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<mods:title id="9E14597DDA5B53A8F4A90D33537FB9EB">A cool-water bryozoan association from the La Pola Formation (Sandbian, Ordovician) of Argentine Precordillera</mods:title>
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<treatment id="865C87FDFFABFFEB396BF9634D52FCDF" ID-DOI="http://doi.org/10.5281/zenodo.6723053" ID-GBIF-Taxon="196202601" ID-Zenodo-Dep="6723053" LSID="urn:lsid:plazi:treatment:865C87FDFFABFFEB396BF9634D52FCDF" httpUri="http://treatment.plazi.org/id/865C87FDFFABFFEB396BF9634D52FCDF" lastPageId="11" lastPageNumber="572" pageId="9" pageNumber="570">
<subSubSection id="46EF6560FFABFFE9396BF9634B2FF93D" box="[866,1402,1718,1746]" pageId="9" pageNumber="570" type="nomenclature">
<paragraph id="0E4A36EBFFABFFE9396BF9634B2FF93D" blockId="9.[866,1402,1718,1777]" box="[866,1402,1718,1746]" pageId="9" pageNumber="570">
<taxonomicName id="C9F54D68FFABFFE9396BF9634B2FF93D" ID-CoL="868BX" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" box="[866,1402,1718,1746]" class="Stenolaemata" family="Monticuliporidae" genus="Monticulipora" higherTaxonomySource="CoL,GBIF" isUncertain="true" kingdom="Animalia" order="Trepostomatida" pageId="9" pageNumber="570" phylum="Bryozoa" rank="species" species="mammulata">
<emphasis id="3C81EAF9FFABFFE9396BF9634A57F93E" bold="true" box="[866,1026,1718,1745]" italics="true" pageId="9" pageNumber="570">Monticulipora</emphasis>
aff.
<emphasis id="3C81EAF9FFABFFE93E3DF9634AEEF93E" bold="true" box="[1076,1211,1718,1745]" italics="true" pageId="9" pageNumber="570">mammulata</emphasis>
dOrbigny, 1850
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="46EF6560FFABFFE939EFF9024AA3F91E" box="[998,1270,1751,1777]" pageId="9" pageNumber="570" type="description">
<paragraph id="0E4A36EBFFABFFE939EFF9024AA3F91E" blockId="9.[866,1402,1718,1777]" box="[998,1270,1751,1777]" pageId="9" pageNumber="570">
(
<figureCitation id="96CE2A6EFFABFFE939E6F9024A7BF91E" box="[1007,1070,1751,1777]" captionStart="FIG" captionStartId="12.[132,143,1940,1957]" captionTargetBox="[132,1455,216,1879]" captionTargetId="figure-25@12.[699,1268,769,1325]" captionTargetPageId="12" captionText="FIG. 6. — A, Dianulites rocklandensis Wilson,1921, tangential section showing autozooecial apertures, CEGH-UNC 27503 a; B-D, Monticulipora aff. mammulata dOrbigny,1850,CEGH-UNC 27516 d; B, C, tangential thin section showing autozooecial apertures;D, longitudinal thin section showing autozooecial chambers with cystiphragms; E-G. Orbignyella multitabulata Coryell, 1921; E, F, longitudinal thin section showing autozooecia with diaphragms, CEGH-UNC 27505 a; G, tangential thin section showing autozooecial apertures, CEGH-UNC 27505 b. Scale bars: A, C, 0.2 mm; B, G, 0.5 mm; D, F, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6725942" httpUri="https://zenodo.org/record/6725942/files/figure.png" pageId="9" pageNumber="570">Fig. 6</figureCitation>
B-D; Appendix 1)
</paragraph>
</subSubSection>
<subSubSection id="46EF6560FFABFFE93924F8DB4AEDF8AF" pageId="9" pageNumber="570" type="reference_group">
<paragraph id="0E4A36EBFFABFFE93924F8DB4AEDF8AF" blockId="9.[811,1456,1806,1856]" pageId="9" pageNumber="570">
For full synonymy of the species
<taxonomicName id="C9F54D68FFABFFE93E5BF8DB4D34F8AF" ID-CoL="868BX" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" class="Stenolaemata" family="Monticuliporidae" genus="Monticulipora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Trepostomatida" pageId="9" pageNumber="570" phylum="Bryozoa" rank="species" species="mammulata">
<emphasis id="3C81EAF9FFABFFE93E5BF8DB4B11F8C9" box="[1106,1348,1806,1830]" italics="true" pageId="9" pageNumber="570">Monticulipora mammulata</emphasis>
dOrbigny, 1850
</taxonomicName>
, see in
<bibRefCitation id="6A644B1AFFABFFE939A2F8FC4AE1F8AF" author="BROWN G. D. &amp; JR &amp; DALY E. J." box="[939,1204,1832,1856]" pageId="9" pageNumber="570" pagination="1 - 95" refId="ref19437" refString="BROWN G. D. JR &amp; DALY E. J. 1985. - Trepostome Bryozoa from the Dillsboro Formation (Cincinnatian Series) of Southeastern Indiana. Indiana Geological Survey Special Report 33: 1 - 95. https: // doi. org / 10.5967 / be 7 p-p 337" type="book chapter" year="1985">Brown &amp; Daly (1985: 68)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46EF6560FFABFFE93924F8824DADF866" pageId="9" pageNumber="570" type="materials_examined">
<paragraph id="0E4A36EBFFABFFE93924F8824DADF866" blockId="9.[813,1455,1878,1929]" pageId="9" pageNumber="570">
<materialsCitation id="BE9D3CB6FFABFFE93924F8824DADF866" ID-GBIF-Occurrence="3818790318" pageId="9" pageNumber="570" specimenCount="1">MATERIAL EXAMINED. — Single colony (three thin sections) CEGH-UNC 27516 a, b, d.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="46EF6560FFABFFE93924F84A4A88F804" pageId="9" pageNumber="570" type="distribution">
<paragraph id="0E4A36EBFFABFFE93924F84A4A88F804" blockId="9.[811,1456,1950,2027]" pageId="9" pageNumber="570">
OCCURRENCE. — La Pola Formation, Upper Ordovician, Sandbian; La Pola creek section near Albardon village,
<collectingRegion id="CC31F809FFABFFE93EE6F86C4BFFF83E" box="[1263,1450,1977,2001]" country="Argentina" name="San Juan" pageId="9" pageNumber="570">San Juan Province</collectingRegion>
,
<collectingCountry id="76E2767BFFABFFE93927F8064DC5F804" box="[814,912,2003,2027]" name="Argentina" pageId="9" pageNumber="570">Argentine</collectingCountry>
Precordillera, western
<collectingCountry id="76E2767BFFABFFE93E7DF8064A8FF804" box="[1140,1242,2003,2027]" name="Argentina" pageId="9" pageNumber="570">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="5A8A6663FFA8FFEA3A8DF8414C9EF806" ID-DOI="http://doi.org/10.5281/zenodo.6725938" ID-Zenodo-Dep="6725938" httpUri="https://zenodo.org/record/6725938/files/figure.png" pageId="10" pageNumber="571" startId="10.[132,143,1940,1957]" targetBox="[132,1455,215,1878]" targetPageId="10">
<paragraph id="0E4A36EBFFA8FFEA3A8DF8414C9EF806" blockId="10.[132,1456,1940,2025]" pageId="10" pageNumber="571">
FIG. 5. —
<emphasis id="3C81EAF9FFA8FFEA3ADCF8414EA0F84A" bold="true" box="[213,245,1940,1957]" pageId="10" pageNumber="571">A-E</emphasis>
,
<taxonomicName id="C9F54D68FFA8FFEA3B08F8414F9CF84A" authority="Ernst &amp; Carrera, 2022" authorityName="Ernst &amp; Carrera" authorityYear="2022" box="[257,457,1940,1957]" class="Stenolaemata" family="Xenotrypidae" genus="Xenotrypa" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Cystoporida" pageId="10" pageNumber="571" phylum="Bryozoa" rank="species" species="argentinensis" status="sp. nov.">
<emphasis id="3C81EAF9FFA8FFEA3B08F8414F9CF84A" box="[257,457,1940,1957]" italics="true" pageId="10" pageNumber="571">Xenotrypa argentinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="27B25782FFA8FFEA3BC6F8414C57F84A" box="[463,514,1940,1957]" pageId="10" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
:
<emphasis id="3C81EAF9FFA8FFEA3800F8414C43F84A" bold="true" box="[521,534,1940,1957]" pageId="10" pageNumber="571">A</emphasis>
, longitudinal thin section, showing autozooecia with diaphragms and vesicles, holotype CEGH-UNC 27523 b;
<emphasis id="3C81EAF9FFA8FFEA3A8DF87E4EFFF853" bold="true" box="[132,170,1963,1980]" pageId="10" pageNumber="571">B, C</emphasis>
, tangential thin section showing autozooecial apertures, acanthostyles, and vesicles, holotype CEGH-UNC 27523 a;
<emphasis id="3C81EAF9FFA8FFEA3E74F87E4AF4F853" bold="true" box="[1149,1185,1963,1980]" pageId="10" pageNumber="571">D, E</emphasis>
, tangential thin section showing autozooecial apertures,acanthostyles,and vesicles, paratype CEGH-UNC 27521 a;
<emphasis id="3C81EAF9FFA8FFEA3933F8174D09F83C" bold="true" box="[826,860,1986,2003]" pageId="10" pageNumber="571">F, G</emphasis>
,
<taxonomicName id="C9F54D68FFA8FFEA396FF8174ACAF83C" authority="Wilson, 1921" authorityName="Wilson" authorityYear="1921" box="[870,1183,1986,2003]" class="Stenolaemata" family="Dianulitidae" genus="Dianulites" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Trepostomatida" pageId="10" pageNumber="571" phylum="Bryozoa" rank="species" species="rocklandensis">
<emphasis id="3C81EAF9FFA8FFEA396FF8174A78F83C" box="[870,1069,1986,2003]" italics="true" pageId="10" pageNumber="571">Dianulites rocklandensis</emphasis>
<bibRefCitation id="6A644B1AFFA8FFEA3E38F8174ACAF83C" author="WILSON A. E." box="[1073,1183,1986,2003]" pageId="10" pageNumber="571" pagination="19 - 57" refId="ref23606" refString="WILSON A. E. 1921. - The range of certain Lower Ordovician faunas of the Ottawa Valley with descriptions of some new species. Canada Geological Survey Bulletin 33: 19 - 57. https: // doi. org / 10.4095 / 104998" type="journal article" year="1921">Wilson, 1921</bibRefCitation>
</taxonomicName>
, longitudinal thin section,CEGH- UNC 27503 c. Scale bars: A, B, E, G, 0.5 mm; C, 0.2 mm; D, F, 1 mm.
</paragraph>
</caption>
<subSubSection id="46EF6560FFA9FFEB3A8DFF0D4F40FDDF" pageId="11" pageNumber="572" type="description">
<paragraph id="0E4A36EBFFA9FFEB3A8DFF0D4F40FF1D" blockId="11.[130,776,216,497]" box="[132,277,216,242]" pageId="11" pageNumber="572">DESCRIPTION</paragraph>
<paragraph id="0E4A36EBFFA9FFEB3A8DFF224C94FE1F" blockId="11.[130,776,216,497]" pageId="11" pageNumber="572">
Submassive colony, multilayered, weakly differentiated into exo- and endozone. Autozooecia prismatic, bearing abundant cystiphragms.Autozooecial diaphragms common to abundant, mainly inclined or cystoid. Autozooecial apertures polygonal. Mesozooecia common, polygonal in tangential section, bearing closely spaced diaphragms, locally beaded. Zooecial walls laminated, serrated,
<quantity id="C90D9B0EFFA9FFEB3B53FE624FA0FE3E" box="[346,501,439,465]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="572" unit="mm" value="0.015" valueMax="0.02" valueMin="0.01">0.01-0.02 mm</quantity>
thick. Acanthostyles rare,
<quantity id="C90D9B0EFFA9FFEB3A8BFE034F69FE1E" box="[130,316,470,497]" metricMagnitude="-5" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="11" pageNumber="572" unit="mm" value="0.0325" valueMax="0.035" valueMin="0.03">
<geoCoordinate id="6BC1502CFFA9FFEB3A8BFE034E97FE1E" box="[130,194,470,497]" degrees="0.030" direction="north" orientation="latitude" pageId="11" pageNumber="572" precision="55" value="0.03">0.030</geoCoordinate>
<geoCoordinate id="6BC1502CFFA9FFEB3ACAFE034F5CFE1E" box="[195,265,470,497]" degrees="0.035" direction="west" orientation="longitude" pageId="11" pageNumber="572" precision="55" value="-0.035">-0.035</geoCoordinate>
mm
</quantity>
in diameter. Maculae not observed.
</paragraph>
<paragraph id="0E4A36EBFFA9FFEB3A8DFDC34F40FDDF" blockId="11.[130,777,534,816]" box="[132,277,534,560]" pageId="11" pageNumber="572">COMPARISON</paragraph>
</subSubSection>
<subSubSection id="46EF6560FFA9FFEB3A8DFDE34D52FCDF" pageId="11" pageNumber="572" type="discussion">
<paragraph id="0E4A36EBFFA9FFEB3A8DFDE34D52FCDF" blockId="11.[130,777,534,816]" pageId="11" pageNumber="572">
The present material is similar to
<taxonomicName id="C9F54D68FFA9FFEB3BE5FDE34F7BFD9E" authority="d'Orbigny, 1850" authorityName="d'Orbigny" authorityYear="1850" class="Stenolaemata" family="Monticuliporidae" genus="Monticulipora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Trepostomatida" pageId="11" pageNumber="572" phylum="Bryozoa" rank="species" species="mammulata">
<emphasis id="3C81EAF9FFA9FFEB3BE5FDE34D52FDBF" box="[492,775,566,592]" italics="true" pageId="11" pageNumber="572">Monticulipora mammulata</emphasis>
dOrbigny, 1850
</taxonomicName>
from the Upper Ordovician of North America. However, the typical monticules consisting of abundant mesozooecia, were not found in the present material.
<taxonomicName id="C9F54D68FFA9FFEB387CFD434FF4FD3F" authority="McKinney, 1971" authorityName="McKinney" authorityYear="1971" class="Stenolaemata" family="Monticuliporidae" genus="Monticulipora" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Trepostomatida" pageId="11" pageNumber="572" phylum="Bryozoa" rank="species" species="parallela">
<emphasis id="3C81EAF9FFA9FFEB387CFD434E8BFD20" italics="true" pageId="11" pageNumber="572">Monticulipora parallela</emphasis>
<bibRefCitation id="6A644B1AFFA9FFEB3AEEFD604FF4FD3F" author="MCKINNEY F. K." box="[231,417,693,720]" pageId="11" pageNumber="572" pagination="195 - 333" refId="ref21788" refString="MCKINNEY F. K. 1971. - Trepostomatous Ectoprocta (Bryozoa) from the lower Chickamauga Group (Middle Ordovician), Wills Valley, Alabama. Bulletins of American Paleontology 60: 195 - 333. https: // www. biodiversitylibrary. org / page / 28721254" type="journal article" year="1971">McKinney, 1971</bibRefCitation>
</taxonomicName>
from the Middle Ordovician of
<collectingCountry id="76E2767BFFA9FFEB3A8FFD004F43FD1F" box="[134,278,725,752]" name="United States of America" pageId="11" pageNumber="572">United States</collectingCountry>
(
<collectingRegion id="CC31F809FFA9FFEB3B2DFD004FD3FD1F" box="[292,390,725,752]" country="United States of America" name="Alabama" pageId="11" pageNumber="572">Alabama</collectingRegion>
) differs from the present material in encrusting colony and larger autozooecial apertures (average aperture width
<quantity id="C90D9B0EFFA9FFEB3B2DFCC04FD3FCDF" box="[292,390,789,816]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="572" unit="mm" value="0.2">0.20 mm</quantity>
vs
<quantity id="C90D9B0EFFA9FFEB3BACFCC04C52FCDF" box="[421,519,789,816]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="11" pageNumber="572" unit="mm" value="0.18">0.18 mm</quantity>
in the present material).
</paragraph>
</subSubSection>
</treatment>
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