446 lines
53 KiB
XML
446 lines
53 KiB
XML
<document id="F7E57088C10F705352B1ACCFE1B4A254" ID-CLB-Dataset="26040" ID-DOI="10.11646/zootaxa.4656.1.1" ID-GBIF-Dataset="a6cd54a1-9476-4ba4-9a29-6362590d223b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3366496" ID-ZooBank="1C0F29BB-0E02-48E4-810F-036F4F2730C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565678587482" checkinUser="plazi" docAuthor="Vilkamaa, Pekka, Halenius, Pentti & Ševčík, Jan" docDate="2019" docId="03F387CAEF2ADC4CFF1CC84EABE2B587" docLanguage="en" docName="zootaxa.4656.1.1.pdf" docOrigin="Zootaxa 4656 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoaerumnosa Rudzinski 2006" docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="FFCAFFB2EF2BDC4FFF8BC903A944B23D" masterDocTitle="Review of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae), with the description of twenty-four new species" masterLastPageNumber="42" masterPageNumber="1" pageNumber="2" updateTime="1698750780509" updateUser="ExternalLinkService">
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<mods:title id="63BA0F90E1C15ED8005C9B0C7DCB73EA">Review of Pseudoaerumnosa Rudzinski (Diptera, Sciaridae), with the description of twenty-four new species</mods:title>
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<mods:namePart id="799C21B2F1A24F3047D7A56CD3AEB7D8">Vilkamaa, Pekka</mods:namePart>
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<mods:namePart id="5294517002A8709DB1AB4D5500E094A1">Halenius, Pentti</mods:namePart>
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<mods:namePart id="26796EE630F4CCFC3E6F3EE96FAD311B">Ševčík, Jan</mods:namePart>
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<mods:typeOfResource id="2D09AA473DB2E72FF6575EF492419653">text</mods:typeOfResource>
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<mods:title id="808B72C27E5A2C2EA543C9CC3B946AD2">Zootaxa</mods:title>
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<mods:part id="DB618093B74992312CC54779398F4D2C">
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<mods:date id="D56B17F4BCBC50190EF45F8CDE15AF8D">2019</mods:date>
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<mods:number id="D6923050D00AD17B0289625B1F92C787">2019-08-13</mods:number>
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<mods:identifier id="CD7D3F1B087D04CADE86EC241671161E" type="DOI">10.11646/zootaxa.4656.1.1</mods:identifier>
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<treatment id="03F387CAEF2ADC4CFF1CC84EABE2B587" ID-GBIF-Taxon="159544498" LSID="urn:lsid:plazi:treatment:03F387CAEF2ADC4CFF1CC84EABE2B587" httpUri="http://treatment.plazi.org/id/03F387CAEF2ADC4CFF1CC84EABE2B587" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="C3406557EF2ADC4EFF1CC84EAB8AB35A" box="[151,718,333,360]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BE536DCEF2ADC4EFF1CC84EAB8AB35A" blockId="1.[151,718,333,360]" box="[151,718,333,360]" pageId="1" pageNumber="2">
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<heading id="D0AD81B0EF2ADC4EFF1CC84EAB8AB35A" bold="true" box="[151,718,333,360]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="B92EEACEEF2ADC4EFF1CC84EAB8AB35A" bold="true" box="[151,718,333,360]" pageId="1" pageNumber="2">
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The genus
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<taxonomicName id="4C5A4D5FEF2ADC4EFE95C84EAB8AB35A" ID-CoL="6Z6Q" ID-ENA="2058813" authority="Rudzinski, 2006" authorityName="Rudzinski" authorityYear="2006" box="[286,718,333,360]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B92EEACEEF2ADC4EFE95C84EA8B9B355" bold="true" box="[286,509,333,360]" italics="true" pageId="1" pageNumber="2">Pseudoaerumnosa</emphasis>
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<bibRefCitation id="EFCB4B2DEF2ADC4EFD8EC84EAB8AB35A" author="Rudzinski, H. - G." box="[517,718,333,359]" pageId="1" pageNumber="2" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski, 2006</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3406557EF2ADC4DFF1CC897A8E6B62E" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="description">
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<paragraph id="8BE536DCEF2ADC4EFF1CC897AD6AB392" blockId="1.[151,1437,404,683]" box="[151,1070,404,431]" pageId="1" pageNumber="2">
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<emphasis id="B92EEACEEF2ADC4EFF1CC897A801B392" bold="true" box="[151,325,404,431]" pageId="1" pageNumber="2">Redescription.</emphasis>
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Male. Small to medium-sized
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<taxonomicName id="4C5A4D5FEF2ADC4EFD15C896AA4DB392" authorityName="Billberg" authorityYear="1820" box="[670,777,405,431]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
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, wing length
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<quantity id="4CA29B39EF2ADC4EFC2BC897AD6EB392" box="[928,1066,404,431]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="3.5" metricValueMin="1.4" pageId="1" pageNumber="2" unit="mm" value="2.45" valueMax="3.5" valueMin="1.4">1.4–3.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BE536DCEF2ADC4EFF4CC8BAAB2BB096" blockId="1.[151,1437,404,683]" pageId="1" pageNumber="2">
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<emphasis id="B92EEACEEF2ADC4EFF4CC8BAA843B3EE" bold="true" box="[199,263,441,467]" pageId="1" pageNumber="2">Head</emphasis>
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. Brown, face concolorous with antenna. Eye bridge 1–4 facets wide. Face with 1–25 setae. Mouth parts strongly reduced, clypeus non-setose (
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<bibRefCitation id="EFCB4B2DEF2ADC4EFDCFC8DEABBEB3CA" author="Rudzinski, H. - G." box="[580,762,477,503]" pageId="1" pageNumber="2" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski 2006</bibRefCitation>
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: Figs. 62, 64). Antennal scape and pedicel normal. Antennal flagellum with 14 flagellomeres, body of flagellomeres asymmetrical, frontal part larger, with distinct apical margin, with dense, short and curved setosity, setae shorter than width of the flagellomeres; with some rod-like sensilla, neck of flagellomeres shorter than wide, body of 4th flagellomere 1.3–2.3x as long as wide in frontal view (
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<figureCitation id="13612A59EF2ADC4EFAD2CB4AA98AB0BA" captionStart="FIGURE 1" captionStartId="1.[151,250,1934,1959]" captionTargetBox="[186,1397,716,1901]" captionTargetId="figure@1.[173,1415,705,1915]" captionTargetPageId="1" captionText="FIGURE 1. A. Apical part of fore tibia, prolateral. B. 4 th antennal flagellomere, frontal. C. 4th antennal flagellomere, lateral. A: Pseudoaerumnosa obovata sp. n. (paratype from Ladan). B, C: P. clavidactyla sp. n. (paratypes from Danum Valley). Scale 0.10 mm." httpUri="https://zenodo.org/record/3366498/files/figure.png" pageId="1" pageNumber="2">Fig. 1 B, C</figureCitation>
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). Mouthparts reduced; maxillary palpus with one segment, with 1–6 pointed setae, with a dorsal patch of long sensilla (
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<bibRefCitation id="EFCB4B2DEF2ADC4EFF71CB92A8F4B096" author="Rudzinski, H. - G." box="[250,432,657,683]" pageId="1" pageNumber="2" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski 2006</bibRefCitation>
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: Fig. 62, 64, 65).
