268 lines
52 KiB
XML
268 lines
52 KiB
XML
<document id="39CAABF81B0C1D69B291E81BA74A701E" ID-DOI="10.11646/zootaxa.5353.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="8430672" ID-ZooBank="16BD810F-4FB7-4D65-995E-1BA6FCBB2788" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697020094218" checkinUser="plazi" docAuthor="Rafael, José A., Marques, Dayse W. A. & Limeira-De-Oliveira, F." docDate="2023" docId="9C3387CEFFAFB95DFF17FE0FA143FE6B" docLanguage="en" docName="zootaxa.5353.4.2.pdf" docOrigin="Zootaxa 5353 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.4.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Czernyola fumialula Rafael & Marques & Limeira-De-Oliveira 2023, sp. nov." docType="treatment" docUuid="DDA72823-2531-49BC-BAFB-53819B4782EB" docUuidSource="ZooBank" docVersion="6" lastPageNumber="334" masterDocId="600AFFB6FFADB959FF80FFCAA246FFE2" masterDocTitle="Insect (Hexapoda) Diversity in the Oceanic Archipelago of Fernando de Noronha, Brazil: Clusiidae (Diptera)" masterLastPageNumber="350" masterPageNumber="332" pageNumber="334" updateTime="1719815327585" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="22183733421DE5E8B65F678FF5659CA5">
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<mods:title id="736F89C6C7C5C950AD67DF772BD6F073">Insect (Hexapoda) Diversity in the Oceanic Archipelago of Fernando de Noronha, Brazil: Clusiidae (Diptera)</mods:title>
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<mods:name id="64FAE57B0EB35FB505715F2D23F516D4" type="personal">
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<mods:roleTerm id="E36BD2EE9C8DF2E66264AAC2FC1506A0">Author</mods:roleTerm>
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<mods:namePart id="A2F2BC6D6278710E419B07B4AAF08A2F">Rafael, José A.</mods:namePart>
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<mods:nameIdentifier id="A2AA830486FA5563B63DB4939A1E4633" type="ORCID">0000-0002-0170-0514</mods:nameIdentifier>
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<mods:affiliation id="0C8619E2A35B7876C67B63F8F862AD1A">Instituto Nacional de Pesquisas da Amazônia, INPA, Coordenação de Biodiversidade, Manaus, Amazonas, Brazil jarafael @ inpa. gov. br; https: // orcid. org / 0000 - 0002 - 0170 - 0514</mods:affiliation>
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<mods:nameIdentifier id="2090624D56DB34F1F5CD11CE8F78A58E" type="email">jarafael@inpa.gov.br</mods:nameIdentifier>
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<mods:namePart id="2F875641997E97C72AB940FD2F1DC28F">Marques, Dayse W. A.</mods:namePart>
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<mods:nameIdentifier id="D84D36137C3BB8FDCDB373D02F59369A" type="ORCID">0000-0001-7260-5760</mods:nameIdentifier>
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<mods:affiliation id="9138983025759FB606AA70F1EFBF183A">Instituto Nacional de Pesquisas da Amazônia, INPA, Programa de Pós-graduação em Entomologia, Manaus, Amazonas, Brazil willkenia @ gmail. com; https: // orcid. org / 0000 - 0001 - 7260 - 5760</mods:affiliation>
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<mods:nameIdentifier id="4EEA0EB72FBB7ED1188C612CB1D7CE49" type="email">willkenia@gmail.com</mods:nameIdentifier>
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<mods:namePart id="41999B155157031689B9924961AD6784">Limeira-De-Oliveira, F.</mods:namePart>
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<mods:affiliation id="BB6BA8549A388D383F1548FFFEB3989B">Universidade Estadual do Maranhão, Laboratório de Estudos dos Invertebrados, 65604 - 380, Caxias, Maranhão, Brazil limeiraf @ gmail. com; https: // orcid. org / 0000 - 0002 - 4980 - 5454</mods:affiliation>
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<mods:nameIdentifier id="4451B09F374AFA3A5055D6FFCA023BAF" type="email">limeiraf@gmail.com</mods:nameIdentifier>
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<mods:title id="0721829F93FBAE702452F0C8D6FD89AE">Zootaxa</mods:title>
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<mods:part id="C4C11ED35EFF6AA83F3B21E6EF390202">
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<mods:date id="AAABA999D40C709F2D939148EEB6666F">2023</mods:date>
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<mods:number id="AE63314DD2B7AFF1073C3596FA697219">2023-10-11</mods:number>
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<mods:number id="94C8239C430E0C7CD79DB30C1CD6D46F">5353</mods:number>
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<mods:url id="9CD5726E464DC2ACDB56736ED5F22AE8">http://dx.doi.org/10.11646/zootaxa.5353.4.2</mods:url>
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<mods:classification id="1CBB5120DB64607259E4CD3975F5BE4F">journal article</mods:classification>
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<mods:identifier id="15201AD06E125A69BA73A114292E2DB4" type="DOI">10.11646/zootaxa.5353.4.2</mods:identifier>
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<treatment id="9C3387CEFFAFB95DFF17FE0FA143FE6B" ID-DOI="http://doi.org/10.5281/zenodo.8430662" ID-Zenodo-Dep="8430662" LSID="urn:lsid:zoobank.org:act:DDA72823-2531-49BC-BAFB-53819B4782EB" httpUri="http://treatment.plazi.org/id/9C3387CEFFAFB95DFF17FE0FA143FE6B" lastPageId="4" lastPageNumber="334" pageId="2" pageNumber="334">
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<subSubSection id="5C806553FFAFB95BFF17FE0FA189FDE6" pageId="2" pageNumber="334" type="nomenclature">
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<paragraph id="142536D8FFAFB95BFF17FE0FA3B3FE02" blockId="2.[151,975,453,552]" box="[151,501,453,480]" pageId="2" pageNumber="334">
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<heading id="4F6D81B4FFAFB95BFF17FE0FA3B3FE02" bold="true" box="[151,501,453,480]" fontSize="11" level="1" pageId="2" pageNumber="334" reason="1">
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<emphasis id="26EEEACAFFAFB95BFF17FE0FA3B3FE02" bold="true" box="[151,501,453,480]" pageId="2" pageNumber="334">
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<taxonomicName id="D39A4D5BFFAFB95BFF17FE0FA3D6FE02" ID-CoL="C2NFB" authority="Rafael & Marques & Limeira-De-Oliveira, 2023" authorityName="Rafael & Marques & Limeira-De-Oliveira" authorityYear="2023" box="[151,400,453,480]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="fumialula" status="sp. nov.">
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<emphasis id="26EEEACAFFAFB95BFF17FE0FA3D6FE02" bold="true" box="[151,400,453,480]" italics="true" pageId="2" pageNumber="334">Czernyola fumialula</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3DDD57B1FFAFB95BFE17FE0CA3B3FE02" box="[407,501,454,480]" pageId="2" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="142536D8FFAFB95BFF17FE23A189FDE6" blockId="2.[151,975,453,552]" box="[151,975,489,516]" pageId="2" pageNumber="334">
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urn:lsid:zoobank.org:act:
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<uuid id="603C0C0DFFAFB95BFE24FE23A189FDE6" box="[420,975,489,516]" pageId="2" pageNumber="334">DDA72823-2531-49BC-BAFB-53819B4782EB</uuid>
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<subSubSection id="5C806553FFAFB95BFF17FDC7A35EFDCA" box="[151,280,525,552]" pageId="2" pageNumber="334" type="description">
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<paragraph id="142536D8FFAFB95BFF17FDC7A35EFDCA" blockId="2.[151,975,453,552]" box="[151,280,525,552]" pageId="2" pageNumber="334">
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(
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<figureCitation id="8CA12A5DFFAFB95BFF1FFDC7A349FDCA" box="[159,271,525,552]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 1–11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="5C806553FFAFB95BFF17FD85A06CFD52" pageId="2" pageNumber="334" type="diagnosis">
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<paragraph id="142536D8FFAFB95BFF17FD85A06CFD52" blockId="2.[151,1437,591,2038]" pageId="2" pageNumber="334">
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<emphasis id="26EEEACAFFAFB95BFF17FD85A34CFD8B" bold="true" box="[151,266,591,617]" pageId="2" pageNumber="334">Diagnosis</emphasis>
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. Postcranium black. Scutum black; mediotergite yellow; laterotergite black. Mesopleuron yellow with anepisternum black. Wing light yellow basally, dusky from the level of the r-m vein to the apex of the vein. Alula brown infuscated. Terminalia black.
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</paragraph>
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</subSubSection>
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<subSubSection id="5C806553FFAFB95BFF47FD73A3E4FA5E" pageId="2" pageNumber="334" type="description">
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<paragraph id="142536D8FFAFB95BFF47FD73A624FA7A" blockId="2.[151,1437,591,2038]" pageId="2" pageNumber="334">
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<emphasis id="26EEEACAFFAFB95BFF47FD73A317FD31" bold="true" box="[199,337,697,723]" pageId="2" pageNumber="334">Description</emphasis>
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.
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<typeStatus id="CB21887AFFAFB95BFEE1FD73A38FFD31" box="[353,457,697,723]" pageId="2" pageNumber="334" type="holotype">Holotype</typeStatus>
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♂ (
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<figureCitation id="8CA12A5DFFAFB95BFE77FD73A07CFD31" box="[503,570,697,723]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 1</figureCitation>
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). Body length
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<quantity id="D3629B3DFFAFB95BFD67FD73A107FD36" box="[743,833,697,724]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="334" unit="mm" value="2.8">2.8 mm</quantity>
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; wing length
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<quantity id="D3629B3DFFAFB95BFC5DFD73A671FD36" box="[989,1079,697,724]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="334" unit="mm" value="2.4">2.4 mm</quantity>
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.
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<emphasis id="26EEEACAFFAFB95BFBC3FD73A6C5FD31" bold="true" box="[1091,1155,697,723]" pageId="2" pageNumber="334">Head</emphasis>
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. Frons yellow, brownish on dorsal half and black ocellar triangle (
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<figureCitation id="8CA12A5DFFAFB95BFDE8FD17A0E8FD15" box="[616,686,733,759]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 2</figureCitation>
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). Face and gena gray pruinose (
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<figureCitation id="8CA12A5DFFAFB95BFB96FD17A63AFD15" box="[1046,1148,733,759]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 2, 3</figureCitation>
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). Frons with five pairs of setae (
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<figureCitation id="8CA12A5DFFAFB95BFF5DFCCAA359FCF8" box="[221,287,768,794]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 3</figureCitation>
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, arrows). Postgena and postcranium margin yellow (
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<figureCitation id="8CA12A5DFFAFB95BFCE5FCCAA1E1FCF8" box="[869,935,768,794]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 3</figureCitation>
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), with postcranium otherwise black.Antenna yellow with distal margin faintly infuscated (
|
||
<figureCitation id="8CA12A5DFFAFB95BFD01FCEEA0A3FCDC" box="[641,741,804,830]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 1–3</figureCitation>
|
||
).
|
||
<emphasis id="26EEEACAFFAFB95BFD77FCEEA116FCDC" bold="true" box="[759,848,804,830]" pageId="2" pageNumber="334">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="8CA12A5DFFAFB95BFCDCFCEEA1F8FCDC" box="[860,958,804,830]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 1, 4</figureCitation>
|
||
). Scutal setae yellow to light brown. Notum black with postpronotum lobe light yellow and brown notopleuron basally light yellow. Acrostichals multiseriate, posterior pair slightly stouter. Scutellum and laterotergite black. Mediotergite yellow (
|
||
<figureCitation id="8CA12A5DFFAFB95BFBDBFCA0A6E7FC67" box="[1115,1185,874,901]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 4</figureCitation>
|
||
). Mesopleuron yellow with anepisternum black (
|
||
<figureCitation id="8CA12A5DFFAFB95BFE39FC44A3BCFC4A" box="[441,506,910,936]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 1</figureCitation>
|
||
). Legs light yellow with coxae and trochanters white, hind tibia with black median band (
|
||
<figureCitation id="8CA12A5DFFAFB95BFF5DFC7BA364FC2E" box="[221,290,945,972]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 4</figureCitation>
|
||
). Wing (
|
||
<figureCitation id="8CA12A5DFFAFB95BFE01FC7BA385FC2E" box="[385,451,945,972]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 5</figureCitation>
|
||
) light yellow basally, dusky from level of vein r-m to apex with a clear spot after vein at dm-m level, and alula brown. Setae along a basal section of Costa lighter in color. Length of cross-vein r-m shorter than the length of cross-vein dm-m.
|
||
<emphasis id="26EEEACAFFAFB95BFDACFC32A0E7FBF0" bold="true" box="[556,673,1016,1042]" pageId="2" pageNumber="334">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="8CA12A5DFFAFB95BFD2FFC32A155FBF1" box="[687,787,1016,1043]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 1, 4</figureCitation>
|
||
) tergites 1 and 2 yellow with one pair of dark lateral spots on tergite 2; tergite 3 to terminalia black. Sternites 1–3 yellow basally turning darker posteriorly.
|
||
<emphasis id="26EEEACAFFAFB95BFA92FBD6A7D3FBD4" bold="true" box="[1298,1429,1052,1078]" pageId="2" pageNumber="334">Terminalia</emphasis>
|
||
. Annulus black, well developed (
|
||
<figureCitation id="8CA12A5DFFAFB95BFD92FBF5A01CFBBB" box="[530,602,1087,1113]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 6</figureCitation>
|
||
). Epandrium 2X longer than high in lateral view (
|
||
<figureCitation id="8CA12A5DFFAFB95BFB30FBF5A6BEFBBB" box="[1200,1272,1087,1113]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 6</figureCitation>
|
||
), C-shaped in dorsal view (
|
||
<figureCitation id="8CA12A5DFFAFB95BFEADFBA9A332FB9F" box="[301,372,1123,1149]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 7</figureCitation>
|
||
). Cerci and surstylus yellowish, distinctly setose. Surstyli somewhat symmetrical with small tubercles apically on the inner face; in dorsal view, slightly inward curved (
|
||
<figureCitation id="8CA12A5DFFAFB95BFC7AFB4CA607FB42" box="[1018,1089,1158,1184]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 7</figureCitation>
|
||
). Cerci fused, shorter than the surstylus, shallow emarginated apically, somewhat heart-shaped, with a pair of longer distal paramedian setae (
|
||
<figureCitation id="8CA12A5DFFAFB95BFAEFFB60A2E1FB05" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 7</figureCitation>
|
||
). Subepandrial sclerite wide with short bacilliform sclerite (
|
||
<figureCitation id="8CA12A5DFFAFB95BFCC7FB07A1CDFB05" box="[839,907,1229,1255]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 8</figureCitation>
|
||
). Hypandrial arm (
|
||
<figureCitation id="8CA12A5DFFAFB95BFBDFFB07A694FB05" box="[1119,1234,1229,1255]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 9–10</figureCitation>
|
||
) upward directed, fused dorsally, articulated with wide concave subepandrial sclerite. Hypandrium with arm loosely attached (
|
||
<figureCitation id="8CA12A5DFFAFB95BFAEFFB3BA2F2FACC" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 10</figureCitation>
|
||
) fused to pregonite. Pregonite with three slender ventrolateral setae. Postgonite, epiphallus and lateral lobe of distiphallus absent. Phallapodeme is somewhat y-shaped, half-length of hypandrium length. Ejaculatory apodeme (
|
||
<figureCitation id="8CA12A5DFFAFB95BFF1FFA91A2B7FA97" box="[159,241,1371,1397]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 10</figureCitation>
|
||
) caliciform at distal end. Phallus (
|
||
<figureCitation id="8CA12A5DFFAFB95BFDF3FA91A0A0FA97" box="[627,742,1371,1397]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Figs 9–10</figureCitation>
|
||
) is somewhat cylindrical and membranous, with well-defined basiphallus and distiphallus, the latter slightly more sclerotized and irregularly shaped.
|
||
</paragraph>
|
||
<paragraph id="142536D8FFAFB95BFF47FA68A3E4FA5E" blockId="2.[151,1437,591,2038]" box="[199,418,1442,1469]" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFF47FA68A358FA5E" bold="true" box="[199,286,1442,1468]" pageId="2" pageNumber="334">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5C806553FFAFB95BFF47FA0FA3D3F98F" pageId="2" pageNumber="334" type="materials_examined">
|
||
<paragraph id="142536D8FFAFB95BFF47FA0FA12EF9C5" blockId="2.[151,1437,591,2038]" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFF47FA0FA3E9FA3D" bold="true" box="[199,431,1477,1503]" pageId="2" pageNumber="334">Material examined</emphasis>
|
||
.
|
||
<collectingCountry id="6C8D7648FFAFB95BFE42FA0FA04EFA3D" box="[450,520,1477,1503]" name="Brazil" pageId="2" pageNumber="334">Brasil</collectingCountry>
|
||
, PE[=
|
||
<collectingRegion id="D65EF83AFFAFB95BFDD1FA0FA0A3FA3D" box="[593,741,1477,1503]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="334">Pernambuco</collectingRegion>
|
||
], Fernando de Noronha, Tr. [=trail] Golfinhos,
|
||
<geoCoordinate id="71AE501FFFAFB95BFA98FA0FA7CDFA02" box="[1304,1419,1477,1504]" degrees="3" direction="south" minutes="51" orientation="latitude" pageId="2" pageNumber="334" precision="15" seconds="17" value="-3.854722">3°51’17”S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="71AE501FFFAFB95BFF17FA23A360F9E1" box="[151,294,1513,1539]" degrees="32" direction="west" minutes="26" orientation="longitude" pageId="2" pageNumber="334" precision="15" seconds="26" value="-32.440556">32°26’26”W</geoCoordinate>
|
||
/
|
||
<date id="60241018FFAFB95BFEB9FA23A3B2F9E1" box="[313,500,1513,1539]" pageId="2" pageNumber="334" value="2019-06-23" valueMax="2019-07-07" valueMin="2019-06-23">23.vi–7.vii.2019</date>
|
||
, Malaise Gd [=Grande; =6 meters Malaise trap], J.A. Rafael, F. Limeira-de-Oliveira, L.C. Castro /
|
||
<typeStatus id="CB21887AFFAFB95BFEAFF9C6A3D1F9C4" box="[303,407,1548,1574]" pageId="2" pageNumber="334" type="holotype">Holotype</typeStatus>
|
||
♂
|
||
<taxonomicName id="D39A4D5BFFAFB95BFE3AF9C6A0D9F9C4" authorityName="Rafael & Marques & Limeira-De-Oliveira" authorityYear="2023" box="[442,671,1548,1574]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="fumialula">
|
||
<emphasis id="26EEEACAFFAFB95BFE3AF9C6A0D9F9C4" box="[442,671,1548,1574]" italics="true" pageId="2" pageNumber="334">Czernyola fumialula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="8CA12A5DFFAFB95BFD2FF9C6A0B8F9C5" box="[687,766,1548,1575]" captionStart="FIGURES 1–11" captionStartId="3.[151,264,1649,1673]" captionTargetBox="[173,1421,190,1619]" captionTargetId="figure-22@3.[160,1427,181,1625]" captionTargetPageId="3" captionText="FIGURES 1–11. Czernyola fumialula sp. nov. Holotype ♂. 1, habitus, lateral view; 2, head, frontal view; 3, head, lateral view (black arrow indicating fronto-orbital setae); 4, habitus, dorsal view; 5, wing; 6, terminalia, lateral view; 7, terminalia, dorsal view; 8, detached left surstylus and subepandrial sclerite; 9, hypandrium and phallus, posterior view; 10, hypandrium and phallus, lateral view (ejaculatory apodeme detached); 11, holotype labels. Abbreviations: cerc = cercus; distph = distiphallus; ej apod = ejaculatory apodeme; epand = epandrium; hypd arm = hypandrial arm; pregnt = pregonite; subepand scl = subepandrial sclerite; sur = surstylus. Veins: dm-m = discal medial crossvein; r-m = radial-medial crossvein." figureDoi="http://doi.org/10.5281/zenodo.8430674" httpUri="https://zenodo.org/record/8430674/files/figure.png" pageId="2" pageNumber="334">Fig. 11</figureCitation>
|
||
) (
|
||
<collectionCode id="728BAE1DFFAFB95BFC96F9C7A11AF9C5" box="[790,860,1549,1575]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="2" pageNumber="334">INPA</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="142536D8FFAFB95BFF47F9FAA3D3F98F" blockId="2.[151,1437,591,2038]" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFF47F9FAA3F5F9A8" bold="true" box="[199,435,1584,1610]" pageId="2" pageNumber="334">
|
||
<typeStatus id="CB21887AFFAFB95BFF47F9FAA372F9A8" box="[199,308,1584,1610]" pageId="2" pageNumber="334" type="holotype">Holotype</typeStatus>
|
||
condition.
|
||
</emphasis>
|
||
Good, pinned, and mounted on card point. Left wing mounted in a microslide; abdomen dissected in microvial.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5C806553FFAFB95BFF47F9BDA354F956" pageId="2" pageNumber="334" type="etymology">
|
||
<paragraph id="142536D8FFAFB95BFF47F9BDA354F956" blockId="2.[151,1437,591,2038]" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFF47F9BDA30EF973" bold="true" box="[199,328,1655,1681]" pageId="2" pageNumber="334">Etymology</emphasis>
|
||
. From the Latin
|
||
<emphasis id="26EEEACAFFAFB95BFD92F9BDA013F973" box="[530,597,1655,1681]" italics="true" pageId="2" pageNumber="334">fumus</emphasis>
|
||
= smoky, infuscated, and
|
||
<emphasis id="26EEEACAFFAFB95BFC0AF9BDA182F973" box="[906,964,1655,1681]" italics="true" pageId="2" pageNumber="334">alula</emphasis>
|
||
= alula, which refers to the alula brown infuscated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5C806553FFAFB95DFF47F974A143FE6B" lastPageId="4" lastPageNumber="336" pageId="2" pageNumber="334" type="discussion">
|
||
<paragraph id="142536D8FFAFB95DFF47F974A143FE6B" blockId="2.[151,1437,591,2038]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="336" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFF47F974A372F93A" bold="true" box="[199,308,1726,1752]" pageId="2" pageNumber="334">Remarks</emphasis>
|
||
. Characteristic of other
|
||
<taxonomicName id="D39A4D5BFFAFB95BFDCEF974A087F93A" authorityName="Berri" authorityYear="1907" box="[590,705,1726,1752]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="26EEEACAFFAFB95BFDCEF974A087F93A" box="[590,705,1726,1752]" italics="true" pageId="2" pageNumber="334">Czernyola</emphasis>
|
||
</taxonomicName>
|
||
, the mid tibia has two dorsal preapical setae and the hind tibia has one; the ejaculatory apodeme is wide and flared at one end; the distiphallus is large, membranous and sac-like. Characteristic of the
|
||
<taxonomicName id="D39A4D5BFFAFB95BFE05F8CFA031F8FC" baseAuthorityName="Czerny" baseAuthorityYear="1903" box="[389,631,1797,1823]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="transversa">
|
||
<emphasis id="26EEEACAFFAFB95BFE05F8CFA031F8FC" box="[389,631,1797,1823]" italics="true" pageId="2" pageNumber="334">Czernyola transversa</emphasis>
|
||
</taxonomicName>
|
||
group, this species has five subequal fronto-orbital setae, of which the third is proclinate, the base of the costa has paler setae, and there is one pair of acrostichal setae. The male specimens from Fernando de Noronha run to couplet
|
||
<quantity id="D3629B3DFFAFB95BFDEEF886A0E0F884" box="[622,678,1868,1894]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="334" unit="in" value="10.0">10 in</quantity>
|
||
the key to
|
||
<taxonomicName id="D39A4D5BFFAFB95BFCA0F886A1ACF884" box="[800,1002,1868,1894]" pageId="2" pageNumber="334">
|
||
<emphasis id="26EEEACAFFAFB95BFCA0F886A1D5F884" box="[800,915,1868,1894]" italics="true" pageId="2" pageNumber="334">Czernyola</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="700B4B29FFAFB95BFB8DF886A70BF887" author="Lonsdale, O. & Marshall, S. A." box="[1037,1357,1867,1894]" pageId="2" pageNumber="334" pagination="1 - 101" refId="ref11608" refString="Lonsdale, O. & Marshall, S. A. (2006) Revision of the New World Craspedochaeta Czerny. Zootaxa, 1291 (1), 1 - 101. https: // doi. org / 10.11646 / zootaxa. 1291.1.1" type="journal article" year="2006">Lonsdale & Marshall (2006)</bibRefCitation>
|
||
, which includes
|
||
<taxonomicName id="D39A4D5BFFAFB95BFF7FF8A5A0C6F86B" authority="(Hennig, 1938)" baseAuthorityName="Hennig" baseAuthorityYear="1938" box="[255,640,1903,1929]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="26EEEACAFFAFB95BFF7FF8A5A381F86B" box="[255,455,1903,1929]" italics="true" pageId="2" pageNumber="334">Czernyola ornata</emphasis>
|
||
(Hennig, 1938)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D39A4D5BFFAFB95BFD3DF8BAA6D3F86B" authority="(Melander & Argo, 1924)" baseAuthorityName="Melander & Argo" baseAuthorityYear="1924" box="[701,1173,1903,1930]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="claripennis">
|
||
<emphasis id="26EEEACAFFAFB95BFD3DF8BAA119F86B" box="[701,863,1903,1930]" italics="true" pageId="2" pageNumber="334">C. claripennis</emphasis>
|
||
(Melander & Argo, 1924)
|
||
</taxonomicName>
|
||
. It fits most
|
||
<taxonomicName id="D39A4D5BFFAFB95BFAAEF8BAA7DBF86B" baseAuthorityName="Hennig" baseAuthorityYear="1938" box="[1326,1437,1904,1930]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="26EEEACAFFAFB95BFAAEF8BAA7DBF86B" box="[1326,1437,1904,1930]" italics="true" pageId="2" pageNumber="334">C. ornata</emphasis>
|
||
</taxonomicName>
|
||
characters: hind tibia with a black median band, anepisternum black, and surstylus produced anteriorly. It differs from
|
||
<taxonomicName id="D39A4D5BFFAFB95BFF54F872A306F833" baseAuthorityName="Hennig" baseAuthorityYear="1938" box="[212,320,1976,2002]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="26EEEACAFFAFB95BFF54F872A306F833" box="[212,320,1976,2002]" italics="true" pageId="2" pageNumber="334">C. ornata</emphasis>
|
||
</taxonomicName>
|
||
by having the postcranium black (versus ventral margin yellow in
|
||
<taxonomicName id="D39A4D5BFFAFB95BFBAAF872A6D3F833" baseAuthorityName="Hennig" baseAuthorityYear="1938" box="[1066,1173,1976,2002]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="334" phylum="Arthropoda" rank="species" species="ornata">
|
||
<emphasis id="26EEEACAFFAFB95BFBAAF872A6D3F833" box="[1066,1173,1976,2002]" italics="true" pageId="2" pageNumber="334">C. ornata</emphasis>
|
||
</taxonomicName>
|
||
); the scutum is entirely black (versus scutum almost always with one pair of yellow spots on the anterolateral margin of the notopleuron); the mediotergite is yellow (versus black); the anepisternum is black (versus yellow); the meron is yellow (versus dark brown to black); the wing is light yellow basally and dusky from the level of vein r-m (versus wing dusky, more heavily infuscated within first radial cell); the terminalia are black (versus light yellow); and the pregonite has three distinct ventrolateral setae on each side (versus one large distal seta and several medial setulae).
|
||
<taxonomicName id="D39A4D5BFFA9B95DFAA9FEC9A344FEA3" authority="Rafael & Marques & Limeira-De-Oliveira, 2023" authorityName="Rafael & Marques & Limeira-De-Oliveira" authorityYear="2023" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="336" phylum="Arthropoda" rank="species" species="fumialula" status="sp. nov.">
|
||
<emphasis id="26EEEACAFFA9B95DFAA9FEC9A344FEA3" italics="true" pageId="4" pageNumber="336">Czernyola fumialula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="26EEEACAFFA9B95DFE88FEEDA327FEA3" bold="true" box="[264,353,295,321]" pageId="4" pageNumber="336">
|
||
<taxonomicNameLabel id="3DDD57B1FFA9B95DFE88FEEDA327FEA3" box="[264,353,295,321]" pageId="4" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="D39A4D5BFFA9B95DFE71FEEDA0C8FEA3" baseAuthorityName="Melander & Argo" baseAuthorityYear="1924" box="[497,654,295,321]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="336" phylum="Arthropoda" rank="species" species="claripennis">
|
||
<emphasis id="26EEEACAFFA9B95DFE71FEEDA0C8FEA3" box="[497,654,295,321]" italics="true" pageId="4" pageNumber="336">C. claripennis</emphasis>
|
||
</taxonomicName>
|
||
by having the notum black with yellow spots (versus notum brown, no yellow spots in
|
||
<taxonomicName id="D39A4D5BFFA9B95DFEC6FE81A3A2FE87" baseAuthorityName="Melander & Argo" baseAuthorityYear="1924" box="[326,484,331,357]" class="Insecta" family="Clusiidae" genus="Czernyola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="336" phylum="Arthropoda" rank="species" species="claripennis">
|
||
<emphasis id="26EEEACAFFA9B95DFEC6FE81A3A2FE87" box="[326,484,331,357]" italics="true" pageId="4" pageNumber="336">C. claripennis</emphasis>
|
||
</taxonomicName>
|
||
); the anepisternum and laterotergite are black (versus yellow); and the wing is almost entirely brown infuscate (versus wing entirely hyaline).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |