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<document id="F5CEF5C7C51647606C8276A4E3AA4529" ID-DOI="10.15298/rusentj.30.4.07" ID-ISSN="0132-8069" ID-Zenodo-Dep="13178230" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722593897160" checkinUser="felipe" docAuthor="Kataev, B. M." docDate="2021" docId="03B087A5F301FFFEE49A6C3C9B4DD87A" docLanguage="en" docName="RussEntomolJ.30.4.448-454.pdf" docOrigin="Russian Entomological Journal 30 (4)" docSource="http://dx.doi.org/10.15298/rusentj.30.4.07" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.3:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="3" docTitle="Coleolissus inessae Kataev 2021, sp.n." docType="treatment" docVersion="2" lastPageNumber="451" masterDocId="FF89FFDDF300FFFDE7076E48983BD06C" masterDocTitle="Description of a remarkable new species of the genus Coleolissus from the Western Ghats (India), with notes on this and related genera (Coleoptera: Carabidae)" masterLastPageNumber="454" masterPageNumber="448" pageNumber="449" updateTime="1722633598588" updateUser="ExternalLinkService">
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<mods:title id="22C4254E661B450F16CFCEE5BB6BA633">Description of a remarkable new species of the genus Coleolissus from the Western Ghats (India), with notes on this and related genera (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="90D17A76E65802C102D70DD1162AA0B7">Kataev, B. M.</mods:namePart>
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<mods:title id="3076427140B9A8BD8C63940E6091326E">Russian Entomological Journal</mods:title>
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<mods:part id="37336A622D49DF1024A01FEDBAD3C2FF">
<mods:date id="BEDEFFD8344C24986A6A806020B50D8A">2021</mods:date>
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<mods:number id="484420BD8FAB51BED272D541925554F2">2021-12-31</mods:number>
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<mods:number id="5EE0FA1779440ABCF269F6E9271DF872">30</mods:number>
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<mods:classification id="6B731CB8749EB63D9BDFA17314C13920">journal article</mods:classification>
<mods:identifier id="9EB34D52C1F395BD392E46897C4A6B80" type="DOI">10.15298/rusentj.30.4.07</mods:identifier>
<mods:identifier id="55B6BDFD6A67838752402AD5100901EE" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="2EAB41E0176391FF2A45470AF4707967" type="Zenodo-Dep">13178230</mods:identifier>
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<treatment id="03B087A5F301FFFEE49A6C3C9B4DD87A" LSID="urn:lsid:plazi:treatment:03B087A5F301FFFEE49A6C3C9B4DD87A" httpUri="http://treatment.plazi.org/id/03B087A5F301FFFEE49A6C3C9B4DD87A" lastPageId="3" lastPageNumber="451" pageId="1" pageNumber="449">
<subSubSection id="C3036538F301FFFCE49A6C3C9D3ED2E0" box="[925,1285,628,652]" pageId="1" pageNumber="449" type="nomenclature">
<paragraph id="8BA636B3F301FFFCE49A6C3C9D3ED2E0" blockId="1.[812,1399,628,2068]" box="[925,1285,628,652]" pageId="1" pageNumber="449">
<heading id="D0EE81DFF301FFFCE49A6C3C9D3ED2E0" box="[925,1285,628,652]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="449" reason="8">
<taxonomicName id="4C194D30F301FFFCE49A6C3C9CFED2E7" authority="Kataev" authorityName="Kataev" authorityYear="2021" box="[925,1221,628,652]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="449" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F301FFFCE49A6C3C9C54D2E0" box="[925,1135,628,652]" italics="true" pageId="1" pageNumber="449">Coleolissus inessae</emphasis>
Kataev
</taxonomicName>
,
<emphasis id="B96DEAA1F301FFFCE3D76C3C9D3ED2E0" bold="true" box="[1232,1285,628,652]" pageId="1" pageNumber="449">
<taxonomicNameLabel id="A25E57DAF301FFFCE3D76C3C9D3ED2E0" box="[1232,1285,628,652]" pageId="1" pageNumber="449" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3036538F301FFFCE3186CD99CB8D2C5" box="[1055,1155,657,681]" pageId="1" pageNumber="449" type="description">
<paragraph id="8BA636B3F301FFFCE3186CD99CB8D2C5" blockId="1.[812,1399,628,2068]" box="[1055,1155,657,681]" pageId="1" pageNumber="449">
<figureCitation id="13222A36F301FFFCE3186CD99C44D2C5" box="[1055,1151,657,681]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Figs 1–6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F301FFFCE4556CFE9BD8D295" pageId="1" pageNumber="449" type="materials_examined">
<paragraph id="8BA636B3F301FFFCE4556CFE9BD8D295" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
<materialsCitation id="3B713CEEF301FFFCE4556CFE9BD8D295" collectingDate="2013-11-02" collectingDateMax="2013-11-05" collectingDateMin="2013-11-02" collectionCode="ZIN" collectorName="S. Saluk" country="India" elevation="530" latitude="14.220667" location="Falls" longLatPrecision="1" longitude="74.80785" municipality="Gats Mts" pageId="1" pageNumber="449" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka" typeStatus="holotype">
MATERIAL.
<typeStatus id="54A28811F301FFFCE4CD6CFE9C25D2A5" box="[970,1054,694,713]" pageId="1" pageNumber="449" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D1FFD3AF301FFFCE3236CF19C03D2A5" box="[1060,1080,697,713]" count="1" pageId="1" pageNumber="449" type="male">♂</specimenCount>
, “
<collectingCountry id="F30E7623F301FFFCE34E6CFE9CB0D2A5" box="[1097,1163,694,713]" name="India" pageId="1" pageNumber="449">INDIA</collectingCountry>
: W
<collectingRegion id="49DDF851F301FFFCE3AD6CFE9D3ED2A5" box="[1194,1285,694,713]" country="India" name="Karnataka" pageId="1" pageNumber="449">Karnataka</collectingRegion>
, W
<collectingMunicipality id="6BC2ACC9F301FFFCE2236CFE9D48D2A6" box="[1316,1395,694,714]" pageId="1" pageNumber="449">Gats Mts</collectingMunicipality>
, jog
<location id="8EC66068F301FFFCE4486C869B40D28D" LSID="urn:lsid:plazi:treatment:03B087A5F301FFFEE49A6C3C9B4DD87A:8EC66068F301FFFCE4486C869B40D28D" box="[847,891,718,737]" country="India" latitude="14.220667" longLatPrecision="1" longitude="74.80785" municipality="Gats Mts" name="Falls" pageId="1" pageNumber="449" stateProvince="Karnataka">Falls</location>
,
<quantity id="4CE19B56F301FFFCE4816C869BFAD28E" box="[902,961,718,738]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="449" unit="m" value="530.0">
<elevation id="0034D180F301FFFCE4816C869BFAD28E" box="[902,961,718,738]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="449" unit="m" value="530.0">530 m</elevation>
</quantity>
, at light,
<geoCoordinate id="EE2D5074F301FFFCE3106C869CBCD28D" box="[1047,1159,718,737]" degrees="14" direction="north" minutes="13.240" orientation="latitude" pageId="1" pageNumber="449" precision="1" value="14.220667">14°13.240´N</geoCoordinate>
<geoCoordinate id="EE2D5074F301FFFCE3886C869CC5D28D" box="[1167,1278,718,737]" degrees="74" direction="east" minutes="48.471" orientation="longitude" pageId="1" pageNumber="449" precision="1" value="74.80785">74°48.471´E</geoCoordinate>
,
<date id="FFA71073F301FFFCE20E6C869D4DD28D" box="[1289,1398,718,737]" pageId="1" pageNumber="449" value="2013-11-02" valueMax="2013-11-05" valueMin="2013-11-02">
<collectingDate id="EFE3E99BF301FFFCE20E6C869D4DD28D" box="[1289,1398,718,737]" pageId="1" pageNumber="449" value="2013-11-02" valueMax="2013-11-05" valueMin="2013-11-02">2–5.XI.2013</collectingDate>
</date>
<collectorName id="26EC5365F301FFFCE42B6CAE9B4ED295" box="[812,885,742,761]" pageId="1" pageNumber="449">S. Saluk</collectorName>
leg.” (
<collectionCode id="ED08AE76F301FFFCE4B66CAE9BE2D295" box="[945,985,742,761]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="449" type="Museum">ZIN</collectionCode>
).
</materialsCitation>
</paragraph>
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<subSubSection id="C3036538F301FFFEE4556CB69988D72D" lastPageId="3" lastPageNumber="451" pageId="1" pageNumber="449" type="description">
<paragraph id="8BA636B3F301FFFCE4556CB69D19D342" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
DESCRIPTION (male). Body elongate, medium-sized, length
<quantity id="4CE19B56F301FFFCE4686D519B86D342" box="[879,957,793,814]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="1" pageNumber="449" unit="mm" value="7.8">7.8 mm</quantity>
, width 3.0 mm. Habitus as in
<figureCitation id="13222A36F301FFFCE3E46D519D26D342" box="[1251,1309,793,815]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="8BA636B3F301FFFCE4556D7B9C58D3D8" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">Body pitch black, shiny and iridescent on dorsal and ventral sides, with labrum, base of mandibles and very narrow lateral margins of pronotum reddish brown; palpi, antennae and legs yellowish brown; antennomeres 3–11 and apices of tibiae very slightly infuscate.</paragraph>
<paragraph id="8BA636B3F301FFFCE4556DF19D37D735" blockId="1.[812,1399,628,2068]" pageId="1" pageNumber="449">
Head medium-sized (HWmax/PWmax 0.68, HWmin/ PWmax 0.53), impunctate. Eyes large, moderately convex (HWmax/HWmin 1.28), separated ventrally from buccal fissure by very short distance about as wide as antennomere 1 basally. Genae glabrous. Tempora short, oblique, gently sloping to neck. Labrum slightly convex, very shallowly concave anteriorly. Clypeus somewhat flat, slightly emarginated along anterior margin, with a pair of lateral setigerous pores and a pair of very short longitudinal wrinkles each adjacent medially to lateral pore. Fronto-clypeal suture fine, slightly impressed, almost straight. Frontal foveae like elongate short pit without prolongation on clypeus. Fronto-ocular furrows very fine, vaguely reaching supraocular furrows. Frons with angulate shallow depression and with a few very fine micropunctures at lateral margins of this depression. Supraorbital furrows very narrow, touching upper margin of eyes. Supraorbital setigerous pores small, situated slightly before level of posterior margin of eyes and removed from supraorbital furrows by diameter of pores.
<collectingRegion id="49DDF851F301FFFCE35E6BD19C90D5C2" box="[1113,1195,1433,1454]" country="Myanmar" name="Chin" pageId="1" pageNumber="449">Mentum</collectingRegion>
separated from submentum by complete transverse suture (
<figureCitation id="13222A36F301FFFCE3B16BFB9CC8D5A5" box="[1206,1267,1459,1481]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="1" pageNumber="449">Fig. 2</figureCitation>
), with a very short and wide median tooth; epilobes markedly widened apically; submentum with one pair of long lateral setae. Ligular sclerite almost parallel-sided, truncate at apex, with one pair of ventroapical setae. Paraglossae glabrous, moderately wide, markedly longer than ligular sclerite and separat- ed from it apically by narrow notches. Basal labial palpomere not carinate; penultimate labial palpomere about as long as ultimate one. Mandibles very moderately elongate, evenly curved apically, with wide bases; left mandible blunted at tip. Dorsal microsculpture very fine, obliterated, consisting of indistinct, more or less isodiametric meshes. Antennae slen- der, surpassing pronotal basal edge by two apical antennomeres, pubescent from apical half of antennomere 3, with antennomeres 4–8 about two times as long as wide and basal antennomere slightly longer than antennomere 3.
</paragraph>
<paragraph id="8BA636B3F301FFFFE45569179CB7D7BB" blockId="1.[812,1399,628,2068]" lastBlockId="2.[812,1399,1799,2061]" lastPageId="2" lastPageNumber="450" pageId="1" pageNumber="449">Pronotum transverse, comparatively long for genus (PWmax/PL 1.34), widest slightly before the middle, very moderately narrowed apically and basally (PWmax/PWmin-ap 1.44, PWmax/PWmin-bas 1.20), with one lateral seta inserted at the end of the anterior third. Sides rounded along entire length; lateral bead complete, throughout very narrow. Apical margin shallowly emarginated, almost straight medially, very narrowly bordered along entire length; border slightly obliterated at the middle. Apical angles rounded, slightly protruding anteriorly. Basal margin almost straight in middle portion, slightly rounded laterally, bordered along entire length, not ciliate, slightly longer than apical margin (PWmin-bas/PWmin-ap 1.20) and almost as long as base of elytra between humeral angles. Basal angles obtuse, rounded at apex. Disc convex. Lateral depressions beginning from lateral setae, very shallow, widened basally and disappearing before the apical quarter. Basal foveae elongate oval, shallow, isolated from lateral margins and from each other by convex areas. Median line distinct, slightly impressed, isolat- ed from apical margin and touching basal margin. Anterior transverse depression wide and shallow. Surface with dense distinct non-setigerous punctures basally and along sides, with sparser and finer punctation at apical margin including anterior transverse depression. Microsculpture highly obliterated, not forming distinct meshes.</paragraph>
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<paragraph id="8BA636B3F302FFFFE7E568CF9BA5D6DE" blockId="2.[189,1399,1671,1766]" pageId="2" pageNumber="450">
Figs 1–6.
<taxonomicName id="4C194D30F302FFFFE63168CF99E6D6F5" authorityName="Kataev" authorityYear="2021" box="[310,477,1671,1690]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F302FFFFE63168CF99E6D6F5" box="[310,477,1671,1690]" italics="true" pageId="2" pageNumber="450">Coleolissus inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F302FFFFE6E668CF9A37D6F6" bold="true" box="[481,524,1671,1690]" pageId="2" pageNumber="450">
<taxonomicNameLabel id="A25E57DAF302FFFFE6E668CF9A37D6F6" box="[481,524,1671,1690]" pageId="2" pageNumber="450" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6).
</paragraph>
<paragraph id="8BA636B3F302FFFFE7E568F39C73D689" blockId="2.[189,1399,1671,1766]" pageId="2" pageNumber="450">
Рис. 1–6.
<taxonomicName id="4C194D30F302FFFFE63868F299D2D6A1" authorityName="Kataev" authorityYear="2021" box="[319,489,1722,1741]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="450" phylum="Arthropoda" rank="species" species="inessae" status="sp. nov.">
<emphasis id="B96DEAA1F302FFFFE63868F299D2D6A1" box="[319,489,1722,1741]" italics="true" pageId="2" pageNumber="450">Coleolissus inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F302FFFFE6F568F29A27D6A1" bold="true" box="[498,540,1722,1741]" pageId="2" pageNumber="450">
<taxonomicNameLabel id="A25E57DAF302FFFFE6F568F29A27D6A1" box="[498,540,1722,1741]" pageId="2" pageNumber="450" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6).
</paragraph>
</caption>
<paragraph id="8BA636B3F302FFFEE45569949A20D3DA" blockId="2.[812,1399,1799,2061]" lastBlockId="3.[189,775,315,2071]" lastPageId="3" lastPageNumber="451" pageId="2" pageNumber="450">
Elytra convex, elongate, only slightly wider than pronotum (EL/EW 1.64, EL/PL 2.61, EW/PWmax 1.19), widest before the apical third; sides very slightly diverging in the apical two thirds, with weak sinuation before the middle, and roundly converging in the apical third; subapical sinuation distinct but shallow. Humeri rounded, with an indistinct denticle recognizable in caudal view. Sutural angle narrowly rounded at tip, not extended posteriorly. Basal edge slightly arched, forming a sharp, very obtuse angle with lateral margin. Striae comparatively wide, impunctate, impressed along entire length, reaching anteriorly basal elytral border, with granulate microsculpture on bottom. Intervals glabrous, almost flat on disc, gradually becoming convex and narrow at apex; intervals 8 and 9 throughout and other intervals just at apex irregularly punctate; punctation of intervals 8 and
<quantity id="4CE19B56F303FFFEE5E76C339B3CD2FD" box="[736,775,635,657]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="451" unit="in" value="9.0">9 in</quantity>
apical half very fine and sparse. Parascutellar (abbreviate) striole long, about 0.2 as long as elytron, with a large setigerous pore basally isolated from basal elytral border. Interval 3 with a series of seven very distinct discal setigerous pores adjoining to stria 2 along its entire length; intervals 5 and 7 without discal pores. Marginal umbilicate series without distinct gap at middle, consisting of 22–25 setigerous pores. Lateral groove narrow and flat throughout, without distinct elongate convexity along this groove apically. Microsculpture highly obliterated on intervals, without distinct meshes or lines, granulate in lateral groove.
</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56DF399F3D3BD" blockId="3.[189,775,315,2071]" box="[226,456,955,977]" pageId="3" pageNumber="451">Wings fully developed.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56D9E9A69D450" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Ventral side of thorax glabrous except for sparse very fine short setae on prosternum anteriorly, almost throughout finely punctate laterally. Metepisternum markedly longer than wide, strongly narrowed posteriorly.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56A099A10D58A" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Metacoxae each only with two obligatory setigerous pores, without additional pores and setae. Profemur with deep longitudinal excavation on inner side. Protibia on dorsal side without longitudinal sulcus, with two similar sized preapical spines on outer margin. Metafemur ventrally with two setae at posterior margin and without setae at anterior margin. Pro- and mesotarsi dorsally with a few very fine and short setae. Protarsomeres 1–4 and mesotarsomeres 2–4 (in male) markedly widened, short, with biseriate adhesive scales ventrally; mesotarsomere 1 not widened, elongate, only slightly shorter than mesotarsomeres 2 and 3 combined, with a pair of very small scales just at apex. Metatarsus slender, about as long as HWmax, with tarsomeres 2 and 3 weakly and tarsomeres 3 and 4 markedly widened distally; metatarsomere 1 slightly longer than metatarsomeres 2 and 3 combined. Tarsomere 5 with two pairs of ventro-lateral setae.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E56BA39AF3D63D" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">Abdominal sternites mostly smooth, only two basal ones with very fine non-setigerous punctures; last visible sternite (VII) (in male) very widely rounded at apex, with one pair of marginal setae; these setae not removed from margin.</paragraph>
<paragraph id="8BA636B3F303FFFEE7E5681E9988D72D" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">
Aedeagus (
<figureCitation id="13222A36F303FFFEE64A681E9999D600" box="[333,418,1622,1644]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Figs 3–6</figureCitation>
) with comparatively large parameres (
<figureCitation id="13222A36F303FFFEE7C468399923D6EA" box="[195,280,1649,1671]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Figs 3, 4</figureCitation>
) and basal bulb. Median lobe of aedeagus in lateral view (
<figureCitation id="13222A36F303FFFEE7FA68C39900D6CD" box="[253,315,1675,1697]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Fig. 5</figureCitation>
) arcuate on dorsal side, with ventral margin almost straight in middle portion and convex in the apical quarter; apex sharply hooged ventrally; in dorsal view medi- an lobe (
<figureCitation id="13222A36F303FFFEE6126893996AD69D" box="[277,337,1755,1777]" captionStart="Figs 1–6" captionStartId="2.[226,263,1671,1690]" captionTargetBox="[180,1389,314,1652]" captionTargetId="figure-163@2.[195,858,326,1652]" captionTargetPageId="2" captionText="Figs 1–6. Coleolissus inessae sp.n. 1 — general view; 2 — labium; 3 — left paramere; 4 — right paramere; 5 — median lobe of aedeagus, lateral view; 6 —same, dorsal view. Scale bars: A = 1.0 mm (1), B = 0.5 mm (2–6). Рис. 1–6. Coleolissus inessae sp.n. 1 — внешний вид; 2 — нижнЯЯ губа; 3 — леваЯ парамера; 4 — праваЯ парамера; 5–6 — срединнаЯ лопасть Эдеагуса (5 — сбоку; 6 — сверху). Масштаб: A = 1,0 мм (1); B = 0,5 мм (2–6)." figureDoi="http://doi.org/10.5281/zenodo.13178232" httpUri="https://zenodo.org/record/13178232/files/figure.png" pageId="3" pageNumber="451">Fig. 6</figureCitation>
) almost straight, with terminal lamella very wide and short, rounded apically. Apical orifice in dorsal position, wide apically. Internal sac with spiny formation medially, without spines.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE7E5690E9C58D107" pageId="3" pageNumber="451" type="etymology">
<paragraph id="8BA636B3F303FFFEE7E5690E9AEBD7C0" blockId="3.[189,775,315,2071]" pageId="3" pageNumber="451">
ETYMOLOGY. Named after Prof. Inessa Kh. Sharova (1931–2021), the famous entomologist and soil zoologist, the teacher of many carabidologists in the former Soviet Union and
<collectingCountry id="F30E7623F303FFFEE7E269DE991DD7C0" box="[229,294,1942,1964]" name="Russia" pageId="3" pageNumber="451">Russia</collectingCountry>
, to whose memory this volume is dedicated.
</paragraph>
<paragraph id="8BA636B3F303FFFEE7E569F99C58D107" blockId="3.[189,775,315,2071]" lastBlockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">COMPARISON. This new species is markedly distinguished from all other congeners by elongate body with comparatively long pronotum which is not depressed latero-basally. In addition, unlike most of other species, its mandibles are shorter and abdominal sternite VII (in male) is with only one pair of marginal setae.</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE4556F399CA9D1CD" pageId="3" pageNumber="451" type="distribution">
<paragraph id="8BA636B3F303FFFEE4556F399CA9D1CD" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">
DISTRIBUTION. Known from the Western Ghats near Jog Falls, Western
<collectingRegion id="49DDF851F303FFFEE4EE6FC39C74D1CD" box="[1001,1103,395,417]" country="India" name="Karnataka" pageId="3" pageNumber="451">Karnataka</collectingRegion>
,
<collectingCountry id="F30E7623F303FFFEE35E6FC39CB5D1CD" box="[1113,1166,395,417]" name="India" pageId="3" pageNumber="451">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036538F303FFFEE4556FEE9B4DD87A" pageId="3" pageNumber="451" type="discussion">
<paragraph id="8BA636B3F303FFFEE4556FEE9B4DD87A" blockId="3.[812,1399,315,2070]" pageId="3" pageNumber="451">
REMARKS. This new species is rather dissimilar to any other described species and at first I hesitated to place it in any described genus. Such characters as the distinct fronto-ocular furrows, glabrous paraglossae, several setae on penultimate labial palpomere, elytral interval 3 with row of setigerous pores, long metatarsomere 1 and pro- and mesotarsi of male with biseriate adhesive scales are clearly indicate that it is a member of the Selenophori group. In having moderately elongate mandibles and glabrous, almost subcylindrical body without latero-basal depressions on pronotum, it looks as somewhat intermediate form between
<taxonomicName id="4C194D30F303FFFEE42B6C839BA6D28C" authorityName="Bates" authorityYear="1892" box="[812,925,715,736]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE42B6C839BA6D28C" box="[812,925,715,736]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE4CB6C849C07D28D" authorityName="Chaudoir" authorityYear="1854" box="[972,1084,716,737]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4CB6C849C07D28D" box="[972,1084,716,737]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
. Although these taxa are always regarded as separate genera, the structural differences between them are not evident [
<bibRefCitation id="EF884B42F303FFFEE3406D499CEAD37A" author="Kataev B. M." box="[1095,1233,768,790]" pageId="3" pageNumber="451" pagination="43 - 55" refId="ref6963" refString="Kataev B. M. 2006. A taxonomic review of the genus Oesyperus (Coleoptera: Carabidae: Harpalini) // Zootaxa. Vol. 1286. P. 43 - 55." type="book chapter" year="2006">Kataev, 2006</bibRefCitation>
] and assignment of some species to one or another genus is difficult since the body form is rather variable in the both genera. For example, two apparently closely related species with elongate body were described by one author [
<bibRefCitation id="EF884B42F303FFFEE3A26D249D29D3ED" author="Ito N." box="[1189,1298,875,897]" pageId="3" pageNumber="451" pagination="39 - 43" refId="ref6259" refString="Ito N. 1997 b. Two new species of the genus Oxycentrus from Vietnam (Coleoptera, Carabidae, Harpalini) // Ent. Rev. Japan. Vol. 52. No. 1. P. 39 - 43." type="journal article" year="1997">Ito, 1997b</bibRefCitation>
,
<bibRefCitation id="EF884B42F303FFFEE2186D239D69D3EC" author="Ito N." box="[1311,1362,875,896]" pageId="3" pageNumber="451" pagination="165 - 168" refId="ref6302" refString="Ito N. 1999. A new species of the genus Coleolissus from Laos (Coleoptera: Carabidae: Harpalini) // Ent. Rev. Japan. Vol. 54. No. 2. P. 165 - 168." type="journal article" year="1999">1999</bibRefCitation>
] as
<taxonomicName id="4C194D30F303FFFEE42B6DCE9CD7D3F7" authority="Ito, 1997" authorityName="Ito" authorityYear="1997" box="[812,1260,902,923]" class="Insecta" family="Carabidae" genus="Oxycentrus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="subarcuaticollis">
<emphasis id="B96DEAA1F303FFFEE42B6DCE9BA7D3F7" box="[812,924,902,923]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
(s. str.)
<emphasis id="B96DEAA1F303FFFEE4E96DCE9CB6D3F7" box="[1006,1165,902,923]" italics="true" pageId="3" pageNumber="451">subarcuaticollis</emphasis>
Ito, 1997
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE21B6DCE9C95D3DA" authority="Ito, 1999" authorityName="Ito" authorityYear="1999" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="iridipennis" subGenus="Tenuistilus">
<emphasis id="B96DEAA1F303FFFEE21B6DCE9B70D3DA" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
(
<emphasis id="B96DEAA1F303FFFEE4596DE99BF1D3DA" box="[862,970,929,950]" italics="true" pageId="3" pageNumber="451">Tenuistilus</emphasis>
)
<emphasis id="B96DEAA1F303FFFEE4DA6DE99C73D3DA" box="[989,1096,929,950]" italics="true" pageId="3" pageNumber="451">iridipennis</emphasis>
<bibRefCitation id="EF884B42F303FFFEE3566DE99C95D3DA" author="Ito N." box="[1105,1198,929,950]" pageId="3" pageNumber="451" pagination="165 - 168" refId="ref6302" refString="Ito N. 1999. A new species of the genus Coleolissus from Laos (Coleoptera: Carabidae: Harpalini) // Ent. Rev. Japan. Vol. 54. No. 2. P. 165 - 168." type="journal article" year="1999">Ito, 1999</bibRefCitation>
</taxonomicName>
without comparing them, and their generic affiliation remains unclear. They both markedly differ from
<taxonomicName id="4C194D30F303FFFEE3306D9E9CA5D380" authorityName="Kataev" authorityYear="2021" box="[1079,1182,982,1004]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3306D9E9CA5D380" box="[1079,1182,982,1004]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE3A26D9E9CEDD387" bold="true" box="[1189,1238,982,1003]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE3A26D9E9CEDD387" box="[1189,1238,982,1003]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
at least in body size larger (about 9.5–12.0 mm) and in having pronotum markedly depressed latero-basally, with more rounded sides and basal angles, and abdominal sternite VII of male with two pairs of marginal setae. The new species is included in the genus
<taxonomicName id="4C194D30F303FFFEE4966A139C39D41C" authorityName="Bates" authorityYear="1892" box="[913,1026,1115,1136]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4966A139C39D41C" box="[913,1026,1115,1136]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
on the basis of the following characters: frontal foveae without prolongation on clypeus, parascutellar striole long, interval 3 with a row of seven setigerous pores, and protibia with two preapical spines on outer margin. Each of these characters occurs in
<taxonomicName id="4C194D30F303FFFEE3D86A8E9D74D4B7" authorityName="Chaudoir" authorityYear="1854" box="[1247,1359,1222,1243]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3D86A8E9D74D4B7" box="[1247,1359,1222,1243]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
but not in such combination. As opposed to
<taxonomicName id="4C194D30F303FFFEE3C06AA99D03D49A" authorityName="Bates" authorityYear="1892" box="[1223,1336,1249,1270]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3C06AA99D03D49A" box="[1223,1336,1249,1270]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
, most species of
<taxonomicName id="4C194D30F303FFFEE4936AB49C3FD57D" authorityName="Chaudoir" authorityYear="1854" box="[916,1028,1276,1297]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4936AB49C3FD57D" box="[916,1028,1276,1297]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
have frontal foveae with more or less distinct prolongation on clypeus, parascutellar striole short or absent, interval 3 with less number of setigerous pores (in some species these pores absent), and protibia with more than two preapical spines on outer margin. The very moderately elongate mandibles of
<taxonomicName id="4C194D30F303FFFEE3476BC99C9DD5FA" authorityName="Kataev" authorityYear="2021" box="[1088,1190,1409,1430]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3476BC99C9DD5FA" box="[1088,1190,1409,1430]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE3AC6BC99CE6D5FA" bold="true" box="[1195,1245,1409,1430]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE3AC6BC99CE6D5FA" box="[1195,1245,1409,1430]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, shorter than in most
<taxonomicName id="4C194D30F303FFFEE4606BD39BE3D5DC" authorityName="Bates" authorityYear="1892" box="[871,984,1435,1456]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE4606BD39BE3D5DC" box="[871,984,1435,1456]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE3136BD49CBFD5DD" authorityName="Chaudoir" authorityYear="1854" box="[1044,1156,1436,1457]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3136BD49CBFD5DD" box="[1044,1156,1436,1457]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
, and its comparatively long pronotum without latero-basal depressions, which are present probably in all congeners, suggest that this species is rather isolated within
<taxonomicName id="4C194D30F303FFFEE31B6BA39CB6D66C" authorityName="Bates" authorityYear="1892" box="[1052,1165,1515,1536]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE31B6BA39CB6D66C" box="[1052,1165,1515,1536]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
. The similarity to
<taxonomicName id="4C194D30F303FFFEE2406BA49B4FD670" authorityName="Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE2406BA49B4FD670" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
may be convergent (most species of
<taxonomicName id="4C194D30F303FFFEE3DB684E9D76D677" authorityName="Bates" authorityYear="1892" box="[1244,1357,1542,1563]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3DB684E9D76D677" box="[1244,1357,1542,1563]" italics="true" pageId="3" pageNumber="451">Coleolissus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE42B68699BA7D65A" authorityName="Chaudoir" authorityYear="1854" box="[812,924,1569,1590]" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE42B68699BA7D65A" box="[812,924,1569,1590]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
have the plesiomorphic state of the abdominal sternite VII with two pairs of marginal setae as opposed to the apomorphic state in the new species with only one, inner, pair of setae), but it is not improbably that
<taxonomicName id="4C194D30F303FFFEE21668399D4CD6EA" authorityName="Kataev" authorityYear="2021" box="[1297,1399,1649,1670]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE21668399D4CD6EA" box="[1297,1399,1649,1670]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE42B68C39B65D6CC" bold="true" box="[812,862,1675,1696]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE42B68C39B65D6CC" box="[812,862,1675,1696]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is phylogenetically close to the branching node of two phyletic lineages sensu
<bibRefCitation id="EF884B42F303FFFEE31D68EE9CFBD6D0" author="Noonan G. R." box="[1050,1216,1702,1724]" pageId="3" pageNumber="451" pagination="1 - 33" refId="ref7311" refString="Noonan G. R. 1985 b. Reconstructed phylogeny and zoogeography of the genera and subgenera of the Selenophori group (Insecta: Coleoptera: Carabidae: Harpalini: Harpalina) // Milwaukee Public Museum, Contributions in Biology and Geology. No. 65. P. 1 - 33." type="book chapter" year="1985">Noonan [1985b]</bibRefCitation>
: the less modified lineage
<taxonomicName id="4C194D30F303FFFEE47C68899BCFD6BA" authorityName="MacLeay" authorityYear="1825" box="[891,1012,1729,1750]" class="Insecta" family="Carabidae" genus="Hyphaereon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE47C68899BCFD6BA" box="[891,1012,1729,1750]" italics="true" pageId="3" pageNumber="451">Hyphaereon</emphasis>
</taxonomicName>
and the more specialized lineage
<taxonomicName id="4C194D30F303FFFEE24068899B4FD69D" authorityName="Chaudoir" authorityYear="1854" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE24068899B4FD69D" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
</taxonomicName>
. The relationships and status of
<taxonomicName id="4C194D30F303FFFEE3D768949D01D69D" authorityName="Kataev" authorityYear="2021" box="[1232,1338,1756,1777]" class="Insecta" family="Carabidae" genus="Coleolissus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="species" species="inessae" status="sp.n.">
<emphasis id="B96DEAA1F303FFFEE3D768949D01D69D" box="[1232,1338,1756,1777]" italics="true" pageId="3" pageNumber="451">C. inessae</emphasis>
</taxonomicName>
<emphasis id="B96DEAA1F303FFFEE24268939D4DD69C" bold="true" box="[1349,1398,1755,1776]" pageId="3" pageNumber="451">
<taxonomicNameLabel id="A25E57DAF303FFFEE24268939D4DD69C" box="[1349,1398,1755,1776]" pageId="3" pageNumber="451" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
need further study. Interestingly, the new species is somewhat similar in appearance to some species of
<taxonomicName id="4C194D30F303FFFEE22469599B8FD72D" authority="Murray, 1859" authorityName="Murray" authorityYear="1859" class="Insecta" family="Carabidae" genus="Siopelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE22469599D4DD74A" box="[1315,1398,1809,1830]" italics="true" pageId="3" pageNumber="451">Siopelus</emphasis>
Murray, 1859
</taxonomicName>
having smooth elytra, but well distinguished from them by having longer mandibles and glabrous genae and outer elytral intervals. The predominantly Ethiopian genus
<taxonomicName id="4C194D30F303FFFEE46969339BFAD7FC" authorityName="Murray" authorityYear="1859" box="[878,961,1915,1936]" class="Insecta" family="Carabidae" genus="Siopelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE46969339BFAD7FC" box="[878,961,1915,1936]" italics="true" pageId="3" pageNumber="451">Siopelus</emphasis>
</taxonomicName>
seems to occupy ancestral position in relation to the phyletic lineages
<taxonomicName id="4C194D30F303FFFEE35169DE9CF4D7C7" authorityName="MacLeay" authorityYear="1825" box="[1110,1231,1942,1963]" class="Insecta" family="Carabidae" genus="Hyphaereon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE35169DE9CF4D7C7" box="[1110,1231,1942,1963]" italics="true" pageId="3" pageNumber="451">Hyphaereon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C194D30F303FFFEE20169DE9C34D7AA" authority="sensu Noonan [1985 b]" authorityName="sensu Noonan" authorityYear="1985" class="Insecta" family="Carabidae" genus="Oxycentrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE20169DE9D4DD7C7" box="[1286,1398,1942,1963]" italics="true" pageId="3" pageNumber="451">Oxycentrus</emphasis>
sensu
<bibRefCitation id="EF884B42F303FFFEE46D69F99C34D7AA" author="Noonan G. R." box="[874,1039,1968,1990]" pageId="3" pageNumber="451" pagination="1 - 33" refId="ref7311" refString="Noonan G. R. 1985 b. Reconstructed phylogeny and zoogeography of the genera and subgenera of the Selenophori group (Insecta: Coleoptera: Carabidae: Harpalini: Harpalina) // Milwaukee Public Museum, Contributions in Biology and Geology. No. 65. P. 1 - 33." type="book chapter" year="1985">Noonan [1985b]</bibRefCitation>
</taxonomicName>
, and, in my opinion, does not differ morphologically from the American genus
<taxonomicName id="4C194D30F303FFFEE3F769839B8AD797" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" class="Insecta" family="Carabidae" genus="Selenophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="451" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEAA1F303FFFEE3F769839D4DD78C" box="[1264,1398,1995,2016]" italics="true" pageId="3" pageNumber="451">Selenophorus</emphasis>
Dejean, 1829
</taxonomicName>
which is ancestral group for many American genera.
</paragraph>
</subSubSection>
</treatment>
</document>
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