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</paragraph>
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<caption id="DF256654EF2ADC4EFF1CCE8DA9BDB5D3" httpUri="https://zenodo.org/record/3366498/files/figure.png" pageId="1" pageNumber="2" startId="1.[151,250,1934,1959]" targetBox="[186,1397,716,1901]" targetPageId="1">
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<paragraph id="8BE536DCEF2ADC4EFF1CCE8DA9BDB5D3" blockId="1.[151,1437,1932,2031]" pageId="1" pageNumber="2">
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<emphasis id="B92EEACEEF2ADC4EFF1CCE8DA877B59A" bold="true" box="[151,307,1934,1959]" pageId="1" pageNumber="2">FIGURE 1. A.</emphasis>
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Apical part of fore tibia, prolateral.
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<emphasis id="B92EEACEEF2ADC4EFD22CE8CAB84B59A" bold="true" box="[681,704,1935,1959]" pageId="1" pageNumber="2">B.</emphasis>
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4
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<superScript id="7C2F9B94EF2ADC4EFD5FCE8FAB9BB5A7" attach="left" box="[724,735,1932,1946]" fontSize="6" pageId="1" pageNumber="2">th</superScript>
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antennal flagellomere, frontal.
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<emphasis id="B92EEACEEF2ADC4EFBAFCE8DAD78B59A" bold="true" box="[1060,1084,1934,1959]" pageId="1" pageNumber="2">C.</emphasis>
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4th antennal flagellomere, lateral.
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<emphasis id="B92EEACEEF2ADC4EFF1CCEB1A9F6B5F6" bold="true" box="[151,178,1970,1995]" pageId="1" pageNumber="2">A:</emphasis>
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<taxonomicName id="4C5A4D5FEF2ADC4EFF33CEB1A88FB5F7" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[184,459,1970,1994]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="obovata" status="sp. nov.">
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<emphasis id="B92EEACEEF2ADC4EFF33CEB1A88FB5F7" box="[184,459,1970,1994]" italics="true" pageId="1" pageNumber="2">Pseudoaerumnosa obovata</emphasis>
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</taxonomicName>
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<emphasis id="B92EEACEEF2ADC4EFE5ACEB0AB4EB5F6" bold="true" box="[465,522,1971,1995]" pageId="1" pageNumber="2">
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<taxonomicNameLabel id="A21D57B5EF2ADC4EFE5ACEB0AB4EB5F6" box="[465,522,1971,1995]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(paratype from Ladan).
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<emphasis id="B92EEACEEF2ADC4EFC8ACEB0AA7DB5F6" bold="true" box="[769,825,1970,1995]" pageId="1" pageNumber="2">B, C:</emphasis>
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<taxonomicName id="4C5A4D5FEF2ADC4EFCB4CEB0AA90B5F7" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[831,980,1970,1994]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="clavidactyla" status="sp. nov.">
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<emphasis id="B92EEACEEF2ADC4EFCB4CEB0AA90B5F7" box="[831,980,1970,1994]" italics="true" pageId="1" pageNumber="2">P. clavidactyla</emphasis>
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</taxonomicName>
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<emphasis id="B92EEACEEF2ADC4EFC51CEB0AD57B5F6" bold="true" box="[986,1043,1971,1995]" pageId="1" pageNumber="2">
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<taxonomicNameLabel id="A21D57B5EF2ADC4EFC51CEB0AD57B5F6" box="[986,1043,1971,1995]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(paratypes from Danum Valley). Scale 0.10 mm.
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</paragraph>
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</caption>
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<paragraph id="8BE536DCEF29DC4DFF4CC99AAB2FB1D2" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="3">
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<emphasis id="B92EEACEEF29DC4DFF4CC99AA864B28E" bold="true" box="[199,288,153,179]" pageId="2" pageNumber="3">Thorax</emphasis>
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. Brown; setae dark. Notum with many short laterals, with robust dorsocentrals. Postpronotum with some setae, anterior pronotum with 1–11 setae, proepisternum with 1–8 setae. Scutellum with 2 or 4 moderately long and many short setae.
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<emphasis id="B92EEACEEF29DC4DFE40C9E3AB58B2C6" bold="true" box="[459,540,224,251]" pageId="2" pageNumber="3">Wing (</emphasis>
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<figureCitation id="13612A59EF29DC4DFD96C9E2AB20B2C6" box="[541,612,225,251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1593,1618]" captionTargetBox="[159,1427,425,1564]" captionTargetId="figure@3.[151,1436,417,1574]" captionTargetPageId="3" captionText="FIGURE 2. Wing, dorsal.A: Pseudoaerumnosa banari sp. n. (paratype). B: P. eminula sp. n. (paratype from Ulu Temburong). C: P. fragilis sp. n. (holotype). D: P. tenuidens sp. n. (holotype). E: P. consuota sp. n. (paratype). F: P. junciseta sp. n. (paratype). Scale 1.0 mm." httpUri="https://zenodo.org/record/3366500/files/figure.png" pageId="2" pageNumber="3">Figs 2</figureCitation>
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,
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<figureCitation id="13612A59EF29DC4DFDFBC9E2ABC4B2C6" box="[624,640,225,251]" captionStart="FIGURE 3" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[188,1390,219,1665]" captionTargetId="figure@4.[173,1415,181,1669]" captionTargetPageId="4" captionText="FIGURE 3. Part of hind margin of wing, dorsal. A: Pseudoaerumnosa fragilis sp. n. (holotype). B: P. tenuidens sp. n. (holotype). Scale 0.10 mm." httpUri="https://zenodo.org/record/3366502/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
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). Fumose. Normal, broad (width/length 0.45–0.60), anal lobe variable. Veins distinct, R1 longer than R, joining before, at or beyond base of M-fork. Wing membrane non-setose, hind margin with dorsal setae, in some species also with ventral setae. Veins C and R5 with dorsal and ventral setae, R1 with dorsal setae, with or without ventral setae, bM and stCu non-setose, r-m non-setose or (rarely) with 1–2 setae. c/w 0.55–0.90, R1/R 1.15–2.65. Haltere with short stem.
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<emphasis id="B92EEACEEF29DC4DFCA2C872AA25B3B6" bold="true" box="[809,865,369,395]" pageId="2" pageNumber="3">Legs</emphasis>
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. Yellow or pale brown, slender, setae dark. Tibial spurs 1:2:2. Fore tibial spur as long as or slightly longer than the subapical width of tibia. Left and right spurs of mid and hind tibia subequal. Tibiae without distinct spinose setae. Fore tibial organ unmodified, with a few slender setae only (
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<figureCitation id="13612A59EF29DC4DFE93C8DEA83DB3CA" box="[280,377,477,503]" captionStart="FIGURE 1" captionStartId="1.[151,250,1934,1959]" captionTargetBox="[186,1397,716,1901]" captionTargetId="figure@1.[173,1415,705,1915]" captionTargetPageId="1" captionText="FIGURE 1. A. Apical part of fore tibia, prolateral. B. 4 th antennal flagellomere, frontal. C. 4th antennal flagellomere, lateral. A: Pseudoaerumnosa obovata sp. n. (paratype from Ladan). B, C: P. clavidactyla sp. n. (paratypes from Danum Valley). Scale 0.10 mm." httpUri="https://zenodo.org/record/3366498/files/figure.png" pageId="2" pageNumber="3">Fig. 1 A</figureCitation>
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;
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<bibRefCitation id="EFCB4B2DEF29DC4DFE0EC8DEAB7EB3CA" author="Rudzinski, H. - G." box="[389,570,477,503]" pageId="2" pageNumber="3" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski 2006</bibRefCitation>
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: Fig. 66). Tarsal claws simple, without teeth.
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<emphasis id="B92EEACEEF29DC4DFBB7C8DFADF5B3CA" bold="true" box="[1084,1201,476,503]" pageId="2" pageNumber="3">Abdomen</emphasis>
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. Pale brown; plump, setae long, dark, and robust.
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<emphasis id="B92EEACEEF29DC4DFE53CB03AB30B026" bold="true" box="[472,628,512,539]" pageId="2" pageNumber="3">Hypopygium</emphasis>
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. Brown, like the abdomen; setae robust and dark. Gonocoxa longer than or as long as gonostylus.
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<emphasis id="B92EEACEEF29DC4DFE4BCB27AB00B002" bold="true" box="[448,580,548,575]" pageId="2" pageNumber="3">Gonocoxae</emphasis>
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basomedially fused, intergonocoxal area broad and short, without lobe(s), with or without microtrichia, medially with or without non-setose area (
|
||
<figureCitation id="13612A59EF29DC4DFC3DCB4AAABEB05E" box="[950,1018,585,611]" captionStart="FIGURE 4" captionStartId="6.[151,250,1098,1123]" captionTargetBox="[151,1435,181,1078]" captionTargetId="figure@6.[151,1435,181,1078]" captionTargetPageId="6" captionText="FIGURE 4. Intergonocoxal area, ventral. A: Pseudoaerumnosa banari sp. n. (holotype). B: P. consuota sp. n. (holotype). C: P. fragilis sp. n. (holotype). D: P. tenuidens sp. n. (paratype from Sabah, Mainline West). Scale 0.10 mm." httpUri="https://zenodo.org/record/3366504/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
), ventromedial margin distinct, semicircular or U-shaped. Gonocoxa with strongly sclerotized apical margin, with long setosity, setae slightly shorter at medial margin, apicomedial seta well developed or indistinguishable from surrounding setae, in most species microtrichosity restricted to basal and lateral parts.
|
||
<emphasis id="B92EEACEEF29DC4DFD53CBB7AA1AB0F2" bold="true" box="[728,862,692,719]" pageId="2" pageNumber="3">Gonostylus</emphasis>
|
||
ventrally and dorsally impressed or not impressed, with sclerotized basolateral apophysis, with a medial lobe, variably developed in the species, or medial side of gonostylus broadly lobe-like produced, the lobe or lobe-like medial side with 2 to 12 megasetae; megasetae slender, hyalinous and nearly straight or curved; with long apical tooth with a distinct basal part, both dark, apical tooth with small megaseta attached to its basoventral side; with rich setosity, usually with some narrow elongated setae on the apical and basal/basoventral side of the tooth, microtrichosity restricted to basolateral parts. Tegmen without basal sclerotization, laterally variably sclerotized, smoothly roundish or curved or acuminate, hyalinous apically, in some species microtrichose, with or without apical finger-like process; with tiny aedeagal teeth in groups, basal apodemes short. Aedeagal apodeme usually distinct.
|
||
</paragraph>
|
||
<paragraph id="8BE536DCEF29DC4DFF4CCAFAA8E6B62E" blockId="2.[151,1437,153,2015]" box="[199,418,1017,1043]" pageId="2" pageNumber="3">
|
||
<emphasis id="B92EEACEEF29DC4DFF4CCAFAA85AB62E" bold="true" box="[199,286,1017,1043]" pageId="2" pageNumber="3">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3406557EF29DC4DFF4CCD1FAAD9B60A" box="[199,925,1052,1079]" pageId="2" pageNumber="3" type="distribution">
|
||
<paragraph id="8BE536DCEF29DC4DFF4CCD1FAAD9B60A" blockId="2.[151,1437,153,2015]" box="[199,925,1052,1079]" pageId="2" pageNumber="3">
|
||
<emphasis id="B92EEACEEF29DC4DFF4CCD1FA825B60A" bold="true" box="[199,353,1052,1079]" pageId="2" pageNumber="3">Distribution.</emphasis>
|
||
<collectingCountry id="F34D764CEF29DC4DFEECCD1EA8B4B60A" box="[359,496,1053,1079]" name="Madagascar" pageId="2" pageNumber="3">Madagascar</collectingCountry>
|
||
, Oriental Region,
|
||
<collectingCountry id="F34D764CEF29DC4DFD36CD1EAADDB60A" box="[701,921,1053,1079]" name="Papua New Guinea" pageId="2" pageNumber="3">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3406557EF29DC4DFF4CCD43AB2EB572" pageId="2" pageNumber="3" type="discussion">
|
||
<paragraph id="8BE536DCEF29DC4DFF4CCD43AD42B46E" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="3">
|
||
<emphasis id="B92EEACEEF29DC4DFF4CCD43A808B666" bold="true" box="[199,332,1088,1115]" pageId="2" pageNumber="3">Discussion.</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFED8CD42AB66B666" authorityName="Rudzinski" authorityYear="2006" box="[339,546,1089,1115]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFED8CD42AB66B666" box="[339,546,1089,1115]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from other genera of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFB88CD42AD2FB666" authorityName="Billberg" authorityYear="1820" box="[1027,1131,1089,1115]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
by the following combination of characters: plump abdomen, maxillary palpus 1-segmented with a dorsal patch of sensilla, antennal flagellomeres asymmetrical with the frontal part larger (unique among the known
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFC59CD8AAD78B699" authorityName="Billberg" authorityYear="1820" box="[978,1084,1161,1188]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
) and with short curved setosity, the apical tooth of the gonostylus with broader basal part and a longer, narrower apical part with a short megaseta attached to the ventral side (such megasetae are also present in some species of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFBB1CDD2AC46B6D6" authority="Menzel" authorityName="Menzel" box="[1082,1282,1233,1259]" class="Insecta" family="Sciaridae" genus="Mohrigia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFBB1CDD2ADE0B6D6" box="[1082,1188,1233,1259]" italics="true" pageId="2" pageNumber="3">Mohrigia</emphasis>
|
||
Menzel
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="EFCB4B2DEF29DC4DFF1CCDF6A82CB72D" author="Rudzinski, H. - G." box="[151,360,1269,1296]" pageId="2" pageNumber="3" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski (2006))</bibRefCitation>
|
||
; a medial (not dorsal or ventral) lobe on the gonostylus (a medial lobe is present in some species of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFF6CCC1AAB13B70E" authority="Frey" authorityName="Frey" box="[231,599,1305,1331]" class="Insecta" family="Sciaridae" genus="Scatopsciara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Xenopygina">
|
||
<emphasis id="B92EEACEEF29DC4DFF6CCC1AA83DB70E" box="[231,377,1305,1331]" italics="true" pageId="2" pageNumber="3">Scatopsciara</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF29DC4DFE01CC19AB54B70E" box="[394,528,1306,1331]" italics="true" pageId="2" pageNumber="3">Xenopygina</emphasis>
|
||
) Frey
|
||
</taxonomicName>
|
||
); hyaline, at least slightly curved gonostylar megasetae (hyaline megasetae present in various genera); a broad hypopygium with a short and evenly broad intergonocoxal area formed by the united gonocoxae (found also in
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFD98CC62ABA0B741" authority="Winnertz" authorityName="Winnertz" authorityYear="1867" box="[531,740,1377,1404]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFD98CC62AB2BB746" box="[531,623,1377,1403]" italics="true" pageId="2" pageNumber="3">Cratyna</emphasis>
|
||
Winnertz
|
||
</taxonomicName>
|
||
); unmodified fore tibial organ (an unmodified tibial organ is also present in
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFEB5CC86AB43B7A2" authority="Haliday" authorityName="Haliday" authorityYear="1851" box="[318,519,1413,1439]" class="Insecta" family="Sciaridae" genus="Epidapus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFEB5CC86A8ECB7A2" box="[318,424,1413,1439]" italics="true" pageId="2" pageNumber="3">Epidapus</emphasis>
|
||
Haliday
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFDB7CC86AA03B79D" authority="Tuomikoski" authorityName="Tuomikoski" box="[572,839,1413,1440]" class="Insecta" family="Sciaridae" genus="Xylosciara" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFDB7CC86ABF1B7A2" box="[572,693,1413,1439]" italics="true" pageId="2" pageNumber="3">Xylosciara</emphasis>
|
||
Tuomikoski
|
||
</taxonomicName>
|
||
); tibiae without distinct spinose setae; a broad wing, a long wing vein R1 (always longer than R; a long R1 is also present in
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFC29CCAAAD2AB7FE" authority="Winnertz" authorityName="Winnertz" authorityYear="1867" box="[930,1134,1449,1475]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFC29CCAAAABAB7FE" box="[930,1022,1449,1475]" italics="true" pageId="2" pageNumber="3">Cratyna</emphasis>
|
||
Winnertz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFBF3CCAAACD8B7FE" authority="Enderlein" authorityName="Enderlein" authorityYear="1911" box="[1144,1436,1449,1475]" class="Insecta" family="Sciaridae" genus="Scythropochroa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFBF3CCAAAC6DB7FE" box="[1144,1321,1449,1475]" italics="true" pageId="2" pageNumber="3">Scythropochroa</emphasis>
|
||
Enderlein
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFF42CCCEAB41B7DA" authority="Frey" authorityName="Frey" box="[201,517,1485,1511]" class="Insecta" family="Dolichopodidae" genus="Trichosia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Mouffetina">
|
||
<emphasis id="B92EEACEEF29DC4DFF42CCCEA877B7DA" box="[201,307,1485,1511]" italics="true" pageId="2" pageNumber="3">Trichosia</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF29DC4DFECECCCEA884B7DA" box="[325,448,1485,1511]" italics="true" pageId="2" pageNumber="3">Mouffetina</emphasis>
|
||
) Frey
|
||
</taxonomicName>
|
||
and in various undescribed groups of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFC34CCCEAD68B7D5" authorityName="Billberg" authorityYear="1820" box="[959,1068,1485,1512]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFBCACCCEAC54B7DA" authorityName="Rudzinski" authorityYear="2006" box="[1089,1296,1485,1511]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFBCACCCEAC54B7DA" box="[1089,1296,1485,1511]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
also always has a long c/w ratio. Like the subgenera of
|
||
<emphasis id="B92EEACEEF29DC4DFDF0CCF2AAF7B436" box="[635,947,1521,1547]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFDF0CCF2AB9EB436" authorityName="Winnertz" authorityYear="1867" box="[635,730,1521,1547]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Cratyna</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFD6FCCF2AAF7B436" authorityName="Rudzinski" authorityYear="2006" box="[740,947,1521,1547]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Pseudoaerumnosa</taxonomicName>
|
||
</emphasis>
|
||
has setae on the postpronotum, although, if broken, their sockets may be difficult to detect in normal slide mounts. Furthermore, the colouration in most species is yellow or pale brown, contrasting strongly with the dark and robust setosity.
|
||
</paragraph>
|
||
<paragraph id="8BE536DCEF29DC4DFF4CCF5FAB2EB572" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="3">
|
||
<emphasis id="B92EEACEEF29DC4DFF4CCF5FABB0B44A" bold="true" box="[199,756,1628,1655]" pageId="2" pageNumber="3">
|
||
Phylogenetic characters of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFD93CF5FABA9B44A" authorityName="Rudzinski" authorityYear="2006" box="[536,749,1628,1655]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFD93CF5FABA9B44A" bold="true" box="[536,749,1628,1655]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EFCB4B2DEF29DC4DFC8BCF5EAA8CB445" author="Rudzinski, H. - G." box="[768,968,1629,1656]" pageId="2" pageNumber="3" pagination="449 - 476" refId="ref24659" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochroa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27, 449 - 476." type="journal article" year="2006">Rudzinski (2006)</bibRefCitation>
|
||
postulated a close relationship between
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFF1CCF82A822B4A6" authorityName="Rudzinski" authorityYear="2006" box="[151,358,1665,1691]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFF1CCF82A822B4A6" box="[151,358,1665,1691]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFE29CF82ABC4B4A6" authority="Mohrig" authorityName="Mohrig" authorityYear="1999" box="[418,640,1665,1691]" class="Insecta" family="Sciaridae" genus="Aerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFE29CF82AB66B4A6" box="[418,546,1665,1691]" italics="true" pageId="2" pageNumber="3">Aerumnosa</emphasis>
|
||
Mohrig
|
||
</taxonomicName>
|
||
on the basis of reduced mouthparts (labrum, labellum and maxillary palpus); a long wing vein R
|
||
<quantity id="4CA29B39EF29DC4DFE47CFA6A8B6B482" box="[460,498,1701,1727]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="3" unit="in" value="1.0">1 in</quantity>
|
||
relation to R; R1 and R5 with ventral setae in addition to the usual dorsal ones, and a wide gonostylus with a group of mesial megasetae. However, the morphology of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFB94CFCAADDBB4DE" authorityName="Mohrig" authorityYear="1999" box="[1055,1183,1737,1763]" class="Insecta" family="Sciaridae" genus="Aerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFB94CFCAADDBB4DE" box="[1055,1183,1737,1763]" italics="true" pageId="2" pageNumber="3">Aerumnosa</emphasis>
|
||
</taxonomicName>
|
||
differs strikingly from
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFF1CCFEEA822B53A" authorityName="Rudzinski" authorityYear="2006" box="[151,358,1773,1799]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFF1CCFEEA822B53A" box="[151,358,1773,1799]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
in many characters: the body is larger, the whole hypopygium is highly modified with complex structures of the gonocoxa, gonostylus and tegmen, the antenna is unmodified and the fore tibial organ is large with a dense patch of setae (see
|
||
<bibRefCitation id="EFCB4B2DEF29DC4DFE4DCE36AB19B572" author="Mohrig, W." box="[454,605,1845,1871]" pageId="2" pageNumber="3" pagination="153 - 203" refId="ref24613" refString="Mohrig, W. (1999) Die Trauermucken (Diptera: Sciaridae) von Papua-Neuguinea. Teil I. Gattungen Sciara, Schwenckfeldina, Aerumnosa gen. nov., Cratyna, Phytosciara und Chaetosciara. Studia dipterologica, 6, 153 - 203." type="journal article" year="1999">Mohrig 1999</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3406557EF29DC4DFF4CCE5AAAD0B586" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="8BE536DCEF29DC4DFF4CCE5AAAD0B586" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="3">
|
||
Based on a phylogenetic analysis using four gene markers, (
|
||
<collectionCode id="ED4BAE19EF29DC4DFCEFCE5AAADCB54E" box="[868,920,1881,1907]" country="Portugal" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14498" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="2" pageNumber="3" type="Herbarium">COI</collectionCode>
|
||
, 16S, 18S and 28S) as characters,
|
||
<bibRefCitation id="EFCB4B2DEF29DC4DFA9DCE5AA841B5AA" author="Vilkamaa, P. & Rudzinski, H. - G. & Burdikova, N. & Sevcik, J." pageId="2" pageNumber="3" pagination="248 - 260" refId="ref24856" refString="Vilkamaa, P., Rudzinski, H. - G., Burdikova, N. & Sevcik, J. (2018) Phylogenetic position of Aerumnosa Mohrig (Diptera, Sciaridae) as revealed by multigene analysis, with the description of four new Oriental species. Zootaxa, 4399 (1), 248 - 260. https: // doi. org / 10.11646 / zootaxa. 4399.2.8" type="journal article" year="2018">
|
||
Vilkamaa
|
||
<emphasis id="B92EEACEEF29DC4DFA03CE59A9E9B5AA" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. (2018)
|
||
</bibRefCitation>
|
||
suggested that both
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFE61CE7EAB2EB5AA" authorityName="Mohrig" authorityYear="1999" box="[490,618,1917,1943]" class="Insecta" family="Sciaridae" genus="Aerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFE61CE7EAB2EB5AA" box="[490,618,1917,1943]" italics="true" pageId="2" pageNumber="3">Aerumnosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFD14CE7EAA2AB5AA" authorityName="Rudzinski" authorityYear="2006" box="[671,878,1917,1943]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFD14CE7EAA2AB5AA" box="[671,878,1917,1943]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
belong to the Cratyninae
|
||
<emphasis id="B92EEACEEF29DC4DFB05CE7DAD88B5AA" box="[1166,1228,1918,1943]" italics="true" pageId="2" pageNumber="3">sensu</emphasis>
|
||
<bibRefCitation id="EFCB4B2DEF29DC4DFB59CE7EACDCB5AA" author="Shin, S. & Jung, S. & Menzel, F. & Heller, K. & Lee, H. & Lee, S." box="[1234,1432,1917,1943]" pageId="2" pageNumber="3" pagination="833 - 846" refId="ref24776" refString="Shin, S., Jung, S., Menzel, F., Heller, K., Lee, H. & Lee, S. (2013) Molecular phylogeny of black fungus gnats (Diptera: Sciaroidea: Sciaridae) and the evolution of larval habitats. Molecular Phylogenetics and Evolution, 66 (3), 833 - 846. https: // doi. org / 10.1016 / j. ympev. 2012.11.008" type="journal article" year="2013">
|
||
Shin
|
||
<emphasis id="B92EEACEEF29DC4DFA87CE7DAC79B5AA" box="[1292,1341,1917,1943]" italics="true" pageId="2" pageNumber="3">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
, with
|
||
<emphasis id="B92EEACEEF29DC4DFF5BCEA2A82BB586" box="[208,367,1953,1979]" italics="true" pageId="2" pageNumber="3">Peyerimhoffia</emphasis>
|
||
Kieffer as the sister group of
|
||
<emphasis id="B92EEACEEF29DC4DFD36CEA2AAD0B586" box="[701,916,1953,1979]" italics="true" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFD36CEA2AAD4B586" authorityName="Rudzinski" authorityYear="2006" box="[701,912,1953,1979]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Pseudoaerumnosa</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3406557EF29DC4CFF4CCEC7ABD7B3B6" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="8BE536DCEF29DC4CFF4CCEC7ABD7B3B6" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1437,153,395]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="B92EEACEEF29DC4DFF4CCEC7AB88B5E2" bold="true" box="[199,716,1988,2015]" pageId="2" pageNumber="3">
|
||
Diagnostic characters of
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFE7BCEC7AB81B5E2" ID-CoL="6Z6Q" ID-ENA="2058813" authorityName="Rudzinski" authorityYear="2006" box="[496,709,1988,2015]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFE7BCEC7AB81B5E2" bold="true" box="[496,709,1988,2015]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Regarding morphology,
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFC6FCEC6ADF7B5E2" authorityName="Rudzinski" authorityYear="2006" box="[996,1203,1989,2015]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF29DC4DFC6FCEC6ADF7B5E2" box="[996,1203,1989,2015]" italics="true" pageId="2" pageNumber="3">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<emphasis id="B92EEACEEF29DC4CFACBCEC6A9F3B28E" italics="true" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C5A4D5FEF29DC4DFACBCEC6ACD8B5E2" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[1344,1436,1989,2015]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" sensu="lato">Cratyna</taxonomicName>
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFF1CC99AA9F3B28E" box="[151,183,153,179]" pageId="3" pageNumber="4" sensu="lato">s.l.</taxonomicNameLabel>
|
||
</emphasis>
|
||
a plump abdomen, a broad hypopygium, a reduced maxillary palpus, a setose postpronotum, a short gonocoxa and the lack of robust spinose setae on the tibiae. Some species of
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFC08C9BEAD16B2EA" authorityName="Rudzinski" authorityYear="2006" box="[899,1106,189,215]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF28DC4CFC08C9BEAD16B2EA" box="[899,1106,189,215]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
