389 lines
55 KiB
XML
389 lines
55 KiB
XML
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<mods:title id="88C388EE7AC9B9E6B7F3992A9F817643">New taxa of Afrotropical Toktokkies (Coleoptera: Tenebrionidae: Molurina) from the Natural History Museum of Basel</mods:title>
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<mods:namePart id="760524FCB3255619870E2CE3B8DC0DA7">Kamiński, Marcin Jan</mods:namePart>
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<mods:date id="44C9E75DFE75A34934DEA298C50EB526">2024</mods:date>
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<mods:identifier id="AD55A03838DAB6797B8037D2FCE54581" type="DOI">10.11646/zootaxa.5446.1.4</mods:identifier>
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<treatment id="03F36C399A1ED467D9FBFBD06694FCC0" ID-DOI="http://doi.org/10.5281/zenodo.11106883" ID-GBIF-Taxon="228888836" ID-Zenodo-Dep="11106883" LSID="urn:lsid:plazi:treatment:03F36C399A1ED467D9FBFBD06694FCC0" httpUri="http://treatment.plazi.org/id/03F36C399A1ED467D9FBFBD06694FCC0" lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78">
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<subSubSection id="C3408EA49A1ED464D9FBFBD062BCFBC8" box="[151,436,1122,1149]" pageId="1" pageNumber="78" type="nomenclature">
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<paragraph id="8BE5DD2F9A1ED464D9FBFBD062BCFBC8" blockId="1.[151,436,1122,1149]" box="[151,436,1122,1149]" pageId="1" pageNumber="78">
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<taxonomicName id="4C5AA6AC9A1ED464D9FBFBD06237FBC8" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[151,319,1122,1149]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92E013D9A1ED464D9FBFBD06237FBC8" bold="true" box="[151,319,1122,1149]" italics="true" pageId="1" pageNumber="78">Bouchardium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21DBC469A1ED464D82AFBD162BCFBC8" box="[326,436,1123,1149]" pageId="1" pageNumber="78" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3408EA49A1ED464D9FBFB1960AEFB70" box="[151,934,1194,1221]" pageId="1" pageNumber="78" type="type_taxon">
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<paragraph id="8BE5DD2F9A1ED464D9FBFB1960AEFB70" blockId="1.[151,1437,1194,2013]" box="[151,934,1194,1221]" pageId="1" pageNumber="78">
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<emphasis id="B92E013D9A1ED464D9FBFB19623AFB71" bold="true" box="[151,306,1194,1221]" pageId="1" pageNumber="78">Type species.</emphasis>
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<taxonomicName id="4C5AA6AC9A1ED464D855FB196188FB70" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[313,640,1195,1221]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="chillygonzalesi" status="sp. nov.">
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<emphasis id="B92E013D9A1ED464D855FB196188FB70" box="[313,640,1195,1221]" italics="true" pageId="1" pageNumber="78">Bouchardium chillygonzalesi</emphasis>
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</taxonomicName>
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<emphasis id="B92E013D9A1ED464DBEBFB1961E8FB70" bold="true" box="[647,736,1195,1221]" pageId="1" pageNumber="78">
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<taxonomicNameLabel id="A21DBC469A1ED464DBEBFB1961E8FB70" box="[647,736,1195,1221]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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; here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3408EA49A1ED467D9ABFB7C62AAFEA8" lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78" type="diagnosis">
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||
<paragraph id="8BE5DD2F9A1ED464D9ABFB7C6060F9C0" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="78">
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<emphasis id="B92E013D9A1ED464D9ABFB7C624AFB5D" bold="true" box="[199,322,1230,1256]" pageId="1" pageNumber="78">Diagnosis.</emphasis>
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The specific structure of the prosternum (elongate and projecting tip of prosternum, in lateral view,
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<figureCitation id="1361C1AA9A1ED464D9FBFB4163F9FAB8" box="[151,241,1267,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1377,1401]" captionTargetBox="[155,1429,208,1352]" captionTargetId="figure-15@3.[151,1436,181,1353]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." figureDoi="http://doi.org/10.5281/zenodo.11084606" httpUri="https://zenodo.org/record/11084606/files/figure.png" pageId="1" pageNumber="78">Figs 1A</figureCitation>
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,
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<figureCitation id="1361C1AA9A1ED464D997FB416231FAB8" box="[251,313,1267,1293]" captionStart="FIGURE 3" captionStartId="5.[151,250,1404,1428]" captionTargetBox="[173,1414,237,1367]" captionTargetId="figure-41@5.[164,1423,237,1381]" captionTargetPageId="5" captionText="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi="http://doi.org/10.5281/zenodo.11084614" httpUri="https://zenodo.org/record/11084614/files/figure.png" pageId="1" pageNumber="78">3F, K</figureCitation>
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), elytra (presence of sharp lateral costae,
