-12.3067" longLatPrecision="7" longitude="43.6641" name="Ouallah" pageId="27" pageNumber="328" stateProvince="Moheli">Ouallah</location>
,
<geoCoordinate id="44A119C0FFD2463BFEC3E4DBFE6C35A3" box="[357,475,439,463]" degrees="12.3067" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.3067">12.3067 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFE4DE4DBFDD535A3" box="[491,610,439,463]" degrees="43.6641" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="43.6641">43.6641 E</geoCoordinate>
, 275 m, rainforest,
<date id="552B59C7FFD2463BFCE1E4DBFC0635A3" box="[839,945,439,463]" pageId="27" pageNumber="328" value="2009-01-21">
<collectingDate id="456FA02FFFD2463BFCE1E4DBFC0635A3" box="[839,945,439,463]" pageId="27" pageNumber="328" value="2009-01-21">21.i.2009</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFC62E4D5FB3435BC" box="[964,1155,440,464]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFC62E4D5FBF235BC" box="[964,1093,440,464]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFB02E4DBFBFB3598" ID-GBIF-Occurrence="3736261400" collectingDate="1999-01-15" collectorName="R. Jocque &amp; G. DeSmet" country="Mayotte" latitude="-12.8333" location="Campus" longLatPrecision="7" longitude="45.1333" municipality="Coconi" pageId="27" pageNumber="328" specimenCount="1">
<emphasis id="13E1A315FFD2463BFB02E4DBFA8535BC" bold="true" box="[1188,1330,439,464]" pageId="27" pageNumber="328">
<collectingCountry id="59823F97FFD2463BFB02E4DBFA8535BC" box="[1188,1330,439,464]" name="Mayotte" pageId="27" pageNumber="328">MAYOTTE</collectingCountry>
</emphasis>
:
<collectingMunicipality id="C14EE57DFFD2463BFAE2E4DBFA2F35A3" box="[1348,1432,439,463]" pageId="27" pageNumber="328">Coconi</collectingMunicipality>
, DAF
<location id="244A29DCFFD2463BFF47E4B0FEF73598" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFF47E4B0FEF73598" box="[225,320,476,500]" country="Mayotte" latitude="-12.8333" longLatPrecision="7" longitude="45.1333" municipality="Coconi" name="Campus" pageId="27" pageNumber="328">Campus</location>
,
<geoCoordinate id="44A119C0FFD2463BFEF0E4B0FE633598" box="[342,468,476,500]" degrees="12.8333" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8333">12.8333 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFE4DE4B0FDDC359F" box="[491,619,476,500]" degrees="45.1333" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1333">45.1333 E</geoCoordinate>
,
<date id="552B59C7FFD2463BFD24E4B0FD5B3598" box="[642,748,476,500]" pageId="27" pageNumber="328" value="1999-01-15">
<collectingDate id="456FA02FFFD2463BFD24E4B0FD5B3598" box="[642,748,476,500]" pageId="27" pageNumber="328" value="1999-01-15">15.i.1999</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFCA1E4B1FB8C3598" box="[775,1083,476,500]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFCA1E4B1FB8C3598" box="[775,1083,476,500]" pageId="27" pageNumber="328">R. Jocque &amp; G. DeSmet</collectorName>
</emphasis>
);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFBF8E4B0FF483650" ID-GBIF-Occurrence="3736261467" collectingDate="1999-02-18" collectorName="R. Jocque &amp; G. DeSmet" latitude="-12.8333" location="Service de Developpement Agricole" longLatPrecision="7" longitude="45.1333" municipality="Coconi" pageId="27" pageNumber="328" specimenCount="1">
<collectingMunicipality id="C14EE57DFFD2463BFBF8E4B0FB063598" box="[1118,1201,476,500]" pageId="27" pageNumber="328">Coconi</collectingMunicipality>
, SDA (
<location id="244A29DCFFD2463BFABCE4B0FE0B367B" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFABCE4B0FE0B367B" latitude="-12.8333" longLatPrecision="7" longitude="45.1333" municipality="Coconi" name="Service de Developpement Agricole" pageId="27" pageNumber="328">Service de Developpement Agricole</location>
),
<geoCoordinate id="44A119C0FFD2463BFE70E493FDF8367B" box="[470,591,511,535]" degrees="12.8333" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8333">12.8333 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFDC4E493FD69367B" box="[610,734,511,535]" degrees="45.1333" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1333">45.1333 E</geoCoordinate>
, wet area with giant bamboo,
<date id="552B59C7FFD2463BFBF0E493FB70367B" box="[1110,1223,511,535]" pageId="27" pageNumber="328" value="1999-02-18">
<collectingDate id="456FA02FFFD2463BFBF0E493FB70367B" box="[1110,1223,511,535]" pageId="27" pageNumber="328" value="1999-02-18">18.ii.1999</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFB78E76DFF593650" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFB78E76DFF593650" pageId="27" pageNumber="328">R. Jocque &amp; G. DeSmet</collectorName>
</emphasis>
);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFEA0E748FB2F3650" ID-GBIF-Occurrence="3736261345" box="[262,1176,548,572]" collectingDate="2007-12-02" collectorName="B. L. Fisher" elevation="135" latitude="-12.9628" location="Dapani" longLatPrecision="7" longitude="45.1504" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFEA0E748FEED3650" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFEA0E748FEED3650" box="[262,346,548,572]" latitude="-12.9628" longLatPrecision="7" longitude="45.1504" name="Dapani" pageId="27" pageNumber="328">Dapani</location>
,
<geoCoordinate id="44A119C0FFD2463BFEC0E748FE6E3650" box="[358,473,548,572]" degrees="12.9628" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.9628">12.9628 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFE40E748FDEC3657" box="[486,603,548,572]" degrees="45.1504" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1504">45.1504 E</geoCoordinate>
, 135 m, rainforest,
<date id="552B59C7FFD2463BFC91E748FC003650" box="[823,951,548,572]" pageId="27" pageNumber="328" value="2007-12-02">
<collectingDate id="456FA02FFFD2463BFC91E748FC003650" box="[823,951,548,572]" pageId="27" pageNumber="328" value="2007-12-02">02.xii.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFC61E749FBC83650" box="[967,1151,548,572]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFC61E749FBF13650" box="[967,1094,548,572]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFB39E748FC343633" ID-GBIF-Occurrence="3736261540" collectingDate="2007-11-30" collectorName="B. L. Fisher" elevation="425" latitude="-12.8758" location="Mont Benara" longLatPrecision="7" longitude="45.1567" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFB39E748FA823657" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFB39E748FA823657" box="[1183,1333,548,571]" latitude="-12.8758" longLatPrecision="7" longitude="45.1567" name="Mont Benara" pageId="27" pageNumber="328">Mont Benara</location>
,
<geoCoordinate id="44A119C0FFD2463BFAE7E748FF1F3633" degrees="12.8758" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8758">12.8758 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFF1EE72BFE863633" box="[184,305,583,607]" degrees="45.1567" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1567">45.1567 E</geoCoordinate>
, 425 m, rainforest,
<date id="552B59C7FFD2463BFDBAE72BFD243633" box="[540,659,583,607]" pageId="27" pageNumber="328" value="2007-11-30">
<collectingDate id="456FA02FFFD2463BFDBAE72BFD243633" box="[540,659,583,607]" pageId="27" pageNumber="328" value="2007-11-30">30.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFD01E725FCDD360C" box="[679,874,584,608]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFD01E725FC9D360C" box="[679,810,584,608]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFC2BE724FDDE36E8" ID-GBIF-Occurrence="3736261585" collectingDate="2007-11-28" collectorName="B. L. Fisher" elevation="470" latitude="-12.9578" location="Mont Chongui" longLatPrecision="54" longitude="45.134" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFC2BE724FB8F3633" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFC2BE724FB8F3633" box="[909,1080,583,607]" latitude="-12.9578" longLatPrecision="54" longitude="45.134" name="Mont Chongui" pageId="27" pageNumber="328">Mont Chongui</location>
,
<geoCoordinate id="44A119C0FFD2463BFBE1E72BFB093633" box="[1095,1214,583,607]" degrees="12.9578" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.9578">12.9578 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFB69E72BFA8D3633" box="[1231,1338,583,607]" degrees="45.134" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="55" value="45.134">45.134 E</geoCoordinate>
, 470 m, rainforest,
<date id="552B59C7FFD2463BFEB6E700FE3F36E8" box="[272,392,620,644]" pageId="27" pageNumber="328" value="2007-11-28">
<collectingDate id="456FA02FFFD2463BFEB6E700FE3F36E8" box="[272,392,620,644]" pageId="27" pageNumber="328" value="2007-11-28">28.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFE3FE701FDE636E8" box="[409,593,620,644]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFE3FE701FDAF36E8" box="[409,536,620,644]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFDD6E700FE8F36CB" ID-GBIF-Occurrence="3736261328" collectingDate="2007-11-28" collectorName="B. L. Fisher" elevation="380" latitude="-12.959" location="Mont Chongui" longLatPrecision="55" longitude="45.1341" municipality="Mont Chongui" pageId="27" pageNumber="328" specimenCount="1">
<collectingMunicipality id="C14EE57DFFD2463BFDD6E700FCA036E8" box="[624,791,620,644]" pageId="27" pageNumber="328">Mont Chongui</collectingMunicipality>
,
<geoCoordinate id="44A119C0FFD2463BFC84E700FC3036E8" box="[802,903,620,644]" degrees="12.959" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="55" value="-12.959">12.959 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFC32E700FBBD36EF" box="[916,1034,620,644]" degrees="45.1341" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1341">45.1341 E</geoCoordinate>
, 380 m, rainforest,
<date id="552B59C7FFD2463BFB43E700FAEA36E8" box="[1253,1373,620,644]" pageId="27" pageNumber="328" value="2007-11-28">
<collectingDate id="456FA02FFFD2463BFB43E700FAEA36E8" box="[1253,1373,620,644]" pageId="27" pageNumber="328" value="2007-11-28">28.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFACBE701FE9736C4" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFACBE701FF5736C4" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFEE5E7FCFAE236CB" ID-GBIF-Occurrence="3736261368" box="[323,1365,655,680]" collectingDate="2007-11-25" collectorName="B. L. Fisher" elevation="370" latitude="-12.8063" location="Mont Combani" longLatPrecision="7" longitude="45.1531" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFEE5E7FCFE4436CB" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFEE5E7FCFE4436CB" box="[323,499,655,679]" latitude="-12.8063" longLatPrecision="7" longitude="45.1531" name="Mont Combani" pageId="27" pageNumber="328">Mont Combani</location>
,
<geoCoordinate id="44A119C0FFD2463BFDA5E7E3FDCD36CB" box="[515,634,655,679]" degrees="12.8063" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8063">12.8063 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFD2CE7E3FCB336CB" box="[650,772,655,679]" degrees="45.1531" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1531">45.1531 E</geoCoordinate>
, 370 m, rainforest,
<date id="552B59C7FFD2463BFC48E7E3FBD236CB" box="[1006,1125,655,679]" pageId="27" pageNumber="328" value="2007-11-25">
<collectingDate id="456FA02FFFD2463BFC48E7E3FBD236CB" box="[1006,1125,655,679]" pageId="27" pageNumber="328" value="2007-11-25">25.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFBDCE7FDFA8B36C4" box="[1146,1340,656,680]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFBDCE7FDFB4B36C4" box="[1146,1276,656,680]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFAF9E7FCFB8A36A0" ID-GBIF-Occurrence="3736261364" collectingDate="2007-11-29" collectorName="B. L. Fisher" elevation="460" latitude="-12.8049" location="Mont Combani" longLatPrecision="7" longitude="45.1527" municipality="Mont Combani" pageId="27" pageNumber="328" specimenCount="1">
<collectingMunicipality id="C14EE57DFFD2463BFAF9E7FCFEB636A0" pageId="27" pageNumber="328">Mont Combani</collectingMunicipality>
,
<geoCoordinate id="44A119C0FFD2463BFEABE7D8FE3736A0" box="[269,384,692,716]" degrees="12.8049" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8049">12.8049 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFE28E7D8FDB336A7" box="[398,516,692,716]" degrees="45.1527" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1527">45.1527 E</geoCoordinate>
, 460 m, rainforest,
<date id="552B59C7FFD2463BFD44E7D8FCEE36A0" box="[738,857,692,716]" pageId="27" pageNumber="328" value="2007-11-29">
<collectingDate id="456FA02FFFD2463BFD44E7D8FCEE36A0" box="[738,857,692,716]" pageId="27" pageNumber="328" value="2007-11-29">29.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFCCDE7D9FB9336A0" box="[875,1060,692,716]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFCCDE7D9FC5D36A0" box="[875,1002,692,716]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFBE2E7D8FC063683" ID-GBIF-Occurrence="3736261496" collectingDate="1999-02-22" collectorName="R. Jocque &amp; G. DeSmet" elevation="470" latitude="-12.8" location="Mt. Combani" longLatPrecision="5555" longitude="45.1333" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFBE2E7D8FB6936A0" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFBE2E7D8FB6936A0" box="[1092,1246,692,716]" latitude="-12.8" longLatPrecision="5555" longitude="45.1333" name="Mt. Combani" pageId="27" pageNumber="328">Mt. Combani</location>
,
<geoCoordinate id="44A119C0FFD2463BFB4CE7D8FA8336A0" box="[1258,1332,692,716]" degrees="12.8" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5555" value="-12.8">12.8 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFAE7E7D8FF1D3683" degrees="45.1333" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1333">45.1333 E</geoCoordinate>
, 470 m / 420 m, forest litter,
<date id="552B59C7FFD2463BFDA1E7BBFDCF3683" box="[519,632,727,751]" pageId="27" pageNumber="328" value="1999-02-22">
<collectingDate id="456FA02FFFD2463BFDA1E7BBFDCF3683" box="[519,632,727,751]" pageId="27" pageNumber="328" value="1999-02-22">22.ii.1999</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFD2AE7B5FC17369C" box="[652,928,728,752]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFD2AE7B5FC17369C" box="[652,928,728,752]" pageId="27" pageNumber="328">R. Jocque &amp; G. DeSmet</collectorName>
</emphasis>
);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFC1DE7B4FCFD3778" ID-GBIF-Occurrence="3736261590" collectingDate="2007-12-03" collectorName="B. L. Fisher" elevation="350" latitude="-12.7689" location="Reserve Forestiere Majimbini" longLatPrecision="7" longitude="45.1902" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFC1DE7B4FAA53683" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFC1DE7B4FAA53683" box="[955,1298,727,751]" latitude="-12.7689" longLatPrecision="7" longitude="45.1902" name="Reserve Forestiere Majimbini" pageId="27" pageNumber="328">Reserve Forestière Majimbini</location>
,
<geoCoordinate id="44A119C0FFD2463BFA86E7BBFA213683" box="[1312,1430,727,751]" degrees="12.7689" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.7689">12.7689 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFF31E790FEBB377F" box="[151,268,764,788]" degrees="45.1902" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1902">45.1902 E</geoCoordinate>
, 350 m, rainforest,
<date id="552B59C7FFD2463BFE4FE790FDDF3778" box="[489,616,764,788]" pageId="27" pageNumber="328" value="2007-12-03">
<collectingDate id="456FA02FFFD2463BFE4FE790FDDF3778" box="[489,616,764,788]" pageId="27" pageNumber="328" value="2007-12-03">03.xii.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFDDFE791FC863778" box="[633,817,764,788]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFDDFE791FD4F3778" box="[633,760,764,788]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFCF7E790FD7D375B" ID-GBIF-Occurrence="3736261390" collectingDate="2007-12-03" collectorName="B. L. Fisher" elevation="525" latitude="-12.768" location="Reserve Forestiere Majimbini" longLatPrecision="55" longitude="45.1861" municipality="Reserve Forestiere Majimbini" pageId="27" pageNumber="328" specimenCount="1">
<collectingMunicipality id="C14EE57DFFD2463BFCF7E790FB143778" box="[849,1187,764,788]" pageId="27" pageNumber="328">Reserve Forestière Majimbini</collectingMunicipality>
,
<geoCoordinate id="44A119C0FFD2463BFB09E790FAA33778" box="[1199,1300,764,788]" degrees="12.768" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="55" value="-12.768">12.768 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFA87E790FA20377F" box="[1313,1431,764,788]" degrees="45.1861" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1861">45.1861 E</geoCoordinate>
, 525 m, rainforest,
<date id="552B59C7FFD2463BFECEE673FE50375B" box="[360,487,799,823]" pageId="27" pageNumber="328" value="2007-12-03">
<collectingDate id="456FA02FFFD2463BFECEE673FE50375B" box="[360,487,799,823]" pageId="27" pageNumber="328" value="2007-12-03">03.xii.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFE5EE64DFD053754" box="[504,690,800,824]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFE5EE64DFDCF3754" box="[504,632,800,824]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.);
</materialsCitation>
<materialsCitation id="91FD755AFFD2463BFD74E64CFD073730" ID-GBIF-Occurrence="3736261463" collectingDate="2007-11-29" collectorName="B. L. Fisher" elevation="20" latitude="-12.8059" location="Reserve Forestiere Sohoa" longLatPrecision="7" longitude="45.1005" pageId="27" pageNumber="328" specimenCount="1">
<location id="244A29DCFFD2463BFD74E64CFC43375B" LSID="urn:lsid:plazi:treatment:A93CCE11FFD1463BFF31E07BFD073730:244A29DCFFD2463BFD74E64CFC43375B" box="[722,1012,799,823]" latitude="-12.8059" longLatPrecision="7" longitude="45.1005" name="Reserve Forestiere Sohoa" pageId="27" pageNumber="328">Reserve Forestière Sohoa</location>
,
<geoCoordinate id="44A119C0FFD2463BFBA7E673FBC2375B" box="[1025,1141,799,823]" degrees="12.8059" direction="south" orientation="latitude" pageId="27" pageNumber="328" precision="5" value="-12.8059">12.8059 S</geoCoordinate>
,
<geoCoordinate id="44A119C0FFD2463BFB25E673FB4F375B" box="[1155,1272,799,823]" degrees="45.1005" direction="east" orientation="longitude" pageId="27" pageNumber="328" precision="5" value="45.1005">45.1005 E</geoCoordinate>
, 20 m, coastal scrub, rainforest,
<date id="552B59C7FFD2463BFEFDE628FE643730" box="[347,467,836,860]" pageId="27" pageNumber="328" value="2007-11-29">
<collectingDate id="456FA02FFFD2463BFEFDE628FE643730" box="[347,467,836,860]" pageId="27" pageNumber="328" value="2007-11-29">29.xi.2007</collectingDate>
</date>
(
<emphasis id="13E1A315FFD2463BFE45E629FD2D3730" box="[483,666,836,860]" italics="true" pageId="27" pageNumber="328">
<collectorName id="8C601AD1FFD2463BFE45E629FDD63730" box="[483,609,836,860]" pageId="27" pageNumber="328">B.L. Fisher</collectorName>
et al
</emphasis>
.).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>