have a finger-like process on their tegmen, like many species of
|
||
<emphasis id="B92EEACEEF28DC4CFD9DC9E2ABF5B2C6" box="[534,689,225,251]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFD9DC9E2AB36B2C6" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[534,626,225,251]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" sensu="stricto">Cratyna</taxonomicName>
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFDFCC9E1ABF5B2C6" box="[631,689,226,251]" pageId="3" pageNumber="4" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and some
|
||
<emphasis id="B92EEACEEF28DC4CFCAEC9E2AAFDB2C6" box="[805,953,225,251]" italics="true" pageId="3" pageNumber="4">Spathobdella</emphasis>
|
||
Frey (see
|
||
<bibRefCitation id="EFCB4B2DEF28DC4CFBA1C9E2AC71B2C6" author="Menzel, F. & Mohrig, W." box="[1066,1333,225,251]" pageId="3" pageNumber="4" pagination="1 - 761" refId="ref24576" refString="Menzel, F. & Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica, Supple- ment, 6 (1999), 1 - 761." type="journal article" year="2000">Menzel & Mohrig 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCB4B2DEF28DC4CFAB5C9E2A9B4B322" author="Hippa, H. & Vilkamaa, P. & Heinakroon, A." pageId="3" pageNumber="4" pagination="1 - 86" refId="ref24480" refString="Hippa, H., Vilkamaa, P. & Heinakroon, A. (1998) The genus Pseudozygoneura Steffan (Diptera, Sciaridae). Acta Zoologica Fennica, 210, 1 - 86." type="journal article" year="1998">
|
||
Hippa
|
||
<emphasis id="B92EEACEEF28DC4CFA03C9E1A9E9B322" italics="true" pageId="3" pageNumber="4">et al</emphasis>
|
||
.1998
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFE88C806A896B322" authorityName="Rudzinski" authorityYear="2006" box="[259,466,261,287]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF28DC4CFE88C806A896B322" box="[259,466,261,287]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFDD5C806AA36B322" authority="" box="[606,882,261,287]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Peyerimhoffia">
|
||
<emphasis id="B92EEACEEF28DC4CFDD5C806ABFEB322" box="[606,698,261,287]" italics="true" pageId="3" pageNumber="4">Cratyna</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF28DC4CFD41C806AA2DB322" box="[714,873,261,287]" italics="true" pageId="3" pageNumber="4">Peyerimhoffia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
and with
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFC6BC806ADADB322" authority="" box="[992,1257,261,287]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Spathobdella">
|
||
<emphasis id="B92EEACEEF28DC4CFC6BC806AD78B322" box="[992,1084,261,287]" italics="true" pageId="3" pageNumber="4">Cratyna</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF28DC4CFBC7C806ADA4B322" box="[1100,1248,261,287]" italics="true" pageId="3" pageNumber="4">Spathobdella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
a reduced maxillary palpus and a modified apical tooth of the gonostylus with a megaseta or megasetae, although the megaseta in
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFF1CC84EA822B35A" authorityName="Rudzinski" authorityYear="2006" box="[151,358,333,359]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF28DC4CFF1CC84EA822B35A" box="[151,358,333,359]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
is attached to the ventral side of the tooth, not inside of it, as in
|
||
<emphasis id="B92EEACEEF28DC4CFBBFC84EAD97B35A" box="[1076,1235,333,359]" italics="true" pageId="3" pageNumber="4">Peyerimhoffia</emphasis>
|
||
and
|
||
<emphasis id="B92EEACEEF28DC4CFA83C84EACD8B35A" box="[1288,1436,333,359]" italics="true" pageId="3" pageNumber="4">Spathobdella</emphasis>
|
||
(and in the
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFE9DC872AB7EB3B6" baseAuthorityName="Frey" baseAuthorityYear="1948" box="[278,570,369,395]" class="Insecta" family="Sciaridae" genus="Corynoptera" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="crassistylata">
|
||
<emphasis id="B92EEACEEF28DC4CFE9DC872AB7EB3B6" box="[278,570,369,395]" italics="true" pageId="3" pageNumber="4">Corynoptera crassistylata</emphasis>
|
||
</taxonomicName>
|
||
group).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF256654EF28DC4CFF1CCF3AA82EB4A7" httpUri="https://zenodo.org/record/3366500/files/figure.png" pageId="3" pageNumber="4" startId="3.[151,250,1593,1618]" targetBox="[159,1427,425,1564]" targetPageId="3">
|
||
<paragraph id="8BE536DCEF28DC4CFF1CCF3AA82EB4A7" blockId="3.[151,1437,1593,1690]" pageId="3" pageNumber="4">
|
||
<emphasis id="B92EEACEEF28DC4CFF1CCF3AA857B46F" bold="true" box="[151,275,1593,1618]" pageId="3" pageNumber="4">FIGURE 2.</emphasis>
|
||
Wing, dorsal. A:
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFE4ECF39AB8EB46F" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[453,714,1594,1618]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="banari" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFE4ECF39AB8EB46F" box="[453,714,1594,1618]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa banari</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFD44CF39AA4CB46F" bold="true" box="[719,776,1594,1618]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFD44CF39AA4CB46F" box="[719,776,1594,1618]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratype).
|
||
<emphasis id="B92EEACEEF28DC4CFC09CF39AAD8B46F" bold="true" box="[898,924,1594,1618]" pageId="3" pageNumber="4">B:</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFC29CF38AD48B46F" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[930,1036,1594,1618]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="eminula" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFC29CF38AD48B46F" box="[930,1036,1594,1618]" italics="true" pageId="3" pageNumber="4">P. eminula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFB99CF39AD0FB46F" bold="true" box="[1042,1099,1594,1618]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFB99CF39AD0FB46F" box="[1042,1099,1594,1618]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratype from Ulu Temburong).
|
||
<emphasis id="B92EEACEEF28DC4CFF1CCF5EA9F6B44B" bold="true" box="[151,178,1629,1654]" pageId="3" pageNumber="4">C:</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFF32CF5CA85FB44B" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[185,283,1630,1654]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fragilis" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFF32CF5CA85FB44B" box="[185,283,1630,1654]" italics="true" pageId="3" pageNumber="4">P. fragilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFEA9CF5DA818B44B" bold="true" box="[290,348,1630,1654]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFEA9CF5DA818B44B" box="[290,348,1630,1654]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype).
|
||
<emphasis id="B92EEACEEF28DC4CFE52CF5DA8B0B44B" bold="true" box="[473,500,1630,1654]" pageId="3" pageNumber="4">D:</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFE70CF5CAB31B44B" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[507,629,1630,1654]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tenuidens" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFE70CF5CAB31B44B" box="[507,629,1630,1654]" italics="true" pageId="3" pageNumber="4">P. tenuidens</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFDF7CF5DABF2B44B" bold="true" box="[636,694,1630,1654]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFDF7CF5DABF2B44B" box="[636,694,1630,1654]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype).
|
||
<emphasis id="B92EEACEEF28DC4CFCB8CF5DAA08B44B" bold="true" box="[819,844,1630,1654]" pageId="3" pageNumber="4">E:</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFCDFCF5CAA8CB44B" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[852,968,1631,1654]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="consuota" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFCDFCF5CAA8CB44B" box="[852,968,1631,1654]" italics="true" pageId="3" pageNumber="4">P. consuota</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFC44CF5DAD4DB44B" bold="true" box="[975,1033,1630,1654]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFC44CF5DAD4DB44B" box="[975,1033,1630,1654]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratype).
|
||
<emphasis id="B92EEACEEF28DC4CFB0ECF5DADD9B44B" bold="true" box="[1157,1181,1630,1654]" pageId="3" pageNumber="4">F:</emphasis>
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFB2FCF5CAC5CB44B" authority="Vilkamaa & Halenius & Ševčík, 2019" authorityName="Vilkamaa & Halenius & Ševčík" authorityYear="2019" box="[1188,1304,1631,1654]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="junciseta" status="sp. nov.">
|
||
<emphasis id="B92EEACEEF28DC4CFB2FCF5CAC5CB44B" box="[1188,1304,1631,1654]" italics="true" pageId="3" pageNumber="4">P. junciseta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92EEACEEF28DC4CFA94CF5DAC1DB44B" bold="true" box="[1311,1369,1630,1654]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A21D57B5EF28DC4CFA94CF5DAC1DB44B" box="[1311,1369,1630,1654]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(paratype). Scale 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3406557EF28DC4CFF4CCFCBABE2B587" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="8BE536DCEF28DC4CFF4CCFCBABE2B587" blockId="3.[151,1436,1736,1979]" pageId="3" pageNumber="4">
|
||
Furthermore,
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFEEFCFCBAB77B4DF" authorityName="Rudzinski" authorityYear="2006" box="[356,563,1736,1762]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF28DC4CFEEFCFCBAB77B4DF" box="[356,563,1736,1762]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
is actually morphologically more similar to
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFBBCCFCAAC07B4DE" authority="" box="[1079,1347,1736,1763]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Spathobdella">
|
||
<emphasis id="B92EEACEEF28DC4CFBBCCFCAADD7B4DE" box="[1079,1171,1737,1763]" italics="true" pageId="3" pageNumber="4">Cratyna</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF28DC4CFB2DCFCBAC7EB4DF" box="[1190,1338,1736,1762]" italics="true" pageId="3" pageNumber="4">Spathobdella</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
than to
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFF1CCFEEA8EFB53A" authority="" box="[151,427,1772,1799]" class="Insecta" family="Sciaridae" genus="Cratyna" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="subGenus" subGenus="Peyerimhoffia">
|
||
<emphasis id="B92EEACEEF28DC4CFF1CCFEEA9B7B53A" box="[151,243,1773,1799]" italics="true" pageId="3" pageNumber="4">Cratyna</emphasis>
|
||
(
|
||
<emphasis id="B92EEACEEF28DC4CFE8FCFEFA8E7B53B" box="[260,419,1772,1798]" italics="true" pageId="3" pageNumber="4">Peyerimhoffia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
in having the apical tooth with a broad basal part and a narrow apical part, the gonostylar megasetae of the similar hyalinous
|
||
<typeStatus id="54E1887EEF28DC4CFD94CE12AB16B516" box="[543,594,1809,1835]" pageId="3" pageNumber="4">type</typeStatus>
|
||
, in having long slender setae apically and subapically on the gonostylus and in having a weakly sclerotized tegmen without basal sclerotization.
|
||
<taxonomicName id="4C5A4D5FEF28DC4CFC2BCE37AD2BB573" authorityName="Rudzinski" authorityYear="2006" box="[928,1135,1844,1870]" class="Insecta" family="Sciaridae" genus="Pseudoaerumnosa" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92EEACEEF28DC4CFC2BCE37AD2BB573" box="[928,1135,1844,1870]" italics="true" pageId="3" pageNumber="4">Pseudoaerumnosa</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<emphasis id="B92EEACEEF28DC4CFA83CE37ACD8B573" box="[1288,1436,1844,1870]" italics="true" pageId="3" pageNumber="4">Spathobdella</emphasis>
|
||
in having the gonocoxae united, not separate, in lacking specialized setae apically on the gonostylus (on the dorsal side) and in having the antennal flagellomeres asymmetric, only a few setae in the fore tibial organ and the attached megaseta on the apical tooth of the gonostylus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |