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<document id="57179710CF31F37AFE4E0CFC2C1931F2" ID-CLB-Dataset="20925" ID-DOI="10.11646/zootaxa.5093.4.2" ID-GBIF-Dataset="d3abaf4f-14fd-4a8b-9362-357792cc0cbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="5912678" ID-ZooBank="440E7DD2-5736-49BB-A59E-9FA494402544" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643354187259" checkinUser="plazi" docAuthor="Daniel, Gimo M., Strümpher, Werner P. &amp; Deschodt, Christian M." docDate="2022" docId="03B087B9FFB5FFB87DD551D5686CFA74" docLanguage="en" docName="zootaxa.5093.4.2.pdf" docOrigin="Zootaxa 5093 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Silvaphilus joselmae Daniel, Strumpher &amp; Deschodt 2022, new species" docType="treatment" docUuid="F290EEB4-901D-4A68-AD23-6F35EF90E7AC" docUuidSource="ZooBank" docVersion="7" lastPageNumber="417" masterDocId="FF89FFC1FFB4FFBB7D4253136B17FFE9" masterDocTitle="An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South Africa" masterLastPageNumber="420" masterPageNumber="414" pageNumber="415" updateTime="1699304596146" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="415B313E48C6D804A15F9BB0C6B09672" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1DDF2401717A90002D0F069AFC1A5860">
<mods:title id="74987C085E01B69298BE49C0B8ED23DD">An unexpected new flightless dung beetle species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Cederberg Mountains, South Africa</mods:title>
</mods:titleInfo>
<mods:name id="00315E539A34AACF54F8012CC9080CD0" type="personal">
<mods:role id="CCF4D84326270C62F89FB171AD44F4F1">
<mods:roleTerm id="7B2CDB64B5613AA8C685FFE9A7578C5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C581084399705A33722019AE68648526">Daniel, Gimo M.</mods:namePart>
<mods:nameIdentifier id="3A133CECD97237E3FC26AD7B2ADA59FC" type="ORCID">0000-0003-3602-5034</mods:nameIdentifier>
<mods:affiliation id="6B4E14F0D8715BF8FA5069F1A4A47674">Department of Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, South Africa. &amp; Department of Biological &amp; Environmental Sciences, Walter Sisulu University, Mthatha, South Africa. gimo. daniel @ nasmus. co. za; https: // orcid. org / 0000 - 0003 - 3602 - 5034</mods:affiliation>
<mods:nameIdentifier id="BBC8AA069492C2CB1F8AE70D2AD1A7BD" type="email">gimo.daniel@nasmus.co.za</mods:nameIdentifier>
</mods:name>
<mods:name id="4E97ED9FF972FC1664483EAB942D7AF4" type="personal">
<mods:role id="93F4E515F847E4F7C0D45D69E47A9609">
<mods:roleTerm id="601EEB3A188ACC1405B942558E342732">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="78CADB77FC4C929200B5F92925283E3C">Strümpher, Werner P.</mods:namePart>
<mods:affiliation id="0B9CB3C1E37A47BDC6CB1807B02E423B">Ditsong National Museum of Natural History, PO Box 413, Pretoria 0001, South Africa.</mods:affiliation>
</mods:name>
<mods:name id="8E6ECC0F91AFD099AE60EEFA64DCBD6B" type="personal">
<mods:role id="20623B1D1A5467B73E7364A77C096851">
<mods:roleTerm id="A10961A0FF39250A1A70B0FAFA62B453">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5918DE43BDDE488FBC1EC3B2BC741564">Deschodt, Christian M.</mods:namePart>
<mods:nameIdentifier id="3E775DFF6015B60C9FA82A24D1F3F186" type="ORCID">0000-0002-3886-1813</mods:nameIdentifier>
<mods:affiliation id="93A0D5B5E4BEEB395FCAB48E38196C2F">Department of Zoology &amp; Entomology, University of Pretoria, Private Bag X 20, Hatfield, 0028 South Africa. cdeschodt @ zoology. up. ac. za; https: // orcid. org / 0000 - 0002 - 3886 - 1813</mods:affiliation>
<mods:nameIdentifier id="E09EB3185DBDBD09229AF0D24F5B3ACE" type="email">cdeschodt@zoology.up.ac.za</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0A1F7C4B706A5401A17CCD7AFCEA4F15">text</mods:typeOfResource>
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<mods:titleInfo id="1E2A275A1AD88CF5E91EC10A4C6D9FAC">
<mods:title id="F166DE7F9F47D91D6B25B19226395759">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="87118D2995F550BF17C4033F7206B9FD">
<mods:date id="EA056FD978AF08877C48BEDBE5AC3757">2022</mods:date>
<mods:detail id="F45CF6BAB5E162F423481B2D0C548076" type="pubDate">
<mods:number id="548152DA3CA4E47271BEB6A51BD97A06">2022-01-28</mods:number>
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<mods:number id="EE6DD323B152DD63631D137B6BA847D2">5093</mods:number>
</mods:detail>
<mods:detail id="36E5B8F1305CE5967EF991D51E966051" type="issue">
<mods:number id="F4DADCEB5B53A4BEE868C6F746471FD7">4</mods:number>
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<mods:classification id="32A1598B99D2E52DB724EAF1342B024B">journal article</mods:classification>
<mods:identifier id="1A7B08D7AF54764296210B5920B814C3" type="CLB-Dataset">20925</mods:identifier>
<mods:identifier id="9F450645140C770E692F3AC76099C374" type="DOI">10.11646/zootaxa.5093.4.2</mods:identifier>
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</mods:mods>
<treatment id="03B087B9FFB5FFB87DD551D5686CFA74" ID-DOI="http://doi.org/10.5281/zenodo.5914107" ID-GBIF-Taxon="192700781" ID-Zenodo-Dep="5914107" LSID="urn:lsid:zoobank.org:act:F290EEB4-901D-4A68-AD23-6F35EF90E7AC" httpUri="http://treatment.plazi.org/id/03B087B9FFB5FFB87DD551D5686CFA74" lastPageId="3" lastPageNumber="417" pageId="1" pageNumber="415">
<subSubSection id="C3036524FFB5FFBA7DD551D568ACFD09" box="[151,955,710,737]" pageId="1" pageNumber="415" type="nomenclature">
<paragraph id="8BA636AFFFB5FFBA7DD551D568ACFD09" blockId="1.[151,955,710,773]" box="[151,955,710,737]" pageId="1" pageNumber="415">
<heading id="D0EE81C3FFB5FFBA7DD551D568ACFD09" bold="true" box="[151,955,710,737]" fontSize="11" level="1" pageId="1" pageNumber="415" reason="1">
<emphasis id="B96DEABDFFB5FFBA7DD551D568ACFD09" bold="true" box="[151,955,710,737]" pageId="1" pageNumber="415">
<taxonomicName id="4C194D2CFFB5FFBA7DD551D56809FD08" ID-CoL="8V23B" authority="Daniel, Strumpher &amp; Deschodt" authorityName="Daniel, Strumpher &amp; Deschodt" authorityYear="2022" box="[151,798,710,737]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB5FFBA7DD551D56A87FD08" bold="true" box="[151,400,710,737]" italics="true" pageId="1" pageNumber="415">Silvaphilus joselmae</emphasis>
Daniel, Strümpher &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C6FFB5FFBA7E6951D468ACFD09" box="[811,955,710,737]" pageId="1" pageNumber="415" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB5FFBA7DD551F868AFFCB6" pageId="1" pageNumber="415" type="description">
<paragraph id="8BA636AFFFB5FFBA7DD551F86A41FCEC" blockId="1.[151,955,710,773]" box="[151,342,747,773]" pageId="1" pageNumber="415">
(
<figureCitation id="13222A2AFFB5FFBA7DDD51F86A0FFCEC" box="[159,280,747,773]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Figs 1A–F</figureCitation>
,
<figureCitation id="13222A2AFFB5FFBA7C6151F86A58FCEC" box="[291,335,747,773]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,260,1304,1330]" captionStartId-1="4.[151,260,1960,1986]" captionTargetBox-0="[151,1436,306,1280]" captionTargetBox-1="[339,1248,1424,1936]" captionTargetId-0="figure-85@4.[151,1436,306,1280]" captionTargetId-1="figure-125@4.[339,1248,1424,1936]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="FIGURE 2. Distribution map of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. In green, the limited extent of the Fynbos riparian vegetation following the valleys in the Cederberg Mountains as shown by Mucina &amp; Rutherford (2006)." captionText-1="FIGURE 3: The type locality of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. Photo by W.P. Strümpher, 9. viii. 2021." figureDoi-0="http://doi.org/10.5281/zenodo.5912684" figureDoi-1="http://doi.org/10.5281/zenodo.5912686" httpUri-0="https://zenodo.org/record/5912684/files/figure.png" httpUri-1="https://zenodo.org/record/5912686/files/figure.png" pageId="1" pageNumber="415">2–3</figureCitation>
)
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7DD5503B6A4EFCA9" blockId="1.[151,952,808,863]" box="[151,345,808,832]" pageId="1" pageNumber="415">Nomenclatural act:</paragraph>
<paragraph id="8BA636AFFFB5FFBA7DD5505468AFFCB6" blockId="1.[151,952,808,863]" box="[151,952,839,863]" pageId="1" pageNumber="415">
LSIDurn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB5FFBA7DD550946E84FB90" pageId="1" pageNumber="415" type="materials_examined">
<paragraph id="8BA636AFFFB5FFBA7DD55094694DFC2C" blockId="1.[151,1436,902,1145]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7DD550946A20FC49" bold="true" box="[151,311,902,929]" pageId="1" pageNumber="415">Type locality.</emphasis>
<materialsCitation id="3B713CF2FFB5FFBA7C0250946941FC2C" ID-GBIF-Occurrence="3458736302" country="Algeria" latitude="-32.408337" location="Cederberg Wilderness Area" longLatPrecision="20" longitude="19.084167" municipality="Forest Station" pageId="1" pageNumber="415" specimenCount="1" stateProvince="Western Cape">
<quantity id="4CE19B4AFFB5FFBA7C0250946A6DFC48" box="[320,378,903,929]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="415" unit="km" value="4.0">4 km</quantity>
SE of
<collectingCountry id="F30E763FFFB5FFBA7C8850946909FC48" box="[458,542,903,929]" name="Algeria" pageId="1" pageNumber="415">Algeria</collectingCountry>
<collectingMunicipality id="6BC2ACD5FFB5FFBA7F65509469D2FC48" box="[551,709,903,929]" pageId="1" pageNumber="415">Forest Station</collectingMunicipality>
(
<geoCoordinate id="EE2D5068FFB5FFBA7F94509B684FFC4A" box="[726,856,904,931]" degrees="32" direction="south" minutes="24" orientation="latitude" pageId="1" pageNumber="415" precision="15" seconds="30" value="-32.408337">32°24′30″S</geoCoordinate>
<geoCoordinate id="EE2D5068FFB5FFBA7E22509B68F1FC4A" box="[864,998,904,931]" degrees="19" direction="east" minutes="05" orientation="longitude" pageId="1" pageNumber="415" precision="15" seconds="03" value="19.084167">19°05′03″E</geoCoordinate>
),
<location id="8EC66074FFB5FFBA7EB950946E22FC48" LSID="urn:lsid:plazi:treatment:03B087B9FFB5FFB87DD551D5686CFA74:8EC66074FFB5FFBA7EB950946E22FC48" box="[1019,1333,903,929]" country="Algeria" latitude="-32.408337" longLatPrecision="20" longitude="19.084167" municipality="Forest Station" name="Cederberg Wilderness Area" pageId="1" pageNumber="415" stateProvince="Western Cape">Cederberg Wilderness Area</location>
,
<collectingRegion id="49DDF84DFFB5FFBA780350946A29FC2C" country="South Africa" name="Western Cape" pageId="1" pageNumber="415">Western Cape Province</collectingRegion>
,
<collectingCountry id="F30E763FFFB5FFBA7C0B50B86ACFFC2C" box="[329,472,939,965]" name="South Africa" pageId="1" pageNumber="415">South Africa</collectingCountry>
(
<figureCitation id="13222A2AFFB5FFBA7CA450B8695AFC2C" box="[486,589,939,965]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,260,1304,1330]" captionStartId-1="4.[151,260,1960,1986]" captionTargetBox-0="[151,1436,306,1280]" captionTargetBox-1="[339,1248,1424,1936]" captionTargetId-0="figure-85@4.[151,1436,306,1280]" captionTargetId-1="figure-125@4.[339,1248,1424,1936]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="FIGURE 2. Distribution map of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. In green, the limited extent of the Fynbos riparian vegetation following the valleys in the Cederberg Mountains as shown by Mucina &amp; Rutherford (2006)." captionText-1="FIGURE 3: The type locality of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. Photo by W.P. Strümpher, 9. viii. 2021." figureDoi-0="http://doi.org/10.5281/zenodo.5912684" figureDoi-1="http://doi.org/10.5281/zenodo.5912686" httpUri-0="https://zenodo.org/record/5912684/files/figure.png" httpUri-1="https://zenodo.org/record/5912686/files/figure.png" pageId="1" pageNumber="415">Figs 2–3</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D8550DC6E12FBBC" blockId="1.[151,1436,902,1145]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8550DC6A6BFC01" bold="true" box="[199,380,974,1001]" pageId="1" pageNumber="415">Type material.</emphasis>
<materialsCitation id="3B713CF2FFB5FFBA7CC950DC6E12FBBC" ID-GBIF-Occurrence="3458736301" collectingDate="2021-08-07" collectingDateMax="2021-08-09" collectingDateMin="2021-08-07" collectionCode="TMSA, SITE, IV" country="South Africa" elevation="790" latitude="-32.408337" location="Cederberg Wilderness Area" longLatPrecision="20" longitude="19.084167" pageId="1" pageNumber="415" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
<emphasis id="B96DEABDFFB5FFBA7CC950DC6916FC0E" bold="true" box="[395,513,975,1001]" pageId="1" pageNumber="415">
<typeStatus id="54A2880DFFB5FFBA7CC950DC6916FC0E" box="[395,513,975,1001]" pageId="1" pageNumber="415" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
,
<specimenCount id="9D1FFD26FFB5FFBA7F5450DC693CFC00" box="[534,555,975,1001]" pageId="1" pageNumber="415" type="male">♂</specimenCount>
(
<collectionCode id="ED08AE6AFFB5FFBA7F0350DC6985FC00" box="[577,658,975,1001]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="1" pageNumber="415">TMSA</collectionCode>
) (
<figureCitation id="13222A2AFFB5FFBA7FF250DC6822FC00" box="[688,821,975,1001]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Figs 1A–F</figureCitation>
): “
<collectingCountry id="F30E763FFFB5FFBA7E1950DC68EAFC0E" box="[859,1021,975,1001]" name="South Africa" pageId="1" pageNumber="415">SOUTH AFRICA</collectingCountry>
:
<collectingRegion id="49DDF84DFFB5FFBA795150DC6E15FC00" box="[1043,1282,975,1001]" country="South Africa" name="Western Cape" pageId="1" pageNumber="415">Western Cape Prov.</collectingRegion>
, |
<location id="8EC66074FFB5FFBA786A50DC6A43FBE4" LSID="urn:lsid:plazi:treatment:03B087B9FFB5FFB87DD551D5686CFA74:8EC66074FFB5FFBA786A50DC6A43FBE4" country="South Africa" latitude="-32.408337" longLatPrecision="20" longitude="19.084167" name="Cederberg Wilderness Area" pageId="1" pageNumber="415" stateProvince="Western Cape">Cederberg Wilderness Area</location>
, |
<locationDeviation id="9A142F8DFFB5FFBA7C2C50E069E2FBE4" box="[366,757,1011,1037]" location="Algeria Forest Station" pageId="1" pageNumber="415">
<quantity id="4CE19B4AFFB5FFBA7C2C50E06ABFFBE4" box="[366,424,1011,1037]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="415" unit="km" value="4.0">4 km</quantity>
SE of Algeria Forest Station
</locationDeviation>
,” || “
<date id="FFA7106FFFB5FFBA7E6B50E068AAFBE4" box="[809,957,1011,1037]" pageId="1" pageNumber="415" value="2021-08-07" valueMax="2021-08-09" valueMin="2021-08-07">
<collectingDate id="EFE3E987FFB5FFBA7E6B50E068AAFBE4" box="[809,957,1011,1037]" pageId="1" pageNumber="415" value="2021-08-07" valueMax="2021-08-09" valueMin="2021-08-07">7-9.viii.2021</collectingDate>
</date>
, altitude
<quantity id="4CE19B4AFFB5FFBA796450E06F67FBE4" box="[1062,1136,1011,1037]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="415" unit="m" value="790.0">
<elevation id="0034D19CFFB5FFBA796450E06F67FBE4" box="[1062,1136,1011,1037]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="415" unit="m" value="790.0">790 m</elevation>
</quantity>
, |
<geoCoordinate id="EE2D5068FFB5FFBA79C950E76E1CFBE6" box="[1163,1291,1012,1039]" degrees="32" direction="south" minutes="24" orientation="latitude" pageId="1" pageNumber="415" precision="15" seconds="30" value="-32.408337">32°24′30″S</geoCoordinate>
<geoCoordinate id="EE2D5068FFB5FFBA785150E76E8FFBE6" box="[1299,1432,1012,1039]" degrees="19" direction="east" minutes="05" orientation="longitude" pageId="1" pageNumber="415" precision="15" seconds="03" value="19.084167">19°05′03″E</geoCoordinate>
, | GM Daniel &amp; WP Strümpher” || &quot;
<collectionCode id="ED08AE6AFFB5FFBA7F6A57046968FBD8" box="[552,639,1047,1073]" pageId="1" pageNumber="415">SITE IV</collectionCode>
:
<quantity id="4CE19B4AFFB5FFBA7FC9570469C2FBD8" box="[651,725,1047,1073]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="415" unit="m" value="790.0">
<elevation id="0034D19CFFB5FFBA7FC9570469C2FBD8" box="[651,725,1047,1073]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="415" unit="m" value="790.0">790 m</elevation>
</quantity>
, Shaded vegetation, | near stream,
<collectingMethod id="52584EB8FFB5FFBA792157046FCCFBD8" box="[1123,1243,1047,1073]" pageId="1" pageNumber="415">pitfall trap</collectingMethod>
| baited with pig dung.” || “
<emphasis id="B96DEABDFFB5FFBA7C4A57286A69FBBA" bold="true" box="[264,382,1083,1109]" pageId="1" pageNumber="415">
<typeStatus id="54A2880DFFB5FFBA7C4A57286A69FBBA" box="[264,382,1083,1109]" pageId="1" pageNumber="415" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
|
<taxonomicName id="4C194D2CFFB5FFBA7CD357296968FBBC" authorityName="Daniel, Strumpher &amp; Deschodt" authorityYear="2022" box="[401,639,1082,1109]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="415" phylum="Arthropoda" rank="species" species="joselmae">
<emphasis id="B96DEABDFFB5FFBA7CD357296968FBBC" bold="true" box="[401,639,1082,1109]" italics="true" pageId="1" pageNumber="415">Silvaphilus joselmae</emphasis>
</taxonomicName>
| Daniel, Strümpher &amp; | Deschodt, 2021 [red cardstock]”.
</materialsCitation>
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D85574C6E84FB90" blockId="1.[151,1436,902,1145]" box="[199,1427,1118,1145]" pageId="1" pageNumber="415">
<materialsCitation id="3B713CF2FFB5FFBA7D85574C6E98FB90" ID-GBIF-Occurrence="3458736303" box="[199,1423,1118,1145]" collectingDate="2021-08-07" collectingDateMax="2021-08-09" collectingDateMin="2021-08-07" collectionCode="BMSA, TMSA, SANC" country="South Africa" elevation="790" latitude="-32.408337" location="Cederberg Wilderness Area" longLatPrecision="20" longitude="19.084167" pageId="1" pageNumber="415" specimenCount="56" specimenCount-female="36" specimenCount-male="20" stateProvince="Western Cape" typeStatus="paratype">
<emphasis id="B96DEABDFFB5FFBA7D85574C6A55FB9E" bold="true" box="[199,322,1119,1145]" pageId="1" pageNumber="415">
<typeStatus id="54A2880DFFB5FFBA7D85574C6A55FB9E" box="[199,322,1119,1145]" pageId="1" pageNumber="415" type="paratype">PARATYPES</typeStatus>
</emphasis>
,
<specimenCount id="9D1FFD26FFB5FFBA7C12574C6A8EFB90" box="[336,409,1119,1145]" pageId="1" pageNumber="415" type="male">10♂♂</specimenCount>
,
<specimenCount id="9D1FFD26FFB5FFBA7CE6574C6AFDFB90" box="[420,490,1119,1145]" pageId="1" pageNumber="415" type="female">18♀♀</specimenCount>
(
<collectionCode id="ED08AE6AFFB5FFBA7CBA574C695FFB90" box="[504,584,1119,1145]" country="South Africa" httpUri="http://grbio.org/cool/7is6-1zj2" name="National Museum Bloemfontein" pageId="1" pageNumber="415">BMSA</collectionCode>
<specimenCount id="9D1FFD26FFB5FFBA7F0D574C6960FB90" box="[591,631,1119,1145]" pageId="1" pageNumber="415" type="male">4♂</specimenCount>
<specimenCount id="9D1FFD26FFB5FFBA7F35574C698FFB90" box="[631,664,1119,1145]" pageId="1" pageNumber="415" type="female">6♀</specimenCount>
;
<collectionCode id="ED08AE6AFFB5FFBA7FE1574C69F8FB90" box="[675,751,1119,1145]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="1" pageNumber="415">TMSA</collectionCode>
<specimenCount id="9D1FFD26FFB5FFBA7FB5574D6809FB90" box="[759,798,1118,1145]" pageId="1" pageNumber="415" type="male">5♂</specimenCount>
<specimenCount id="9D1FFD26FFB5FFBA7E5C574D6828FB90" box="[798,831,1118,1145]" pageId="1" pageNumber="415" type="female">9♀</specimenCount>
;
<collectionCode id="ED08AE6AFFB5FFBA7E09574C6881FB90" box="[843,918,1119,1145]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" lsid="urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="1" pageNumber="415" type="Museum">SANC</collectionCode>
<specimenCount id="9D1FFD26FFB5FFBA7EDF574C68D5FB90" box="[925,962,1119,1145]" pageId="1" pageNumber="415" type="male">1♂</specimenCount>
;
<specimenCount id="9D1FFD26FFB5FFBA7E8D574C6F10FB90" box="[975,1031,1119,1145]" pageId="1" pageNumber="415" type="female">3♀♀</specimenCount>
); same collection data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB5FFB87DD557B36956FCD0" lastPageId="3" lastPageNumber="417" pageId="1" pageNumber="415" type="description">
<paragraph id="8BA636AFFFB5FFBA7DD557B36AF4FB52" blockId="1.[151,1437,1184,2039]" box="[151,483,1184,1211]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7DD557B36AA9FB53" bold="true" box="[151,446,1184,1210]" pageId="1" pageNumber="415">
Description of
<typeStatus id="54A2880DFFB5FFBA7C0557B36AB9FB53" box="[327,430,1184,1210]" pageId="1" pageNumber="415" type="holotype">holotype</typeStatus>
(
</emphasis>
<specimenCount id="9D1FFD26FFB5FFBA7CFC57B26AC2FB52" box="[446,469,1185,1211]" pageId="1" pageNumber="415" type="male">♂</specimenCount>
).
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D8557D76960FB36" blockId="1.[151,1437,1184,2039]" box="[199,631,1220,1247]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8557D76BE9FB37" box="[199,254,1220,1246]" italics="true" pageId="1" pageNumber="415">Size:</emphasis>
Length:
<quantity id="4CE19B4AFFB5FFBA7C2157D76AABFB36" box="[355,444,1220,1247]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="415" unit="mm" value="3.4">3.4 mm</quantity>
. Width:
<quantity id="4CE19B4AFFB5FFBA7F5B57D76965FB36" box="[537,626,1220,1247]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="415" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D8557FB6E8BFAEA" blockId="1.[151,1437,1184,2039]" box="[199,1436,1256,1283]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8557FB6A01FAEB" box="[199,278,1256,1282]" italics="true" pageId="1" pageNumber="415">Colour</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C6757FB6AB2FAEB" box="[293,421,1256,1282]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Figs 1A–B</figureCitation>
): Black; mouthparts and antenna yellowish. Body surface covered with recurved tan setae.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D85561F6F62FACE" blockId="1.[151,1437,1184,2039]" box="[199,1141,1292,1319]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D85561F6A5EFACF" box="[199,329,1292,1318]" italics="true" pageId="1" pageNumber="415">Body shape</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C1A561F6AA5FACF" box="[344,434,1292,1318]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Fig. 1A</figureCitation>
): Round (in dorsal view) and strongly convex (in lateral view).
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D85562369F7FA32" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8556236A13FAA3" box="[199,260,1328,1354]" italics="true" pageId="1" pageNumber="415">Head</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C5456236A8DFAA3" box="[278,410,1328,1354]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Figs 1A–B</figureCitation>
): Rounded, surface densely and coarsely punctate with few scattered tan setae. Clypeus broadly rounded, somewhat depressed anterolaterally. Anterior clypeal margin bidentate; edge between denticles broadly W-shaped with small triangular medial denticle present; lateral clypeal margin continuous; fronto-clypeal suture indistinct. Clypeo-genal margin continuously rounded. Dorsal part of eyes elongated and curved. Antennae with nine antennomeres; mouthparts with few setae.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D8556F669B9F9AF" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8556F66A21FA17" box="[199,310,1509,1534]" italics="true" pageId="1" pageNumber="415">Pronotum</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C0756F76A8AFA17" box="[325,413,1508,1534]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Fig. 1A</figureCitation>
): Strongly convex, wider than long, coarsely punctate, some punctures bearing recurved tan setae; basal margin with single clear line of uniform elliptical punctures arranged in parallel; basal margin wider than apical; lateral margin somewhat explanate.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D8555436F42F8F7" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D8555436A1AF983" box="[199,269,1616,1642]" italics="true" pageId="1" pageNumber="415">Elytra</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C5F55436A88F982" box="[285,415,1616,1643]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Figs 1A, D</figureCitation>
): Strongly convex, fused, with nine striae. Striae 1–7 formed by continuous row of double semi-circles; stria 8 with a carina on either side, modified as deep and obvious groove; stria 9 adjacent to epipleuron. Epipleuron surface smooth, well-defined, wider basally and narrower apically. Elytral interstriae with single row of recurved tan setae paralleling outer edge of each interval; seta spaced every four or five punctures. Medial edges impunctate; interstriae surface densely punctate on first basal half, while on second apical half punctate only on lateral edges. Elytral humerus and apical callosity absent. Metathoracic wings absent.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D85543A6E15F8AA" blockId="1.[151,1437,1184,2039]" box="[199,1282,1832,1859]" pageId="1" pageNumber="415">
<taxonomicName id="4C194D2CFFB5FFBA7D85543A6A5DF8AB" box="[199,330,1833,1858]" class="Insecta" family="Elateridae" genus="Prosternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEABDFFB5FFBA7D85543A6A5DF8AB" box="[199,330,1833,1858]" italics="true" pageId="1" pageNumber="415">Prosternum</emphasis>
</taxonomicName>
and
<emphasis id="B96DEABDFFB5FFBA7CC3543B6AEAF8AB" box="[385,509,1832,1858]" italics="true" pageId="1" pageNumber="415">propleuron</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7F4E543B6974F8AB" box="[524,611,1832,1858]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Fig. 1B</figureCitation>
): Surface dull, with few irregular sets of ocellate punctures.
</paragraph>
<paragraph id="8BA636AFFFB5FFBA7D85545E6F87F81E" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="415">
<emphasis id="B96DEABDFFB5FFBA7D85545E6A3EF88F" box="[199,297,1869,1894]" italics="true" pageId="1" pageNumber="415">Ventrites</emphasis>
(
<figureCitation id="13222A2AFFB5FFBA7C7B545F6A87F88F" box="[313,400,1868,1894]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="1" pageNumber="415">Fig. 1B</figureCitation>
): Mesepimere with few punctures; surface dull. Mesoventrite with surface shining; virtually glabrous; punctures almost touching and about twice larger than those on the metaventrite. Mesometaventral suture very slightly curving anteriorly. Metaventrite surface shining; surface completely punctate with punctures separated by almost one puncture diameter, glabrous anteriorly, setose posteriorly. Six abdominal ventrites visible, punctate laterally, last abdominal ventrite twice wider than others with single row of recurved setae.
</paragraph>
<caption id="DF666627FFB6FFB97DD5544C6917F828" ID-DOI="http://doi.org/10.5281/zenodo.5912680" ID-Zenodo-Dep="5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="2" pageNumber="416" startId="2.[151,275,1887,1913]" targetBox="[160,1426,189,1855]" targetPageId="2">
<paragraph id="8BA636AFFFB6FFB97DD5544C6917F828" blockId="2.[151,1436,1887,1986]" pageId="2" pageNumber="416">
<emphasis id="B96DEABDFFB6FFB97DD5544C6A4CF890" bold="true" box="[151,347,1887,1913]" pageId="2" pageNumber="416">FIGURES 1A–F</emphasis>
.
<taxonomicName id="4C194D2CFFB6FFB97C2A544C68A5F890" authority="Daniel, Strumpher &amp; Deschodt" authorityName="Daniel, Strumpher &amp; Deschodt" authorityYear="2022" box="[360,946,1887,1914]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="416" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB6FFB97C2A544C6958F890" box="[360,591,1887,1913]" italics="true" pageId="2" pageNumber="416">Silvaphilus joselmae</emphasis>
Daniel, Strümpher &amp; Deschodt
</taxonomicName>
,
<taxonomicNameLabel id="A25E57C6FFB6FFB97EFE54736F52F890" box="[956,1093,1887,1914]" pageId="2" pageNumber="416" rank="species">new species</taxonomicNameLabel>
, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels.
</paragraph>
</caption>
<paragraph id="8BA636AFFFB7FFB87D8553846E8FFF3C" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8553846A27FF58" box="[199,304,151,177]" italics="true" pageId="3" pageNumber="417">Pygidium</emphasis>
(
<figureCitation id="13222A2AFFB7FFB87C0153846A8CFF58" box="[323,411,151,177]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="3" pageNumber="417">Fig. 1C</figureCitation>
): About twice as wide as high, surface shagreened with punctures, lateral edges with few punctures, each bearing tan seta. Complete and strongly marginate, marginal bead obtusely angulate at basal midline.
</paragraph>
<paragraph id="8BA636AFFFB7FFB87D8553CC6AA5FD24" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8553CC6BEBFF11" box="[199,252,223,248]" italics="true" pageId="3" pageNumber="417">Legs</emphasis>
(
<figureCitation id="13222A2AFFB7FFB87C4953CC6A9CFF10" box="[267,395,223,249]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="3" pageNumber="417">Figs 1A–B</figureCitation>
): Profemora ventrally punctate, with some punctures bearing seta, surface micro-shagreened, shining; upper anterior and posterior edges strongly marginate; lower anterior edge weakly marginate; distinct pit on trochanterofemoral segments present. Protibiae, slender basally and stouter apically, truncate apically with three prominent teeth on outer margin; entire margin distinctly serrated, area between teeth serrate; apical tooth largest, medial tooth somewhat smaller than basal one. In dorsal view, a row of six-aligned setae running parallel to outer margin visible; outer margin carina present, extending from basal edge of tibia to basal tooth. In ventral view, median row of seven-aligned setae running in between inner and outer margin; shallowly carinate on basal half of outer margin. Protibial spur present. Protarsi with four visible tarsomeres and tiny pair of claws; protarsomeres 1–3 triangular and short; protarsomere 4 cylindrical at least twice longer than others, with pair of tiny apical claws. Meso- and metafemora ventrally sparsely, finely punctate hind edges, punctures bearing setae, surfaces microreticulate, shiny; anterior and posterior edges marginate. Meso- and metatibiae flattened, gradually expanded nearly to apex, with prominent spur on inner margin. Meso- and metatarsomeres with five tarsomeres, protarsomeres 1–4 triangular, short, covered with few stiff setae; protarsomere 5 cylindrical at least twice longer than others, with pair of very tiny apical claws.
</paragraph>
<paragraph id="8BA636AFFFB7FFB87D8551C46FA2FD18" blockId="3.[151,1437,151,1437]" box="[199,1205,727,753]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8551C46A23FD18" box="[199,308,727,753]" italics="true" pageId="3" pageNumber="417">Aedeagus</emphasis>
(
<figureCitation id="13222A2AFFB7FFB87C0151C46A8DFD18" box="[323,410,727,753]" captionStart="FIGURES 1" captionStartId="2.[151,275,1887,1913]" captionTargetBox="[160,1426,189,1855]" captionTargetId="figure-18@2.[151,1436,179,1863]" captionTargetPageId="2" captionText="FIGURES 1A–F. Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species, holotype (TMSA). A, habitus, dorsal view; B, habitus, ventral view; C, pygidium; D, elytron, lateral view; E, aedeagus (paratype), lateral and dorsal views; F, holotype labels." figureDoi="http://doi.org/10.5281/zenodo.5912680" httpUri="https://zenodo.org/record/5912680/files/figure.png" pageId="3" pageNumber="417">Fig. 1E</figureCitation>
): Symmetrical, parameres with the apex tapered and inwardly curving.
</paragraph>
<paragraph id="8BA636AFFFB7FFB87D8551E96956FCD0" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8551E96AE3FCFD" bold="true" box="[199,500,762,788]" pageId="3" pageNumber="417">Morphological variation:</emphasis>
Length:
<quantity id="4CE19B4AFFB7FFB87F1551E869F6FCFC" box="[599,737,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="3" pageNumber="417" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.3–3.8 mm</quantity>
. Width:
<quantity id="4CE19B4AFFB7FFB87E7F51E868DAFCFC" box="[829,973,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="3" pageNumber="417" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1. 9 mm</quantity>
. There is no clear sexual dimorphism, but females are slightly larger than males.
</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB7FFB87D8550516A93FB94" pageId="3" pageNumber="417" type="diagnosis">
<paragraph id="8BA636AFFFB7FFB87D8550516A93FB94" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8550516A53FCB5" bold="true" box="[199,324,834,860]" pageId="3" pageNumber="417">Diagnosis:</emphasis>
<taxonomicName id="4C194D2CFFB7FFB87C1350506929FCB4" authority="Daniel &amp; Strümpher &amp; Deschodt, 2022" authorityName="Daniel &amp; Strümpher &amp; Deschodt" authorityYear="2022" box="[337,574,835,861]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB7FFB87C1350506929FCB4" box="[337,574,835,861]" italics="true" pageId="3" pageNumber="417">Silvaphilus joselmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C6FFB7FFB87F09505069C0FCB4" box="[587,727,835,861]" pageId="3" pageNumber="417" rank="species">new species</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C194D2CFFB7FFB8796750506FDDFCB4" authorityName="Roets &amp; Oberlander" authorityYear="2010" box="[1061,1226,835,861]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB8796750506FDDFCB4" box="[1061,1226,835,861]" italics="true" pageId="3" pageNumber="417">S. oubosiensis</emphasis>
</taxonomicName>
by the following combination of characters: males of the new species lack the lateral elytral spine which is present in males of
<taxonomicName id="4C194D2CFFB7FFB878C550746A0DFC4C" authorityName="Roets &amp; Oberlander" authorityYear="2010" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB878C550746A0DFC4C" italics="true" pageId="3" pageNumber="417">S. oubosiensis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C194D2CFFB7FFB87C6850986924FC4C" authorityName="Roets &amp; Oberlander" authorityYear="2010" box="[298,563,907,933]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB87C6850986924FC4C" box="[298,563,907,933]" italics="true" pageId="3" pageNumber="417">Silvaphilus oubosiensis</emphasis>
</taxonomicName>
shows a pronotal depression on the basal-medial edge, which is not present in the new species.
<taxonomicName id="4C194D2CFFB7FFB87C3150BC694CFC20" authority="Daniel &amp; Strümpher &amp; Deschodt, 2022" authorityName="Daniel &amp; Strümpher &amp; Deschodt" authorityYear="2022" box="[371,603,943,969]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB7FFB87C3150BC694CFC20" box="[371,603,943,969]" italics="true" pageId="3" pageNumber="417">Silvaphilus joselmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C6FFB7FFB87F2050BC69FEFC20" box="[610,745,943,969]" pageId="3" pageNumber="417" rank="species">new species</taxonomicNameLabel>
is much smaller in size than
<taxonomicName id="4C194D2CFFB7FFB8797350BC6FC6FC20" authorityName="Roets &amp; Oberlander" authorityYear="2010" box="[1073,1233,943,969]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB8797350BC6FC6FC20" box="[1073,1233,943,969]" italics="true" pageId="3" pageNumber="417">S. oubosiensis</emphasis>
</taxonomicName>
; in the former the medial width of its mesoventrite is twice wider and with two rows of punctures, in the latter the mesoventrite is narrower and with one row of punctures. Furthermore, the mesometaventral suture curves very slightly anteriorly in the new species, whereas it is rounded in
<taxonomicName id="4C194D2CFFB7FFB87FD457086820FBDC" authorityName="Roets &amp; Oberlander" authorityYear="2010" box="[662,823,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB87FD457086820FBDC" box="[662,823,1051,1077]" italics="true" pageId="3" pageNumber="417">S. oubosiensis</emphasis>
</taxonomicName>
. Lastly, the eighth stria in
<taxonomicName id="4C194D2CFFB7FFB8792F57086FF8FBDC" authority="Daniel &amp; Strümpher &amp; Deschodt, 2022" authorityName="Daniel &amp; Strümpher &amp; Deschodt" authorityYear="2022" box="[1133,1263,1051,1077]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB7FFB8792F57086FF8FBDC" box="[1133,1263,1051,1077]" italics="true" pageId="3" pageNumber="417">S. joselmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C6FFB7FFB879BA57086E96FBDC" box="[1272,1409,1051,1077]" pageId="3" pageNumber="417" rank="species">new species</taxonomicNameLabel>
is modified to form a deep and obvious groove, but in
<taxonomicName id="4C194D2CFFB7FFB87FB7572C6881FBB0" authorityName="Roets &amp; Oberlander" authorityYear="2010" box="[757,918,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="oubosiensis">
<emphasis id="B96DEABDFFB7FFB87FB7572C6881FBB0" box="[757,918,1087,1113]" italics="true" pageId="3" pageNumber="417">S. oubosiensis</emphasis>
</taxonomicName>
it is formed of deep and clear elongated and connected punctures.
</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB7FFB87D8557946895FB2C" pageId="3" pageNumber="417" type="etymology">
<paragraph id="8BA636AFFFB7FFB87D8557946895FB2C" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8557946A45FB48" bold="true" box="[199,338,1159,1185]" pageId="3" pageNumber="417">Etymology:</emphasis>
The new species is named after Joselma G. Pereira, the wife of the first author, Gimo M. Daniel, for her unconditional love and support in his entomological career.
</paragraph>
</subSubSection>
<subSubSection id="C3036524FFB7FFB87D8557DD686CFA74" pageId="3" pageNumber="417" type="distribution">
<paragraph id="8BA636AFFFB7FFB87D8557DD686CFA74" blockId="3.[151,1437,151,1437]" pageId="3" pageNumber="417">
<emphasis id="B96DEABDFFB7FFB87D8557DD6A4EFB01" bold="true" box="[199,345,1230,1256]" pageId="3" pageNumber="417">Distribution</emphasis>
(
<figureCitation id="13222A2AFFB7FFB87C2857DC6AC3FB00" box="[362,468,1231,1257]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,260,1304,1330]" captionStartId-1="4.[151,260,1960,1986]" captionTargetBox-0="[151,1436,306,1280]" captionTargetBox-1="[339,1248,1424,1936]" captionTargetId-0="figure-85@4.[151,1436,306,1280]" captionTargetId-1="figure-125@4.[339,1248,1424,1936]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="FIGURE 2. Distribution map of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. In green, the limited extent of the Fynbos riparian vegetation following the valleys in the Cederberg Mountains as shown by Mucina &amp; Rutherford (2006)." captionText-1="FIGURE 3: The type locality of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. Photo by W.P. Strümpher, 9. viii. 2021." figureDoi-0="http://doi.org/10.5281/zenodo.5912684" figureDoi-1="http://doi.org/10.5281/zenodo.5912686" httpUri-0="https://zenodo.org/record/5912684/files/figure.png" httpUri-1="https://zenodo.org/record/5912686/files/figure.png" pageId="3" pageNumber="417">Figs 2–3</figureCitation>
):
<taxonomicName id="4C194D2CFFB7FFB87CA857DC69C4FB00" authority="Daniel &amp; Strümpher &amp; Deschodt, 2022" authorityName="Daniel &amp; Strümpher &amp; Deschodt" authorityYear="2022" box="[490,723,1231,1257]" class="Insecta" family="Scarabaeidae" genus="Silvaphilus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="417" phylum="Arthropoda" rank="species" species="joselmae" status="sp. nov.">
<emphasis id="B96DEABDFFB7FFB87CA857DC69C4FB00" box="[490,723,1231,1257]" italics="true" pageId="3" pageNumber="417">Silvaphilus joselmae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25E57C6FFB7FFB87F9E57DC6872FB00" box="[732,869,1231,1257]" pageId="3" pageNumber="417" rank="species">new species</taxonomicNameLabel>
is known from a single locality in the Cederberg Mountains where it was recorded in Fynbos riparian vegetation (Aza 1;
<bibRefCitation id="EF884B5EFFB7FFB87E9157E06E18FAE7" author="Mucina, L. &amp; Rutherford, M. C." box="[979,1295,1267,1294]" pageId="3" pageNumber="417" refId="ref4652" refString="Mucina, L. &amp; Rutherford, M. C. (Eds.) (2006) The vegetation of South Africa, Lesotho and Swaziland. Strelitzi. Vol. 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) (
<figureCitation id="13222A2AFFB7FFB8786557E06E79FAE4" box="[1319,1390,1267,1293]" captionStart="FIGURE 3" captionStartId="4.[151,260,1960,1986]" captionTargetBox="[339,1248,1424,1936]" captionTargetId="figure-125@4.[339,1248,1424,1936]" captionTargetPageId="4" captionText="FIGURE 3: The type locality of Silvaphilus joselmae Daniel, Strümpher &amp; Deschodt, new species. Photo by W.P. Strümpher, 9. viii. 2021." figureDoi="http://doi.org/10.5281/zenodo.5912686" httpUri="https://zenodo.org/record/5912686/files/figure.png" pageId="3" pageNumber="417">Fig. 3</figureCitation>
). In the Cederberg area there are less than ten severely fragmented patches of the Fynbos riparian vegetation covering a total area of less than
<quantity id="4CE19B4AFFB7FFB87CE856296AE5FABC" box="[426,498,1338,1365]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="417" unit="km" value="50.0">50 km</quantity>
<superScript id="7C6C9BE7FFB7FFB87CB056296AEDFAA1" attach="left" box="[498,506,1338,1352]" fontSize="6" pageId="3" pageNumber="417">2</superScript>
. Although the vegetation unit is well protected in the Cederberg Wilderness Area dedicated collecting in the area is required to determine if this species occurs in any of the other patches of riparian scrub vegetation or if it exhibits broader vegetation specialisation.
</paragraph>
</subSubSection>
</treatment>
</document>
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