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<document id="2E353135E00B8B5D37774E564F0A1FAB" ID-DOI="10.11646/zootaxa.5026.4.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5300736" ID-ZooBank="44594360-3408-4C4C-A4E6-1025F171A49F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630039285316" checkinUser="plazi" docAuthor="Michie, Laura A., Barnes, R. S. K., Clark, Paul F., Bennett, Wayne A. &amp; Cragg, Simon M." docDate="2021" docId="947287DFFFACFF837EAA132FFE9DF966" docLanguage="en" docName="zootaxa.5026.4.2.pdf" docOrigin="Zootaxa 5026 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gelasimus tetragonon" docType="treatment" docVersion="4" lastPageNumber="493" masterDocId="684BFFA7FFA1FF8E7E3D124CFF94FFB1" masterDocTitle="Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics" masterLastPageNumber="506" masterPageNumber="480" pageNumber="493" updateTime="1699139032236" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="F01468B479CFF7176C4A818D52E76DDA">Distinguishing ten sympatric species of fiddler crab (Decapoda: Ocypodidae) using a suite of phenotypic characteristics</mods:title>
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<mods:namePart id="EBE24C3AED78FCAC8FB20704B092AE0B">Michie, Laura A.</mods:namePart>
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<mods:namePart id="D7C1CEB01D4EC5DD477253140D5D7D51">Barnes, R. S. K.</mods:namePart>
<mods:affiliation id="B9A72E90C7C7232F9172B461F35BE41A">Department of Zoology and Conservation Research Institute, University of Cambridge, CB 2 3 EJ, UK.</mods:affiliation>
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<mods:namePart id="F8D195BFBEDA9505DFC09AB62B766A66">Clark, Paul F.</mods:namePart>
<mods:affiliation id="97FADF737883958B8696AFB4F9C770D9">Department of Life Sciences, The Natural History Museum, Cromwell Rad, London, SW 7 5 BD, UK.</mods:affiliation>
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<mods:namePart id="B815D1F5167F472F8A1A0DCBA3E772F8">Bennett, Wayne A.</mods:namePart>
<mods:affiliation id="BC4483DFC2BED944EABB97FB91CB310F">University of West Florida, 11000 University Parkway, Pensacola, US.</mods:affiliation>
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<mods:namePart id="7D05BD21BCCB0BD7A337C7627C0CCC2C">Cragg, Simon M.</mods:namePart>
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<mods:affiliation id="2EB237D685792F42E6CD52A6D7C79BB3">simon. cragg @ port. ac. uk; https: // orcid. org / 0000 - 0003 - 1082 - 7653</mods:affiliation>
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<mods:date id="E219536346D42E7BECF018AB70C6DD1C">2021</mods:date>
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<treatment id="947287DFFFACFF837EAA132FFE9DF966" ID-DOI="http://doi.org/10.5281/zenodo.5307764" ID-Zenodo-Dep="5307764" LSID="urn:lsid:plazi:treatment:947287DFFFACFF837EAA132FFE9DF966" httpUri="http://treatment.plazi.org/id/947287DFFFACFF837EAA132FFE9DF966" lastPageNumber="493" pageId="13" pageNumber="493">
<subSubSection id="54C16542FFACFF837EAA132FFDC8FECF" box="[151,604,355,382]" pageId="13" pageNumber="493" type="nomenclature">
<paragraph id="1C6436C9FFACFF837EAA132FFDC8FECF" blockId="13.[151,604,355,419]" box="[151,604,355,382]" pageId="13" pageNumber="493">
<heading id="472C81A5FFACFF837EAA132FFDC8FECF" bold="true" box="[151,604,355,382]" fontSize="11" level="1" pageId="13" pageNumber="493" reason="1">
<taxonomicName id="DBDB4D4AFFACFF837EAA132FFDC8FECF" ID-CoL="3FHCW" authority="(Herbst, 1790)" baseAuthorityName="Herbst" baseAuthorityYear="1790" box="[151,604,355,382]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="493" phylum="Arthropoda" rank="species" species="tetragonon">
<emphasis id="2EAFEADBFFACFF837EAA132FFDC8FECF" bold="true" box="[151,604,355,382]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EAA132FFE36FECF" bold="true" box="[151,418,355,382]" italics="true" pageId="13" pageNumber="493">Gelasimus tetragonon</emphasis>
(Herbst, 1790)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="54C16542FFACFF837EAA13C4FEBEFE12" box="[151,298,392,419]" pageId="13" pageNumber="493" type="description">
<paragraph id="1C6436C9FFACFF837EAA13C4FEBEFE12" blockId="13.[151,604,355,419]" box="[151,298,392,419]" pageId="13" pageNumber="493">
(
<figureCitation id="84E02A4CFFACFF837EA213C4FEB6FE12" box="[159,290,392,419]" captionStart="FIGURE 8" captionStartId="14.[151,250,1875,1899]" captionTargetBox="[163,1425,181,1850]" captionTargetId="figure-17@14.[163,1425,181,1852]" captionTargetPageId="14" captionText="FIGURE 8. Gelasimus tetragonon (Herbst, 1790): ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300752" httpUri="https://zenodo.org/record/5300752/files/figure.png" pageId="13" pageNumber="493">Figure 8aj</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFACFF837EAA139CFDE2FD83" pageId="13" pageNumber="493" type="materials_examined">
<paragraph id="1C6436C9FFACFF837EAA139CFDE2FD83" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EAA139CFEEBFE5B" bold="true" box="[151,383,464,490]" pageId="13" pageNumber="493">Material examined.</emphasis>
<materialsCitation id="ACB33C94FFACFF837FB8139CFE5FFDBF" collectingDate="2014-08-27" collectorName="L. Michie" country="Indonesia" latitude="-5.4969444" location="Kaledupa" longLatPrecision="21" longitude="123.75584" pageId="13" pageNumber="493" specimenCount="20" specimenCount-female="10" specimenCount-male="10">
<specimenCount id="0ADDFD40FFACFF837FB8139CFE28FE5A" box="[389,444,464,491]" pageId="13" pageNumber="493" type="female">10 ♀</specimenCount>
,
<specimenCount id="0ADDFD40FFACFF837FF4139CFD96FE5A" box="[457,514,464,491]" pageId="13" pageNumber="493" type="male">10 ♂</specimenCount>
,
<location id="19046012FFACFF837C30139CFDEFFE5B" LSID="urn:lsid:plazi:treatment:947287DFFFACFF837EAA132FFE9DF966:19046012FFACFF837C30139CFDEFFE5B" box="[525,635,464,490]" country="Indonesia" latitude="-5.4969444" longLatPrecision="21" longitude="123.75584" name="Kaledupa" pageId="13" pageNumber="493">Kaledupa</location>
,
<collectingCountry id="64CC7659FFACFF837CB8139CFD61FE5B" box="[645,757,464,490]" name="Indonesia" pageId="13" pageNumber="493">Indonesia</collectingCountry>
,
<geoCoordinate id="79EF500EFFACFF837CC3139CFC16FE5B" box="[766,898,464,491]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="13" pageNumber="493" precision="15" seconds="49" value="-5.4969444">5º 29 49S</geoCoordinate>
,
<geoCoordinate id="79EF500EFFACFF837DB0139CFBBBFE5B" box="[909,1071,464,491]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="13" pageNumber="493" precision="15" seconds="21" value="123.75584">123º 45 21E</geoCoordinate>
, mud flat southwest of boat jetty, coll.
<collectorName id="B12E531FFFACFF837EF313B9FEAAFDBF" box="[206,318,500,527]" pageId="13" pageNumber="493">L. Michie</collectorName>
,
<date id="68651009FFACFF837F7413B8FE5FFDBF" box="[329,459,500,526]" pageId="13" pageNumber="493" value="2014-08-27">
<collectingDate id="7821E9E1FFACFF837F7413B8FE5FFDBF" box="[329,459,500,526]" pageId="13" pageNumber="493" value="2014-08-27">27.08.2014</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="ACB33C94FFACFF837FE913B8FDE6FD83" collectingDate="2014-08-28" collectorName="L. Michie" country="Indonesia" latitude="-5.497222" location="Kaledupa" longLatPrecision="21" longitude="123.75584" pageId="13" pageNumber="493" specimenCount="20" specimenCount-female="10" specimenCount-male="10">
<specimenCount id="0ADDFD40FFACFF837FE913B8FD9FFDBE" box="[468,523,500,527]" pageId="13" pageNumber="493" type="female">10 ♀</specimenCount>
,
<specimenCount id="0ADDFD40FFACFF837C2513B8FDC5FDBE" box="[536,593,500,527]" pageId="13" pageNumber="493" type="male">10 ♂</specimenCount>
,
<location id="19046012FFACFF837C6113B8FD5EFDBF" LSID="urn:lsid:plazi:treatment:947287DFFFACFF837EAA132FFE9DF966:19046012FFACFF837C6113B8FD5EFDBF" box="[604,714,500,526]" country="Indonesia" latitude="-5.497222" longLatPrecision="21" longitude="123.75584" name="Kaledupa" pageId="13" pageNumber="493">Kaledupa</location>
,
<collectingCountry id="64CC7659FFACFF837CE913B8FCD0FDBF" box="[724,836,500,526]" name="Indonesia" pageId="13" pageNumber="493">Indonesia</collectingCountry>
,
<geoCoordinate id="79EF500EFFACFF837D7313B8FC46FDBE" box="[846,978,500,527]" degrees="5" direction="south" minutes="29" orientation="latitude" pageId="13" pageNumber="493" precision="15" seconds="50" value="-5.497222">5º 29 50S</geoCoordinate>
,
<geoCoordinate id="79EF500EFFACFF837DE013B8FBEBFDBF" box="[989,1151,500,527]" degrees="123" direction="east" minutes="45" orientation="longitude" pageId="13" pageNumber="493" precision="15" seconds="21" value="123.75584">123º 45 21E</geoCoordinate>
, mud flat between houses and boat jetty, coll.
<collectorName id="B12E531FFFACFF837F491055FE71FD83" box="[372,485,536,563]" pageId="13" pageNumber="493">L. Michie</collectorName>
,
<date id="68651009FFACFF837FCD1054FDE6FD83" box="[496,626,536,562]" pageId="13" pageNumber="493" value="2014-08-28">
<collectingDate id="7821E9E1FFACFF837FCD1054FDE6FD83" box="[496,626,536,562]" pageId="13" pageNumber="493" value="2014-08-28">28.08.2014</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFACFF837EFA1070FEBEFA23" pageId="13" pageNumber="493" type="diagnosis">
<paragraph id="1C6436C9FFACFF837EFA1070FB22FD73" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA1070FED6FDE7" bold="true" box="[199,322,572,598]" pageId="13" pageNumber="493">Diagnosis.</emphasis>
Carapace: broader than long; smooth; front narrow; anterolateral angles acute, slightly produced; anterolateral margins short, converging from angles; dorsolateral margins short, sometimes absent; carapace lateral margins slightly converging, practically straight posteriorly. Major cheliped: long, slender dactyl and pollex; pollex almost straight; dactyl moderately arched; dactyl and pollex with one or more sub-distal teeth.
</paragraph>
<paragraph id="1C6436C9FFACFF837EFA1080FEDDFC0E" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA1080FECCFD57" bold="true" box="[199,344,716,742]" pageId="13" pageNumber="493">Colouration</emphasis>
(
<figureCitation id="84E02A4CFFACFF837F5A1080FE66FD56" box="[359,498,716,743]" captionStart="FIGURE 8" captionStartId="14.[151,250,1875,1899]" captionTargetBox="[163,1425,181,1850]" captionTargetId="figure-17@14.[163,1425,181,1852]" captionTargetPageId="14" captionText="FIGURE 8. Gelasimus tetragonon (Herbst, 1790): ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300752" httpUri="https://zenodo.org/record/5300752/files/figure.png" pageId="13" pageNumber="493">Figure 8ad</figureCitation>
). Adult male. Carapace: blue with black mottled pattern varying in size and shape, anterior ventral surface of carapace generally black, dark brown or navy blue and ventral surface of third maxillipeds normally same blue as carapace, anterolateral angles can be pale straw coloured, occasionally pale anterolateral areas can be extensive. Major cheliped: orange, with dactyl often possessing a lighter tip, proximally pollex with darker hue of orange which extends onto part of manus, proximally manus with dark brown mottled pattern extending to carpus and merus. Minor cheliped &amp; ambulatory legs: black or dark brown, 4
<superScript id="EBAE9B81FFACFF837DD311CCFC6FFC3F" attach="left" box="[1006,1019,896,910]" fontSize="6" pageId="13" pageNumber="493">th</superScript>
pair of legs often with black and blue mottled pattern.
</paragraph>
<paragraph id="1C6436C9FFACFF837EFA1184FD87FBB7" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">Adult female. Carapace: similar to male, often with a darker hue of blue. Chelae &amp; ambulatory legs: orange, often have blue dactyl and pollex.</paragraph>
<paragraph id="1C6436C9FFACFF837EFA165CFE6FFBC3" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">Juvenile male. Carapace: mottled black and white pattern with colours developing with maturity. Major cheliped: similar to adults but with white distal end to dactyl and pollex. Minor cheliped: blue dactyl, pollex and manus. Ambulatory legs: black.</paragraph>
<paragraph id="1C6436C9FFACFF837EFA1630FCA7FB27" blockId="13.[151,1437,464,1751]" box="[199,819,1148,1175]" pageId="13" pageNumber="493">Juvenile female: same colour pattern as juvenile males.</paragraph>
<paragraph id="1C6436C9FFACFF837EFA16ECFD8AFAB3" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA16ECFE5AFB0B" bold="true" box="[199,462,1184,1210]" pageId="13" pageNumber="493">Gonopod morphology</emphasis>
(
<figureCitation id="84E02A4CFFACFF837FE116ECFDF2FB0A" box="[476,614,1184,1211]" captionStart="FIGURE 8" captionStartId="14.[151,250,1875,1899]" captionTargetBox="[163,1425,181,1850]" captionTargetId="figure-17@14.[163,1425,181,1852]" captionTargetPageId="14" captionText="FIGURE 8. Gelasimus tetragonon (Herbst, 1790): ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300752" httpUri="https://zenodo.org/record/5300752/files/figure.png" pageId="13" pageNumber="493">Figure 8eh</figureCitation>
). G1 strongly curved in dorso-lateral direction; distal end slightly curved laterally; anterior flange slightly narrower than posterior; pore of sperm channel large; distal protuberance moderately large, not reaching base of flanges.
</paragraph>
<paragraph id="1C6436C9FFACFF837EFA1740FEBEFA23" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA1740FEC2FA97" bold="true" box="[199,342,1292,1318]" pageId="13" pageNumber="493">Gastric mill</emphasis>
(
<figureCitation id="84E02A4CFFACFF837F581740FE71FA96" box="[357,485,1292,1319]" captionStart="FIGURE 8" captionStartId="14.[151,250,1875,1899]" captionTargetBox="[163,1425,181,1850]" captionTargetId="figure-17@14.[163,1425,181,1852]" captionTargetPageId="14" captionText="FIGURE 8. Gelasimus tetragonon (Herbst, 1790): ad Sulawesi Tenggara, Indonesia. Adult male colouration; a, large chela; b, carapace dorsal view. Adult female colouration; c, frontal view; d, carapace dorsal view. Photos credit L. Michie. Line drawing of left G1; e, lateral view; f, mesial view; CLSM images of apical part of G1; g, lateral view; h, mesial view; SEM images of male gastric mill; i, median tooth plate, ventral view; j, left lateral tooth plate, mesial view." figureDoi="http://doi.org/10.5281/zenodo.5300752" httpUri="https://zenodo.org/record/5300752/files/figure.png" pageId="13" pageNumber="493">Figure 8i, j</figureCitation>
). Urocardiac ossicles moderately complex, with 4 transverse ridges of median teeth, decreasing in size distally, different in shape, first two arched, 3 and 4 straighter, fourth large and attached to plate dorsally; stem with setae present on lateral margins of posterior stem region. Lateral tooth plate with 19 combshaped teeth.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFACFF837EFA17D0FBDBFA4F" pageId="13" pageNumber="493" type="biology_ecology">
<paragraph id="1C6436C9FFACFF837EFA17D0FBDBFA4F" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA17D0FEB5FA07" bold="true" box="[199,289,1436,1462]" pageId="13" pageNumber="493">Biology</emphasis>
.
<taxonomicName id="DBDB4D4AFFACFF837F0D17D0FDBDFA07" baseAuthorityName="Herbst" baseAuthorityYear="1790" box="[304,553,1436,1462]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="493" phylum="Arthropoda" rank="species" species="tetragonon">
<emphasis id="2EAFEADBFFACFF837F0D17D0FDBDFA07" box="[304,553,1436,1462]" italics="true" pageId="13" pageNumber="493">Gelasimus tetragonon</emphasis>
</taxonomicName>
is a medium-size species (largest male CL =
<quantity id="DB239B2CFFACFF837A1417D0FB1BFA06" box="[1065,1167,1436,1463]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="13" pageNumber="493" unit="mm" value="16.8">16.8 mm</quantity>
, CB =
<quantity id="DB239B2CFFACFF837AE217D0FAD2FA06" box="[1247,1350,1436,1463]" metricMagnitude="-2" metricUnit="m" metricValue="2.31" pageId="13" pageNumber="493" unit="mm" value="23.1">23.1 mm</quantity>
, largest female CL =
<quantity id="DB239B2CFFACFF837F12178CFE1CFA6A" box="[303,392,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="13" pageNumber="493" unit="mm" value="9.4">9.4 mm</quantity>
, CB =
<quantity id="DB239B2CFFACFF837FE6178CFDD0FA6A" box="[475,580,1472,1499]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="13" pageNumber="493" unit="mm" value="14.3">14.3 mm</quantity>
); present in mid and low intertidal zones, often associated with presence of coral rock where crabs were seen feeding on small patches of algae on rock surfaces.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFACFF837EFA1444FD0CF9F7" pageId="13" pageNumber="493" type="distribution">
<paragraph id="1C6436C9FFACFF837EFA1444FD0CF9F7" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA1444FECDF993" bold="true" box="[199,345,1544,1570]" pageId="13" pageNumber="493">Distribution</emphasis>
.
<collectingCountry id="64CC7659FFACFF837F541444FE6DF993" box="[361,505,1544,1570]" name="South Africa" pageId="13" pageNumber="493">South Africa</collectingCountry>
to
<collectingCountry id="64CC7659FFACFF837C221445FDDBF992" box="[543,591,1545,1571]" name="Iran" pageId="13" pageNumber="493">Iran</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837C611444FD71F993" box="[604,741,1544,1570]" name="Madagascar" pageId="13" pageNumber="493">Madagascar</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837CCC1444FCC3F993" box="[753,855,1544,1570]" name="Thailand" pageId="13" pageNumber="493">Thailand</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837D591444FC59F993" box="[868,973,1544,1570]" name="Malaysia" pageId="13" pageNumber="493">Malaysia</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837DE51444FBD6F993" box="[984,1090,1544,1570]" name="Australia" pageId="13" pageNumber="493">Australia</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837A731444FB2AF993" box="[1102,1214,1544,1570]" name="Indonesia" pageId="13" pageNumber="493">Indonesia</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837AF71444FADFF993" box="[1226,1355,1544,1570]" name="Philippines" pageId="13" pageNumber="493">Philippines</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837B651445FEB3F9F7" name="Papua New Guinea" pageId="13" pageNumber="493">Papua New Guinea</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837F0C1460FE12F9F7" box="[305,390,1580,1606]" name="Taiwan" pageId="13" pageNumber="493">Taiwan</collectingCountry>
,
<collectingCountry id="64CC7659FFACFF837FAC1460FD86F9F7" box="[401,530,1580,1606]" name="Micronesia" pageId="13" pageNumber="493">Micronesia</collectingCountry>
, Melanesia.
</paragraph>
</subSubSection>
<subSubSection id="54C16542FFACFF837EFA141CFE9DF966" pageId="13" pageNumber="493" type="discussion">
<paragraph id="1C6436C9FFACFF837EFA141CFE9DF966" blockId="13.[151,1437,464,1751]" pageId="13" pageNumber="493">
<emphasis id="2EAFEADBFFACFF837EFA141CFEAFF9DB" bold="true" box="[199,315,1616,1642]" pageId="13" pageNumber="493">Remarks.</emphasis>
<taxonomicName id="DBDB4D4AFFACFF837F79141CFDAAF9DB" baseAuthorityName="Herbst" baseAuthorityYear="1790" box="[324,574,1616,1642]" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="493" phylum="Arthropoda" rank="species" species="tetragonon">
<emphasis id="2EAFEADBFFACFF837F79141CFDAAF9DB" box="[324,574,1616,1642]" italics="true" pageId="13" pageNumber="493">Gelasimus tetragonon</emphasis>
</taxonomicName>
is distinguished by the strong curve of the G1, the long, slender dactyl and pollex and the distinct blue carapace and orange major cheliped. The specimens examined fit the original description and the moderately complex gastric mill conforms to the description by
<bibRefCitation id="784A4B38FFACFF837DE514D4FBF4F902" author="Shih, H. - T." box="[984,1120,1688,1715]" pageId="13" pageNumber="493" pagination="151 - 169" refId="ref14702" refString="Shih, H. - T. (2015) Uca (Xeruca), a new subgenus for the Taiwanese fiddler crab Uca formosensis Rathbun, 1921 (Crustacea: Decapoda: Ocypodidae), based on morphological and molecular evidence. Zootaxa, 3974 (2), 151 - 169. https: // doi. org / 10.11646 / zootaxa. 3974.2.1" type="journal article" year="2015">Shih (2015)</bibRefCitation>
and is similar to that of
<taxonomicName id="DBDB4D4AFFACFF837BBC14D4FE96F967" baseAuthorityName="H. -T. Shih, Naruse &amp; P. K. L. Ng" baseAuthorityYear="2010" class="Malacostraca" family="Ocypodidae" genus="Gelasimus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="493" phylum="Arthropoda" rank="species" species="jocelynae">
<emphasis id="2EAFEADBFFACFF837BBC14D4FE96F967" italics="true" pageId="13" pageNumber="493">G. jocelynae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>