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<figureCitation id="1361C1AA9A1ED464DB91FB416056FAB8" box="[765,862,1267,1293]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1377,1401]" captionStartId-1="4.[151,250,1685,1709]" captionStartId-2="5.[151,250,1404,1428]" captionTargetBox-0="[155,1429,208,1352]" captionTargetBox-1="[159,1428,309,1652]" captionTargetBox-2="[173,1414,237,1367]" captionTargetId-0="figure-15@3.[151,1436,181,1353]" captionTargetId-1="figure-67@4.[151,1436,309,1661]" captionTargetId-2="figure-41@5.[164,1423,237,1381]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." captionText-1="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." captionText-2="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi-0="http://doi.org/10.5281/zenodo.11084606" figureDoi-1="http://doi.org/10.5281/zenodo.11084610" figureDoi-2="http://doi.org/10.5281/zenodo.11084614" httpUri-0="https://zenodo.org/record/11084606/files/figure.png" httpUri-1="https://zenodo.org/record/11084610/files/figure.png" httpUri-2="https://zenodo.org/record/11084614/files/figure.png" pageId="1" pageNumber="78">Figs 1–3</figureCitation>
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), and scutellum (apex with deep transverse grooves,
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<figureCitation id="1361C1AA9A1ED464D9FBFAA56206FA84" box="[151,270,1303,1329]" captionStart="FIGURE 3" captionStartId="5.[151,250,1404,1428]" captionTargetBox="[173,1414,237,1367]" captionTargetId="figure-41@5.[164,1423,237,1381]" captionTargetPageId="5" captionText="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi="http://doi.org/10.5281/zenodo.11084614" httpUri="https://zenodo.org/record/11084614/files/figure.png" pageId="1" pageNumber="78">Fig. 3A–C</figureCitation>
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) place
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<taxonomicName id="4C5AA6AC9A1ED464D832FAA562FDFA84" authorityName="Kamiński" authorityYear="2024" box="[350,501,1303,1329]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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<emphasis id="B92E013D9A1ED464D832FAA562FDFA84" box="[350,501,1303,1329]" italics="true" pageId="1" pageNumber="78">Bouchardium</emphasis>
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</taxonomicName>
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close to the following genera (
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<bibRefCitation id="EFCBA0DE9A1ED464DA20FAA56732FA84" author="Kaminski, M. J. & Gearner, O. M. & Kanda, K & Swichtenberg, K. & Purchart, L. & Smith, A. D." box="[844,1082,1303,1329]" pageId="1" pageNumber="78" pagination="883 - 901" refId="ref6154" refString="Kaminski, M. J., Gearner, O. M., Kanda, K .., Swichtenberg, K., Purchart, L. & Smith, A. D. (2021) First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina), and examination of the status of the Psammodes vialis species-group. Zoological Journal of the Linnean Society, 191, 883 - 901. https: // doi. org / 10.1093 / zoolinnean / zlaa 052" type="journal article" year="2021">
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||
Kamiński
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||
<emphasis id="B92E013D9A1ED464DAADFAA560F1FA84" box="[961,1017,1303,1329]" italics="true" pageId="1" pageNumber="78">et al.</emphasis>
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||
2021
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||
</bibRefCitation>
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||
,
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||
<bibRefCitation id="EFCBA0DE9A1ED464DD29FAA56629FA84" author="Gearner, O. M. & Lumen, R. & Kaminski, M. J." box="[1093,1313,1303,1329]" pageId="1" pageNumber="78" pagination="123 - 127" refId="ref5849" refString="Gearner, O. M., Lumen, R. & Kaminski, M. J. (2022) New species of Toktokkus from Mozambique (Tenebrionidae: Sepidiini: Molurina). Annales Zoologici, 72, 123 - 127. https: // doi. org / 10.3161 / 00034541 ANZ 2022.72.1.010" type="journal article" year="2022">
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||
Gearner
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||
<emphasis id="B92E013D9A1ED464DDC9FAA567D5FA84" box="[1189,1245,1303,1329]" italics="true" pageId="1" pageNumber="78">et al.</emphasis>
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||
2022
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||
</bibRefCitation>
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||
):
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||
<taxonomicName id="4C5AA6AC9A1ED464DC5EFAA56244FAE0" authority="Fahraeus, 1870" authorityName="Fahraeus" authorityYear="1870" class="Insecta" family="Tenebrionidae" genus="Amiantus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92E013D9A1ED464DC5EFAA56694FA84" box="[1330,1436,1303,1329]" italics="true" pageId="1" pageNumber="78">Amiantus</emphasis>
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||
Fåhraeus, 1870
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</taxonomicName>
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,
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||
<taxonomicName id="4C5AA6AC9A1ED464D832FA896157FAE0" authority="Koch, 1951" authorityName="Koch" authorityYear="1951" box="[350,607,1338,1365]" class="Insecta" family="Tenebrionidae" genus="Arturium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92E013D9A1ED464D832FA8962CDFAE0" box="[350,453,1339,1365]" italics="true" pageId="1" pageNumber="78">Arturium</emphasis>
|
||
<bibRefCitation id="EFCBA0DE9A1ED464D8BFFA896157FAE0" author="Koch, C." box="[467,607,1338,1365]" pageId="1" pageNumber="78" pagination="89 - 96" refId="ref6369" refString="Koch, C. (1951) Die Tenebrioniden des sudlichen Afikas VII - Arturium nov. gen. Molurinorum ex aff. Phrynocolus Lac. Atti della Societa Italiana di Scienze Naturali, 90, 89 - 96." type="journal article" year="1951">Koch, 1951</bibRefCitation>
|
||
</taxonomicName>
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||
,
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||
<taxonomicName id="4C5AA6AC9A1ED464DB1EFA8960E0FAE0" authority="Fairmaire, 1882" authorityName="Fairmaire" authorityYear="1882" box="[626,1000,1339,1365]" class="Insecta" family="Tenebrionidae" genus="Brachyphrynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92E013D9A1ED464DB1EFA896015FAE0" box="[626,797,1339,1365]" italics="true" pageId="1" pageNumber="78">Brachyphrynus</emphasis>
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||
Fairmaire, 1882
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B92E013D9A1ED464DA96FA896753FAE0" box="[1018,1115,1339,1365]" italics="true" pageId="1" pageNumber="78">
|
||
<taxonomicName id="4C5AA6AC9A1ED464DA96FA89675FFAE0" authorityName="Haag-Rutenberg" authorityYear="1871" box="[1018,1111,1339,1365]" class="Insecta" family="Tenebrionidae" genus="Dichtha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Dichtha</taxonomicName>
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||
,
|
||
</emphasis>
|
||
<taxonomicName id="4C5AA6AC9A1ED464DD05FA8963D9FACC" authority="Haag-Rutenberg, 1871" authorityName="Haag-Rutenberg" authorityYear="1871" class="Insecta" family="Tenebrionidae" genus="Distretus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DD05FA8967C7FAE1" box="[1129,1231,1339,1364]" italics="true" pageId="1" pageNumber="78">Distretus</emphasis>
|
||
<bibRefCitation id="EFCBA0DE9A1ED464DDB1FA8963D9FACC" author="Haag-Rutenberg, G." pageId="1" pageNumber="78" pagination="21 - 111" refId="ref5905" refString="Haag-Rutenberg, G. (1871) Beitrage zur Familie der Tenebrioniden (II. Stuck). Coleopterologische Hefte, VII, 21 - 111." type="journal article" year="1871">Haag-Rutenberg, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1ED464D98CFAED6113FACC" authority="Fairmaire, 1897" authorityName="Fairmaire" authorityYear="1897" box="[224,539,1375,1401]" class="Insecta" family="Tenebrionidae" genus="Euphrynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92E013D9A1ED464D98CFAED6251FACC" box="[224,345,1375,1401]" italics="true" pageId="1" pageNumber="78">Euphrynus</emphasis>
|
||
Fairmaire, 1897
|
||
</taxonomicName>
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||
,
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||
<taxonomicName id="4C5AA6AC9A1ED464DB45FAED6099FACC" authority="Fairmaire, 1899" authorityName="Fairmaire" authorityYear="1899" box="[553,913,1375,1401]" class="Insecta" family="Tenebrionidae" genus="Glyptophrynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B92E013D9A1ED464DB45FAED61C7FACC" box="[553,719,1375,1401]" italics="true" pageId="1" pageNumber="78">Glyptophrynus</emphasis>
|
||
Fairmaire, 1899
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||
</taxonomicName>
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||
,
|
||
<taxonomicName id="4C5AA6AC9A1ED464DAF3FAED6609FACC" authority="Fairmaire, 1882" authorityName="Fairmaire" authorityYear="1882" box="[927,1281,1375,1401]" class="Insecta" family="Tenebrionidae" genus="Melanolophus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DAF3FAED6737FACC" box="[927,1087,1375,1401]" italics="true" pageId="1" pageNumber="78">Melanolophus</emphasis>
|
||
Fairmaire, 1882
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1ED464DC63FAED6256FA28" authority="Lacordaire, 1859" authorityName="Lacordaire" authorityYear="1859" class="Insecta" family="Tenebrionidae" genus="Phrynocolus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DC63FAED6695FACC" box="[1295,1437,1375,1401]" italics="true" pageId="1" pageNumber="78">Phrynocolus</emphasis>
|
||
Lacordaire, 1859
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1ED464D801FA3161ABFA28" authority="Koch, 1951" authorityName="Koch" authorityYear="1951" box="[365,675,1410,1437]" class="Insecta" family="Tenebrionidae" genus="Phrynophanes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464D801FA316107FA28" box="[365,527,1411,1437]" italics="true" pageId="1" pageNumber="78">Phrynophanes</emphasis>
|
||
<bibRefCitation id="EFCBA0DE9A1ED464DB76FA3161ABFA28" author="Koch, C." box="[538,675,1410,1437]" pageId="1" pageNumber="78" pagination="89 - 96" refId="ref6369" refString="Koch, C. (1951) Die Tenebrioniden des sudlichen Afikas VII - Arturium nov. gen. Molurinorum ex aff. Phrynocolus Lac. Atti della Societa Italiana di Scienze Naturali, 90, 89 - 96." type="journal article" year="1951">Koch, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB8BFA316744FA28" authority="Fairmaire, 1882" authorityName="Fairmaire" authorityYear="1882" box="[743,1100,1411,1437]" class="Insecta" family="Tenebrionidae" genus="Physophrynus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DB8BFA31608EFA28" box="[743,902,1411,1437]" italics="true" pageId="1" pageNumber="78">Physophrynus</emphasis>
|
||
Fairmaire, 1882
|
||
</taxonomicName>
|
||
. Among these,
|
||
<taxonomicName id="4C5AA6AC9A1ED464DC69FA316694FA28" authorityName="Kamiński" authorityYear="2024" box="[1285,1436,1411,1437]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DC69FA316694FA28" box="[1285,1436,1411,1437]" italics="true" pageId="1" pageNumber="78">Bouchardium</emphasis>
|
||
</taxonomicName>
|
||
displays the greatest affinity towards
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB59FA156193FA75" authorityName="Haag-Rutenberg" authorityYear="1871" box="[565,667,1447,1472]" class="Insecta" family="Tenebrionidae" genus="Distretus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DB59FA156193FA75" box="[565,667,1447,1472]" italics="true" pageId="1" pageNumber="78">Distretus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AA6AC9A1ED464DBA3FA156021FA74" authorityName="Haag-Rutenberg" authorityYear="1871" box="[719,809,1447,1473]" class="Insecta" family="Tenebrionidae" genus="Dichtha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DBA3FA156021FA74" box="[719,809,1447,1473]" italics="true" pageId="1" pageNumber="78">Dichtha</emphasis>
|
||
</taxonomicName>
|
||
as all share a similar structure of elytra (sharply carinate humeral edge and flattened surface of elytra). Nevertheless,
|
||
<taxonomicName id="4C5AA6AC9A1ED464DA3BFA7960E6FA50" authorityName="Kamiński" authorityYear="2024" box="[855,1006,1483,1509]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DA3BFA7960E6FA50" box="[855,1006,1483,1509]" italics="true" pageId="1" pageNumber="78">Bouchardium</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all of the abovementioned genera, including
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB4EFA5D6174F9BC" authorityName="Haag-Rutenberg" authorityYear="1871" box="[546,636,1519,1545]" class="Insecta" family="Tenebrionidae" genus="Dichtha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DB4EFA5D6174F9BC" box="[546,636,1519,1545]" italics="true" pageId="1" pageNumber="78">Dichtha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AA6AC9A1ED464DBDFFA5D6011F9BD" authorityName="Haag-Rutenberg" authorityYear="1871" box="[691,793,1519,1544]" class="Insecta" family="Tenebrionidae" genus="Distretus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1ED464DBDFFA5D6011F9BD" box="[691,793,1519,1544]" italics="true" pageId="1" pageNumber="78">Distretus</emphasis>
|
||
</taxonomicName>
|
||
, by a unique arrangement of pronotal punctures (sides of the pronotal disc with large, confluent punctures; center of the disc with small punctures, 4–5 diameters apart vs pronotal disc evenly covered with large sometimes confluent punctures—
|
||
<figureCitation id="1361C1AA9A1ED464DABBF9856747F9E4" box="[983,1103,1591,1617]" captionStart="FIGURE 2" captionStartId="4.[151,250,1685,1709]" captionTargetBox="[159,1428,309,1652]" captionTargetId="figure-67@4.[151,1436,309,1661]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." figureDoi="http://doi.org/10.5281/zenodo.11084610" httpUri="https://zenodo.org/record/11084610/files/figure.png" pageId="1" pageNumber="78">Figs 2A, F</figureCitation>
|
||
,
|
||
<figureCitation id="1361C1AA9A1ED464DD37F98567DCF9E4" box="[1115,1236,1591,1617]" captionStart="FIGURE 3" captionStartId="5.[151,250,1404,1428]" captionTargetBox="[173,1414,237,1367]" captionTargetId="figure-41@5.[164,1423,237,1381]" captionTargetPageId="5" captionText="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi="http://doi.org/10.5281/zenodo.11084614" httpUri="https://zenodo.org/record/11084614/files/figure.png" pageId="1" pageNumber="78">3D, F, G, I</figureCitation>
|
||
) and subquadrate shape of the body (frontal/posterior views) (
|
||
<figureCitation id="1361C1AA9A1ED464DBE9F9E961D3F9C0" box="[645,731,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1377,1401]" captionTargetBox="[155,1429,208,1352]" captionTargetId="figure-15@3.[151,1436,181,1353]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." figureDoi="http://doi.org/10.5281/zenodo.11084606" httpUri="https://zenodo.org/record/11084606/files/figure.png" pageId="1" pageNumber="78">Fig. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="1361C1AA9A1ED464DB84F9E96025F9C0" box="[744,813,1627,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1685,1709]" captionTargetBox="[159,1428,309,1652]" captionTargetId="figure-67@4.[151,1436,309,1661]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." figureDoi="http://doi.org/10.5281/zenodo.11084610" httpUri="https://zenodo.org/record/11084610/files/figure.png" pageId="1" pageNumber="78">2C, D</figureCitation>
|
||
,
|
||
<figureCitation id="1361C1AA9A1ED464DA55F9E96053F9C0" box="[825,859,1627,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1404,1428]" captionTargetBox="[173,1414,237,1367]" captionTargetId="figure-41@5.[164,1423,237,1381]" captionTargetPageId="5" captionText="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi="http://doi.org/10.5281/zenodo.11084614" httpUri="https://zenodo.org/record/11084614/files/figure.png" pageId="1" pageNumber="78">3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A1ED464D9ABF9CD6780F80C" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="78">
|
||
<emphasis id="B92E013D9A1ED464D9ABF9CD6219F92C" bold="true" box="[199,273,1663,1689]" pageId="1" pageNumber="78">Notes.</emphasis>
|
||
According to
|
||
<bibRefCitation id="EFCBA0DE9A1ED464D8C2F9CD61A4F92C" author="Kaminski, M. J. & Gearner, O. M. & Ras, M. & Hunsinger, E. T. & Smith, A. L. & Mas-Peinado, P & Giron, J. C. & Bilska, A. G. & Strumpher, W. P. & Wirth, C. C. & Kanda, K. & Swichtenberg, K. & Iwan, D. & Smith, A. D." box="[430,684,1663,1689]" pageId="1" pageNumber="78" pagination="623 - 648" refId="ref6245" refString="Kaminski, M. J., Gearner, O. M., Ras, M., Hunsinger, E. T., Smith, A. L., Mas-Peinado, P, Giron, J. C., Bilska, A. G., Strumpher, W. P., Wirth, C. C., Kanda, K., Swichtenberg, K., Iwan, D. & Smith, A. D. (2022) Female terminalia morphology and cladistic relations among Tok-Tok beetles (Tenebrionidae: Sepidiini). Cladistics, 38, 623 - 648. https: // doi. org / 10.1111 / cla. 12510" type="journal article" year="2022">
|
||
Kamiński
|
||
<emphasis id="B92E013D9A1ED464DB4DF9CD6151F92C" box="[545,601,1663,1689]" italics="true" pageId="1" pageNumber="78">et al.</emphasis>
|
||
(2022)
|
||
</bibRefCitation>
|
||
, the morphology of female terminalia is relatively invariable within the
|
||
<emphasis id="B92E013D9A1ED464D9AFF911622EF908" box="[195,294,1699,1725]" italics="true" pageId="1" pageNumber="78">
|
||
‘
|
||
<taxonomicName id="4C5AA6AC9A1ED464D9ABF911622EF908" authorityName="Haag-Rutenberg" authorityYear="1871" box="[199,294,1699,1725]" class="Insecta" family="Tenebrionidae" genus="Dichtha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Dichtha</taxonomicName>
|
||
</emphasis>
|
||
clade’. All analyzed members of this grouping, including
|
||
<taxonomicName id="4C5AA6AC9A1ED464DAAAF9116607F908" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[966,1295,1699,1725]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="chillygonzalesi" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1ED464DAAAF9116607F908" box="[966,1295,1699,1725]" italics="true" pageId="1" pageNumber="78">Bouchardium chillygonzalesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1ED464DC74F911667CF908" bold="true" box="[1304,1396,1699,1725]" pageId="1" pageNumber="78">
|
||
<taxonomicNameLabel id="A21DBC469A1ED464DC74F911667CF908" box="[1304,1396,1699,1725]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1ED464DCE8F911612FF954" authority="(Guerin-Meneville, 1845)" authorityName="Kamiński" authorityYear="2024" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="cubicum" status="comb. nov.">
|
||
<emphasis id="B92E013D9A1ED464DCE8F91163FEF954" italics="true" pageId="1" pageNumber="78">B. cubicum</emphasis>
|
||
(Guérin-Méneville, 1845)
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1ED464DB5EF97561BEF954" bold="true" box="[562,694,1735,1761]" pageId="1" pageNumber="78">
|
||
<taxonomicNameLabel id="A21DBC469A1ED464DB5EF97561BEF954" box="[562,694,1735,1761]" pageId="1" pageNumber="78" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB99F975606FF955" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[757,871,1735,1760]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1ED464DB99F975606FF955" box="[757,871,1735,1760]" italics="true" pageId="1" pageNumber="78">B. mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1ED464DA1FF97560D9F954" bold="true" box="[883,977,1735,1761]" pageId="1" pageNumber="78">
|
||
<taxonomicNameLabel id="A21DBC469A1ED464DA1FF97560D9F954" box="[883,977,1735,1761]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
possess 3-lobed ovipositors, where the merged lobes 1 (valvifer) and 2 are strongly emarginate basally. Proctigers were reported to be widely indented medially, while bursae were simple (no sclerites were detected) and accompanied by multibranched spermatheca. Finally, spicula ventrali were reported to be Y-shaped (
|
||
<figureCitation id="1361C1AA9A1ED464DB94F8816059F8F8" box="[760,849,1843,1869]" captionStart="FIGURE 4" captionStartId="6.[151,250,1414,1438]" captionTargetBox="[159,1430,389,1382]" captionTargetId="figure-97@6.[151,1436,381,1390]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of female terminalia within the genus Bouchardium gen. nov. Bouchardium chillygonzalesi sp. nov. (A, B), B. mariae sp. nov. (C), B. cubicum (D). Ovipositor (A, B, D), spiculum ventrale (C). Abbreviations: c- coxities, c2-c4- subsequent lobes of coxities, v- valvifer, ip- inner plate of paraprocts, op- outer plate of paraprocts, pa - paraprocts." figureDoi="http://doi.org/10.5281/zenodo.11084619" httpUri="https://zenodo.org/record/11084619/files/figure.png" pageId="1" pageNumber="78">Fig. 4C</figureCitation>
|
||
). Nevertheless, some variability of the apical coxites (c4 plates) has been recovered for different
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB10F8E5607BF8C4" box="[636,883,1879,1905]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B92E013D9A1ED464DB10F8E5601BF8C4" box="[636,787,1879,1905]" italics="true" pageId="1" pageNumber="78">Bouchardium</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
In particular, the c4 plates of the newly described
|
||
<taxonomicName id="4C5AA6AC9A1ED464D9FBF8C96256F820" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[151,350,1915,1941]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="chillygonzalesi" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1ED464D9FBF8C96256F820" box="[151,350,1915,1941]" italics="true" pageId="1" pageNumber="78">B. chillygonzalesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1ED464D808F8C962B5F820" bold="true" box="[356,445,1915,1941]" pageId="1" pageNumber="78">
|
||
<taxonomicNameLabel id="A21DBC469A1ED464D808F8C962B5F820" box="[356,445,1915,1941]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are relatively short and only slightly projected above the outline of the c3 plate (
|
||
<figureCitation id="1361C1AA9A1ED464DC53F8C963A4F80C" captionStart="FIGURE 4" captionStartId="6.[151,250,1414,1438]" captionTargetBox="[159,1430,389,1382]" captionTargetId="figure-97@6.[151,1436,381,1390]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of female terminalia within the genus Bouchardium gen. nov. Bouchardium chillygonzalesi sp. nov. (A, B), B. mariae sp. nov. (C), B. cubicum (D). Ovipositor (A, B, D), spiculum ventrale (C). Abbreviations: c- coxities, c2-c4- subsequent lobes of coxities, v- valvifer, ip- inner plate of paraprocts, op- outer plate of paraprocts, pa - paraprocts." figureDoi="http://doi.org/10.5281/zenodo.11084619" httpUri="https://zenodo.org/record/11084619/files/figure.png" pageId="1" pageNumber="78">Fig. 4A, B</figureCitation>
|
||
). In the case of
|
||
<taxonomicName id="4C5AA6AC9A1ED464D832F82D62D4F80C" authorityName="Kamiński" authorityYear="2024" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" box="[350,476,1951,1977]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="cubicum">
|
||
<emphasis id="B92E013D9A1ED464D832F82D62D4F80C" box="[350,476,1951,1977]" italics="true" pageId="1" pageNumber="78">B. cubicum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5AA6AC9A1ED464DB7FF82D6189F80D" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[531,641,1951,1976]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1ED464DB7FF82D6189F80D" box="[531,641,1951,1976]" italics="true" pageId="1" pageNumber="78">B. mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1ED464DBE4F82D61EAF80C" bold="true" box="[648,738,1951,1977]" pageId="1" pageNumber="78">
|
||
<taxonomicNameLabel id="A21DBC469A1ED464DBE4F82D61EAF80C" box="[648,738,1951,1977]" pageId="1" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the c4 plates are elongated (
|
||
<figureCitation id="1361C1AA9A1ED464DD4DF82D6774F80C" box="[1057,1148,1951,1977]" captionStart="FIGURE 4" captionStartId="6.[151,250,1414,1438]" captionTargetBox="[159,1430,389,1382]" captionTargetId="figure-97@6.[151,1436,381,1390]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of female terminalia within the genus Bouchardium gen. nov. Bouchardium chillygonzalesi sp. nov. (A, B), B. mariae sp. nov. (C), B. cubicum (D). Ovipositor (A, B, D), spiculum ventrale (C). Abbreviations: c- coxities, c2-c4- subsequent lobes of coxities, v- valvifer, ip- inner plate of paraprocts, op- outer plate of paraprocts, pa - paraprocts." figureDoi="http://doi.org/10.5281/zenodo.11084619" httpUri="https://zenodo.org/record/11084619/files/figure.png" pageId="1" pageNumber="78">Fig. 4D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A1ED467D9ABF87162AAFEA8" blockId="1.[151,1437,1194,2013]" lastBlockId="2.[151,1437,151,537]" lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78">
|
||
The aedeagal morphology is relatively invariable within the subtribe
|
||
<taxonomicName id="4C5AA6AC9A1ED464DA81F8716664F868" authority="(Kaminski et al. 2021)" baseAuthorityName="Kaminski" baseAuthorityYear="2021" box="[1005,1388,1987,2013]" pageId="1" pageNumber="78" rank="subTribe" subTribe="Molurina">
|
||
Molurina (
|
||
<bibRefCitation id="EFCBA0DE9A1ED464DD0BF871666BF868" author="Kaminski, M. J. & Gearner, O. M. & Kanda, K & Swichtenberg, K. & Purchart, L. & Smith, A. D." box="[1127,1379,1987,2013]" pageId="1" pageNumber="78" pagination="883 - 901" refId="ref6154" refString="Kaminski, M. J., Gearner, O. M., Kanda, K .., Swichtenberg, K., Purchart, L. & Smith, A. D. (2021) First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina), and examination of the status of the Psammodes vialis species-group. Zoological Journal of the Linnean Society, 191, 883 - 901. https: // doi. org / 10.1093 / zoolinnean / zlaa 052" type="journal article" year="2021">
|
||
Kamiński
|
||
<emphasis id="B92E013D9A1ED464DD8DF8716615F868" box="[1249,1309,1987,2013]" italics="true" pageId="1" pageNumber="78">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Up to now, the shape of male terminalia has not been used for diagnostic purposes at the species or generic levels. Morphology of aedeagal tegmen and spiculum gastrale of
|
||
<taxonomicName id="4C5AA6AC9A1DD467DA41FF096713FF61" authorityName="Kamiński" authorityYear="2024" box="[813,1051,187,213]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="mariae">
|
||
<emphasis id="B92E013D9A1DD467DA41FF096713FF61" box="[813,1051,187,213]" italics="true" pageId="2" pageNumber="79">Bouchardium mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1DD467DD4FFF09677FFF60" bold="true" box="[1059,1143,187,213]" pageId="2" pageNumber="79">sp nov.</emphasis>
|
||
is illustrated in
|
||
<figureCitation id="1361C1AA9A1DD467DC5CFF096690FF60" box="[1328,1432,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1685,1709]" captionTargetBox="[159,1428,309,1652]" captionTargetId="figure-67@4.[151,1436,309,1661]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." figureDoi="http://doi.org/10.5281/zenodo.11084610" httpUri="https://zenodo.org/record/11084610/files/figure.png" pageId="2" pageNumber="79">Fig. 2I, J</figureCitation>
|
||
. Present investigation did not reveal any substantial differences in male terminalia morphology between that species and its other congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A1DD467D9ABFE946771FED0" pageId="2" pageNumber="79" type="distribution">
|
||
<paragraph id="8BE5DD2F9A1DD467D9ABFE946771FED0" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="79">
|
||
<emphasis id="B92E013D9A1DD467D9ABFE94625BFEF4" bold="true" box="[199,339,294,321]" italics="true" pageId="2" pageNumber="79">Distribution</emphasis>
|
||
. Botswana (Eastern part of the Country), Mozambique, South Africa (Provinces: Gauteng, Limpopo, Mpumalanga, North West), Zimbabwe (Southern part of the Country) (
|
||
<figureCitation id="1361C1AA9A1DD467DD4BFEF96763FED0" box="[1063,1131,330,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1305,1329]" captionTargetBox="[151,1436,181,1281]" captionTargetId="figure-15@7.[151,1436,181,1281]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of the genus Bouchardium gen. nov. and its representatives." figureDoi="http://doi.org/10.5281/zenodo.11084621" httpUri="https://zenodo.org/record/11084621/files/figure.png" pageId="2" pageNumber="79">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A1DD467D9ABFEDD662EFE64" pageId="2" pageNumber="79" type="etymology">
|
||
<paragraph id="8BE5DD2F9A1DD467D9ABFEDD662EFE64" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="79">
|
||
<emphasis id="B92E013D9A1DD467D9ABFEDD6246FE3C" bold="true" box="[199,334,367,393]" pageId="2" pageNumber="79">Etymology.</emphasis>
|
||
This genus is named in honor of Dr. Patrice Bouchard (Canadian National Collection of Insects, Arachnids and Nematodes,Agriculture andAgri-Food Canada,Ottawa,Ontario,Canada), a specialist in
|
||
<taxonomicName id="4C5AA6AC9A1DD467DD90FE216694FE18" authorityName="Latreille" authorityYear="1802" box="[1276,1436,403,429]" class="Insecta" family="Tenebrionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="family">Tenebrionidae</taxonomicName>
|
||
and zoological nomenclature in gratitude for his professional support during the first years of my career.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A1DD467D9ABFE6963F9FDAC" pageId="2" pageNumber="79" type="taxon_list">
|
||
<paragraph id="8BE5DD2F9A1DD467D9ABFE6963F9FDAC" blockId="2.[151,1437,151,537]" pageId="2" pageNumber="79">
|
||
<emphasis id="B92E013D9A1DD467D9ABFE6962A7FE41" bold="true" box="[199,431,474,501]" pageId="2" pageNumber="79">
|
||
<emphasis id="B92E013D9A1DD467D9ABFE696280FE40" bold="true" box="[199,392,474,501]" italics="true" pageId="2" pageNumber="79">Species included</emphasis>
|
||
(3)
|
||
</emphasis>
|
||
.
|
||
<taxonomicName id="4C5AA6AC9A1DD467D8D1FE69618CFE40" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[445,644,475,501]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="chillygonzalesi" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1DD467D8D1FE69618CFE40" box="[445,644,475,501]" italics="true" pageId="2" pageNumber="79">B. chillygonzalesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1DD467DBE7FE6961ECFE40" bold="true" box="[651,740,475,501]" pageId="2" pageNumber="79">
|
||
<taxonomicNameLabel id="A21DBC469A1DD467DBE7FE6961ECFE40" box="[651,740,475,501]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1DD467DB9EFE696790FE40" authority="(Guerin-Meneville, 1845)" authorityName="Kamiński" authorityYear="2024" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" box="[754,1176,474,501]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="cubicum" status="comb. nov.">
|
||
<emphasis id="B92E013D9A1DD467DB9EFE696078FE40" box="[754,880,475,501]" italics="true" pageId="2" pageNumber="79">B. cubicum</emphasis>
|
||
(Guérin-Méneville, 1845)
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1DD467DDF3FE696628FE40" bold="true" box="[1183,1312,475,501]" pageId="2" pageNumber="79">
|
||
<taxonomicNameLabel id="A21DBC469A1DD467DDF3FE696628FE40" box="[1183,1312,475,501]" pageId="2" pageNumber="79" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C5AA6AC9A1DD467DC42FE696694FE41" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[1326,1436,475,500]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1DD467DC42FE696694FE41" box="[1326,1436,475,500]" italics="true" pageId="2" pageNumber="79">B. mariae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92E013D9A1DD467D9FBFE4D63F9FDAC" bold="true" box="[151,241,511,537]" pageId="2" pageNumber="79">
|
||
<taxonomicNameLabel id="A21DBC469A1DD467D9FBFE4D63F9FDAC" box="[151,241,511,537]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A1DD467D9FBFDD9614DFD30" box="[151,581,618,645]" pageId="2" pageNumber="79" type="multiple">
|
||
<paragraph id="8BE5DD2F9A1DD467D9FBFDD9614DFD30" blockId="2.[151,581,618,645]" box="[151,581,618,645]" pageId="2" pageNumber="79">
|
||
<heading id="D0AD6A439A1DD467D9FBFDD9614DFD30" bold="true" box="[151,581,618,645]" fontSize="11" level="1" pageId="2" pageNumber="79" reason="1">
|
||
<emphasis id="B92E013D9A1DD467D9FBFDD9614DFD30" bold="true" box="[151,581,618,645]" pageId="2" pageNumber="79">
|
||
Key to the species of
|
||
<taxonomicName id="4C5AA6AC9A1DD467D8F1FDD8614DFD30" authorityName="Kamiński" authorityYear="2024" box="[413,581,618,645]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92E013D9A1DD467D8F1FDD8614DFD30" bold="true" box="[413,581,618,645]" italics="true" pageId="2" pageNumber="79">Bouchardium</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3408EA49A1DD467D9FBFD026694FCC0" pageId="2" pageNumber="79" type="key">
|
||
<keyStep id="30AEC58A9A1DD467D9FBFD026694FC8F" pageId="2" pageNumber="79">
|
||
<paragraph id="8BE5DD2F9A1DD467D9FBFD026694FD4A" blockId="2.[151,1437,688,886]" pageId="2" pageNumber="79">
|
||
<keyLead id="30AB7E1A9A1DD467D9FBFD026694FD4A" pageId="2" pageNumber="79">
|
||
1. Prosternal process with elongate appendages (
|
||
<figureCitation id="1361C1AA9A1DD467DBF0FD0261EDFD70" box="[668,741,688,710]" captionStart="FIGURE 1" captionStartId="3.[151,250,1377,1401]" captionTargetBox="[155,1429,208,1352]" captionTargetId="figure-15@3.[151,1436,181,1353]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." figureDoi="http://doi.org/10.5281/zenodo.11084606" httpUri="https://zenodo.org/record/11084606/files/figure.png" pageId="2" pageNumber="79">Fig. 1E</figureCitation>
|
||
). Elytral disc relatively narrow leaving sides of the epipleura exposed in dorsal view (
|
||
<figureCitation id="1361C1AA9A1DD467D832FD7F62A1FD57" box="[350,425,717,739]" captionStart="FIGURE 1" captionStartId="3.[151,250,1377,1401]" captionTargetBox="[155,1429,208,1352]" captionTargetId="figure-15@3.[151,1436,181,1353]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." figureDoi="http://doi.org/10.5281/zenodo.11084606" httpUri="https://zenodo.org/record/11084606/files/figure.png" pageId="2" pageNumber="79">Fig. 1B</figureCitation>
|
||
). Ovipositor with short c4 plates (not extending beyond the tip of c3) (
|
||
<figureCitation id="1361C1AA9A1DD467DD3CFD7F67BFFD57" box="[1104,1207,717,739]" captionStart="FIGURE 4" captionStartId="6.[151,250,1414,1438]" captionTargetBox="[159,1430,389,1382]" captionTargetId="figure-97@6.[151,1436,381,1390]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of female terminalia within the genus Bouchardium gen. nov. Bouchardium chillygonzalesi sp. nov. (A, B), B. mariae sp. nov. (C), B. cubicum (D). Ovipositor (A, B, D), spiculum ventrale (C). Abbreviations: c- coxities, c2-c4- subsequent lobes of coxities, v- valvifer, ip- inner plate of paraprocts, op- outer plate of paraprocts, pa - paraprocts." figureDoi="http://doi.org/10.5281/zenodo.11084619" httpUri="https://zenodo.org/record/11084619/files/figure.png" pageId="2" pageNumber="79">Fig. 4A, B</figureCitation>
|
||
).................................................................................................
|
||
<emphasis id="B92E013D9A1DD467DDF7FD596694FD4A" bold="true" box="[1179,1436,746,768]" pageId="2" pageNumber="79">
|
||
<taxonomicName id="4C5AA6AC9A1DD467DDF7FD596642FCB5" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[1179,1354,746,768]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="chillygonzalesi" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1DD467DDF7FD596642FCB5" bold="true" box="[1179,1354,746,768]" italics="true" pageId="2" pageNumber="79">B. chillygonzalesi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DBC469A1DD467DC3CFD586694FD4A" box="[1360,1436,746,767]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A1DD467D9FBFCBA6694FC8F" blockId="2.[151,1437,688,886]" pageId="2" pageNumber="79">
|
||
<keyLead id="30AB7E1A9A1DD467D9FBFCBA6694FC8F" pageId="2" pageNumber="79">
|
||
-. Prosternal process rounded (in lateral view) (
|
||
<figureCitation id="1361C1AA9A1DD467DBF4FCBA61EFFCA8" box="[664,743,776,798]" captionStart="FIGURE 2" captionStartId="4.[151,250,1685,1709]" captionTargetBox="[159,1428,309,1652]" captionTargetId="figure-67@4.[151,1436,309,1661]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." figureDoi="http://doi.org/10.5281/zenodo.11084610" httpUri="https://zenodo.org/record/11084610/files/figure.png" pageId="2" pageNumber="79">Fig. 2G</figureCitation>
|
||
). Elytral disc relatively wide covering sides of epipleura in dorsal view (
|
||
<figureCitation id="1361C1AA9A1DD467D985FC97623CFC8F" box="[233,308,805,827]" captionStart="FIGURE 1" captionStartId="3.[151,250,1377,1401]" captionTargetBox="[155,1429,208,1352]" captionTargetId="figure-15@3.[151,1436,181,1353]" captionTargetPageId="3" captionText="FIGURE 1. Morphology of Bouchardium chillygonzalesi sp. nov. (holotype). Lateral (A), dorsal (B), posterior (C), and ventral (D) views of the body; prosternal process (E); epipleuron (F); label (G). Arrow indicates edge of epipleuron." figureDoi="http://doi.org/10.5281/zenodo.11084606" httpUri="https://zenodo.org/record/11084606/files/figure.png" pageId="2" pageNumber="79">Fig. 1B</figureCitation>
|
||
). Ovipositor with elongate c4 plates (extending beyond the tip of c3) (
|
||
<figureCitation id="1361C1AA9A1DD467DABBFC97672AFC8F" box="[983,1058,805,827]" captionStart="FIGURE 4" captionStartId="6.[151,250,1414,1438]" captionTargetBox="[159,1430,389,1382]" captionTargetId="figure-97@6.[151,1436,381,1390]" captionTargetPageId="6" captionText="FIGURE 4. Morphology of female terminalia within the genus Bouchardium gen. nov. Bouchardium chillygonzalesi sp. nov. (A, B), B. mariae sp. nov. (C), B. cubicum (D). Ovipositor (A, B, D), spiculum ventrale (C). Abbreviations: c- coxities, c2-c4- subsequent lobes of coxities, v- valvifer, ip- inner plate of paraprocts, op- outer plate of paraprocts, pa - paraprocts." figureDoi="http://doi.org/10.5281/zenodo.11084619" httpUri="https://zenodo.org/record/11084619/files/figure.png" pageId="2" pageNumber="79">Fig. 4D</figureCitation>
|
||
)............................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30AEC58A9A1DD467D9FBFCF06694FCC0" pageId="2" pageNumber="79">
|
||
<paragraph id="8BE5DD2F9A1DD467D9FBFCF06694FCE2" blockId="2.[151,1437,688,886]" box="[151,1436,834,856]" pageId="2" pageNumber="79">
|
||
<keyLead id="30AB7E1A9A1DD467D9FBFCF06694FCE2" box="[151,1436,834,856]" pageId="2" pageNumber="79">
|
||
2. Elytra entirely black. Elytral disc relatively flat (
|
||
<figureCitation id="1361C1AA9A1DD467DBD8FCF061F3FCE2" box="[692,763,834,856]" captionStart="FIGURE 3" captionStartId="5.[151,250,1404,1428]" captionTargetBox="[173,1414,237,1367]" captionTargetId="figure-41@5.[164,1423,237,1381]" captionTargetPageId="5" captionText="FIGURE 3. Morphology of selected species representing ‘Dichtha clade’: Bouchardium cubicum (A, D–F), Amiantus gibbosus Fåhraeus, 1870 (B), Bouchardium mariae (C), Dichtha inflata (Gerstaecker, 1854) (G, H), Dichtha quedenfeldti Kolbe, 1886 (I, J), Distretus (Perdistretus) mormolyce Koch, 1953, holotype (K). Pronotum (D, G, I); posterior (E, H, J), lateral (F), and ventral (K) views of the body." figureDoi="http://doi.org/10.5281/zenodo.11084614" httpUri="https://zenodo.org/record/11084614/files/figure.png" pageId="2" pageNumber="79">Fig. 3E</figureCitation>
|
||
)....................................
|
||
<emphasis id="B92E013D9A1DD467DDDBFCF16694FCE2" bold="true" box="[1207,1436,834,856]" pageId="2" pageNumber="79">
|
||
<taxonomicName id="4C5AA6AC9A1DD467DDDBFCF16620FCED" authorityName="Kamiński" authorityYear="2024" baseAuthorityName="Guerin-Meneville" baseAuthorityYear="1845" box="[1207,1320,834,856]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="cubicum" status="comb. nov.">
|
||
<emphasis id="B92E013D9A1DD467DDDBFCF16620FCED" bold="true" box="[1207,1320,834,856]" italics="true" pageId="2" pageNumber="79">B. cubicum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DBC469A1DD467DC42FCF06694FCE2" box="[1326,1436,834,855]" pageId="2" pageNumber="79" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE5DD2F9A1DD467D9FBFCD26694FCC0" blockId="2.[151,1437,688,886]" box="[151,1436,863,886]" pageId="2" pageNumber="79">
|
||
<keyLead id="30AB7E1A9A1DD467D9FBFCD26694FCC0" box="[151,1436,863,886]" pageId="2" pageNumber="79">
|
||
-. Elytra with two red stripes along the sides of the disc. Elytral disc convex (
|
||
<figureCitation id="1361C1AA9A1DD467DADEFCD2673EFCC0" box="[946,1078,864,886]" captionStart="FIGURE 2" captionStartId="4.[151,250,1685,1709]" captionTargetBox="[159,1428,309,1652]" captionTargetId="figure-67@4.[151,1436,309,1661]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of Bouchardium mariae sp. nov. (holotype and paratype). Lateral (A), dorsal (B), frontal (C), posterior (D), and ventral (E) views of the body; pronotum (F); prosternal process (G); label (H); aedeagal tegmen (I); spiculum gastrale (J)." figureDoi="http://doi.org/10.5281/zenodo.11084610" httpUri="https://zenodo.org/record/11084610/files/figure.png" pageId="2" pageNumber="79">Fig. 2A, C, D</figureCitation>
|
||
)..............
|
||
<emphasis id="B92E013D9A1DD467DD84FCD26694FCC0" bold="true" box="[1256,1436,863,885]" pageId="2" pageNumber="79">
|
||
<taxonomicName id="4C5AA6AC9A1DD467DD84FCD26641FCC1" authority="Kamiński, 2024" authorityName="Kamiński" authorityYear="2024" box="[1256,1353,863,885]" class="Insecta" family="Tenebrionidae" genus="Bouchardium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
|
||
<emphasis id="B92E013D9A1DD467DD84FCD26641FCC1" bold="true" box="[1256,1353,863,885]" italics="true" pageId="2" pageNumber="79">B. mariae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21DBC469A1DD467DC3CFCD26694FCC0" box="[1360,1436,864,885]" pageId="2" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |