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<document id="8CD82A146E7E17993FB577102BBC89B5" ID-DOI="10.5852/ejt.2024.926.2469" ID-ISSN="2118-9773" ID-Zenodo-Dep="10847806" ID-ZooBank="A00D7669-2151-46AE-A066-9AF4D0F2BEEE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1711011351298" checkinUser="plazi" docAuthor="Jochum, Adrienne, Michalik, Peter, Inäbnit, Thomas, Kneubühler, Jeannette, Slapnik, Rajko, Vrabec, Marko, Schilthuizen, Menno &amp; Ruthensteiner, Bernhard" docDate="2024" docId="03B0879DFF95FFF2FE19D249FE15FC23" docLanguage="en" docName="EJT.2024.926.1-62.pdf" docOrigin="European Journal of Taxonomy 926" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zospeum simplex Inabnit, Jochum &amp; Neubert 2021" docType="treatment" docVersion="5" lastPageNumber="51" masterDocId="FF89FFE5FFBAFFC0FFFFD700FFD4FFD4" masterDocTitle="3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)" masterLastPageNumber="62" masterPageNumber="1" pageNumber="48" updateTime="1711118001397" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FA8E2E1FB03D75BD56A842BAEB9E7796">3 D X-ray microscopy (Micro-CT) and SEM reveal Zospeum troglobalcanicum Absolon, 1916 and allied species from the Western Balkans (Ellobioidea: Carychiidae)</mods:title>
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<mods:namePart id="7B8CEBF9AC9B0DC4588E33BF75E984EA">Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier id="129EF5789C7BC7268C5FDC2368A71247" type="email">nne.jochum@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="DE217EF4C5203BB39128921118EDCE6A">Michalik, Peter</mods:namePart>
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<mods:namePart id="1BDC2B094C58D8B29FDFB98C241F9B35">Inäbnit, Thomas</mods:namePart>
<mods:nameIdentifier id="C1DBE0C3B68D0ABEFEE967904C1CE148" type="LSID">95F0148B-E147-4EB9-903A-6EEA7F55EB47</mods:nameIdentifier>
<mods:affiliation id="91BC353FD46CE76800245AF2D33BA648">Institute for Biochemistry &amp; Biology, University of Potsdam, Karl-Liebknecht-Strasse 24 - 25, House 26, D- 14476 Potsdam, Germany.</mods:affiliation>
<mods:nameIdentifier id="E1A3BEEAFA75EBCFB67784640D137528" type="email">inaebnit.thomas@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="9D74490121C27B02B22E9CC6A53C28AB" type="personal">
<mods:role id="980F5A9B48F3551B7303988635959024">
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<mods:namePart id="CD0E9114E792D3321C4928F9EE088978">Kneubühler, Jeannette</mods:namePart>
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<mods:affiliation id="7F68E6AE5C3D31C924625CC8DC8FB672">Naturhistorisches Museum Bern, Bernastrasse 15, CH- 3005 Bern, Switzerland. &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, CH- 3012 Bern, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="984737D71C5D448EAA59DA3C6EF4797C" type="email">jeannette.kneubuehler@nmbe.ch</mods:nameIdentifier>
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<mods:namePart id="2084CDD27BB1E21D44F8C7C319CAA3E2">Slapnik, Rajko</mods:namePart>
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<mods:affiliation id="DA20A51F19D9D62A57FF898F7DEDDDE0">ZOSPEUM, Molluscs, Cave &amp; Karst Biological Consulting, Drnovškova pot 2, Mekinje, SI 1240 Kamnik, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="BCAC7E5C98075132BC8843A6936DD72C" type="email">rajko.slapnik@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5876FC8B4AB05D7C0AC0CC2516FC53A5">Vrabec, Marko</mods:namePart>
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<mods:affiliation id="1EFBD594B1914C8FAA3A18F3924B2C98">Department of Geology, Faculty of Natural Sciences and Engineering, Aškerčeva 12, University of Ljubljana, 1000 Ljubljana, Slovenia.</mods:affiliation>
<mods:nameIdentifier id="ECE544D7D8D30A1DA647C6BAED7ACFA6" type="email">marko.vrabec@geo.ntf.uni-lj.si</mods:nameIdentifier>
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<mods:namePart id="9DABA972C77B1FE8B1CD58050654C1F7">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="AE5B47B452F22A2A181D7AAF100591D3">Naturalis Biodiversity Center and Leiden University, Vondellaan 55, NL- 2332 AA Leiden, The Netherlands.</mods:affiliation>
<mods:nameIdentifier id="2EAE359C8E5B9C01A8EF2057CE565CB1" type="email">menno.schilthuizen@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="6EF04FA6C1C1E76A7985F76C51169E37">Ruthensteiner, Bernhard</mods:namePart>
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<mods:affiliation id="0CB7C958FC63B54B5C0BBDC32A0C9259">SNSB - Zoologische Staatssammlung München (ZSM), 81247 München, Germany.</mods:affiliation>
<mods:nameIdentifier id="6AEBAED225348C983722ECC59525F830" type="email">ruthensteiner@snsb.de</mods:nameIdentifier>
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<mods:typeOfResource id="9FA9D9F7453A73702433EE564A12126E">text</mods:typeOfResource>
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<mods:title id="089EBE02E69E4141145C1683EA91550C">European Journal of Taxonomy</mods:title>
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<mods:part id="5E17BDD80D0444E94F7D7676F4C46127">
<mods:date id="7D5526F52201CA76A94F42860DDEE55A">2024</mods:date>
<mods:detail id="C8745B1485AB3B69D2855FD6A639CAEC" type="pubDate">
<mods:number id="E657D1FF1C2F0F7D580C46C5618F6028">2024-03-20</mods:number>
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<mods:detail id="1444197C9C9EC29276D5A54FB3A083E8" type="volume">
<mods:number id="19279B41A984D9490B9CA1F99BCD5F11">926</mods:number>
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<mods:start id="7CA9BC63AA1982234243169B3FD90CDF">1</mods:start>
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<mods:url id="BFB61DE380E512BDF27A05485A56659D">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2469/10973</mods:url>
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<mods:classification id="AA6505294D8B5FC678E672CB6C1651ED">journal article</mods:classification>
<mods:identifier id="03B701AE97153F7DD28D9201F0E52A6D" type="DOI">10.5852/ejt.2024.926.2469</mods:identifier>
<mods:identifier id="AC103E079CB68DC11A43C5D9939E560B" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="6A41BDFDA524E651A1DF1CA3DFCF5611" type="Zenodo-Dep">10847806</mods:identifier>
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<treatment id="03B0879DFF95FFF2FE19D249FE15FC23" LSID="urn:lsid:plazi:treatment:03B0879DFF95FFF2FE19D249FE15FC23" httpUri="http://treatment.plazi.org/id/03B0879DFF95FFF2FE19D249FE15FC23" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48">
<subSubSection id="C3036500FF95FFEFFE19D249FB99FAB7" box="[486,1101,1352,1379]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BA6368BFF95FFEFFE19D249FB99FAB7" blockId="47.[486,1101,1352,1414]" box="[486,1101,1352,1379]" pageId="47" pageNumber="48">
<heading id="D0EE81E7FF95FFEFFE19D249FB99FAB7" box="[486,1101,1352,1379]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="48" reason="2">
<taxonomicName id="4C194D08FF95FFEFFE19D249FB99FAB7" ID-CoL="8SN4W" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[486,1101,1352,1379]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF95FFEFFE19D249FD68FAB7" bold="true" box="[486,700,1352,1379]" italics="true" pageId="47" pageNumber="48">Zospeum simplex</emphasis>
<bibRefCitation id="EF884B7AFF95FFEFFD3CD249FB99FAB7" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[707,1101,1353,1379]" pageId="47" pageNumber="48" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3036500FF95FFEFFD27D26CFC88FA52" box="[728,860,1388,1414]" pageId="47" pageNumber="48" type="description">
<paragraph id="8BA6368BFF95FFEFFD27D26CFC88FA52" blockId="47.[486,1101,1352,1414]" box="[728,860,1388,1414]" pageId="47" pageNumber="48">
<figureCitation id="13222A0EFF95FFEFFD27D26CFC88FA52" box="[728,860,1388,1414]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="48.[189,232,1857,1883]" captionStartId-1="49.[189,232,1856,1882]" captionTargetBox-0="[186,1395,563,1819]" captionTargetBox-1="[189,1398,1262,1816]" captionTargetId-0="figure-152@48.[186,1395,563,1819]" captionTargetId-1="figure-452@49.[189,1398,1262,1816]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionText-0="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." captionText-1="Fig. 19. Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (SMF 349425) Scanning Electron Microscopy images. A. Protoconch and upper teleoconch showing microstructure of superficial pitting. B. Close up view of pitting microstructure. C. Last whorl with axial ribbing extending beyond peristome lip. D. Close up view of second whorl showing rows of interrupted dashes of radial pitting. E. Growth lines and radial banding on teleoconch. F. Close up view of growth lines and shell microstructure." figureDoi-0="http://doi.org/10.5281/zenodo.10847858" figureDoi-1="http://doi.org/10.5281/zenodo.10847860" httpUri-0="https://zenodo.org/record/10847858/files/figure.png" httpUri-1="https://zenodo.org/record/10847860/files/figure.png" pageId="47" pageNumber="48">Figs 18–19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF95FFEFFF42D2ADFCDEF9CE" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BA6368BFF95FFEFFF42D2ADFE76FA13" blockId="47.[189,418,1453,1479]" box="[189,418,1453,1479]" pageId="47" pageNumber="48">
<heading id="D0EE81E7FF95FFEFFF42D2ADFE76FA13" bold="true" box="[189,418,1453,1479]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
<emphasis id="B96DEA99FF95FFEFFF42D2ADFE76FA13" bold="true" box="[189,418,1453,1479]" pageId="47" pageNumber="48">Original diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D2DCFCDEF9CE" blockId="47.[189,1398,1500,1562]" pageId="47" pageNumber="48">
Shell usually ca
<quantity id="4CE19B6EFF95FFEFFE66D2DCFE28FA23" box="[409,508,1500,1527]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="47" pageNumber="48" unit="mm" value="1.3">1.3 mm</quantity>
in height, transparent, conical, peristome thickened, roundish, with a differentiated parietal shield, lamellae not present.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF95FFEFFF42D141FDC2F87D" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BA6368BFF95FFEFFF42D141FEB8F98E" blockId="47.[189,364,1600,1627]" box="[189,364,1600,1627]" pageId="47" pageNumber="48">
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<emphasis id="B96DEA99FF95FFEFFF42D141FEB8F98E" bold="true" box="[189,364,1600,1627]" pageId="47" pageNumber="48">Type material</emphasis>
</heading>
</paragraph>
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<emphasis id="B96DEA99FF95FFEFFF15D16FFE88F95D" bold="true" box="[234,348,1647,1673]" pageId="47" pageNumber="48">
<typeStatus id="54A28829FF95FFEFFF15D16FFE88F95D" box="[234,348,1647,1673]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D192FEDDF927" blockId="47.[189,1398,1647,1779]" pageId="47" pageNumber="48">
<materialsCitation id="3B713CD6FF95FFEFFF42D192FEDDF927" collectingDate="2019-08-25" collectionCode="NMBE, RSC" collectorName="R. Slapnik &amp; J. Valentincic" country="Bosnia and Herzegovina" county="Gornji Brisnik" latitude="43.6347" location="Jama Dobravljevac" longLatPrecision="6" longitude="17.2328" municipality="Tomislavgrad" pageId="47" pageNumber="48" specimenCode="NMBE 568060" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F30E761BFF95FFEFFF42D192FD9CF978" box="[189,584,1682,1709]" name="Bosnia and Herzegovina" pageId="47" pageNumber="48">BOSNIA AND HERZEGOVINA</collectingCountry>
• municipality of
<collectingMunicipality id="6BC2ACF1FF95FFEFFCDFD192FC10F978" box="[800,964,1682,1708]" pageId="47" pageNumber="48">Tomislavgrad</collectingMunicipality>
,
<collectingCounty id="62C74E07FF95FFEFFC2FD192FBAAF978" box="[976,1150,1682,1708]" pageId="47" pageNumber="48">Gornji Brišnik</collectingCounty>
,
<location id="8EC66050FF95FFEFFB73D193FAA6F978" LSID="urn:lsid:plazi:treatment:03B0879DFF95FFF2FE19D249FE15FC23:8EC66050FF95FFEFFB73D193FAA6F978" box="[1164,1394,1682,1709]" country="Bosnia and Herzegovina" county="Gornji Brisnik" latitude="43.6347" longLatPrecision="6" longitude="17.2328" municipality="Tomislavgrad" name="Jama Dobravljevac" pageId="47" pageNumber="48">Jama Dobravljevac</location>
;
<geoCoordinate id="EE2D504CFF95FFEFFF41D1B5FE9EF904" box="[190,330,1717,1744]" degrees="43.6347" direction="north" orientation="latitude" pageId="47" pageNumber="48" precision="5" value="43.6347">43.6347° N</geoCoordinate>
,
<geoCoordinate id="EE2D504CFF95FFEFFEA6D1B6FE37F904" box="[345,483,1718,1744]" degrees="17.2328" direction="east" orientation="longitude" pageId="47" pageNumber="48" precision="5" value="17.2328">17.2328° E</geoCoordinate>
;
<date id="FFA7104BFF95FFEFFE0DD1B5FD4EF904" box="[498,666,1717,1744]" pageId="47" pageNumber="48" value="2019-08-25">
<collectingDate id="EFE3E9A3FF95FFEFFE0DD1B5FD4EF904" box="[498,666,1717,1744]" pageId="47" pageNumber="48" value="2019-08-25">25 Aug. 2019</collectingDate>
</date>
;
<collectorName id="26EC535DFF95FFEFFD56D1B6FCFCF91B" box="[681,808,1717,1744]" pageId="47" pageNumber="48">R. Slapnik</collectorName>
and
<collectorName id="26EC535DFF95FFEFFC99D1B6FBD3F91B" box="[870,1031,1717,1744]" pageId="47" pageNumber="48">J. Valentinčič</collectorName>
leg.;
<specimenCode id="DBBF9EF0FF95FFEFFBB1D1B6FAD1F904" box="[1102,1285,1717,1744]" collectionCode="NMBE" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="47" pageNumber="48">NMBE 568060</specimenCode>
(ex RSC 3760).
</materialsCitation>
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D01AFCFEF8E0" blockId="47.[189,810,1818,1844]" box="[189,810,1818,1844]" pageId="47" pageNumber="48">
<emphasis id="B96DEA99FF95FFEFFF42D01AFE2DF8E0" bold="true" box="[189,505,1818,1844]" pageId="47" pageNumber="48">Other material examined</emphasis>
(via Micro-CT and SEM)
</paragraph>
<paragraph id="8BA6368BFF95FFEFFF42D049FDC2F87D" blockId="47.[189,1372,1865,1962]" pageId="47" pageNumber="48">
<materialsCitation id="3B713CD6FF95FFEFFF42D049FDB9F852" collectingDate="2019-08-25" collectionCode="RSC" collectorName="R. Slapnik &amp; J. Valentincic" country="Bosnia and Herzegovina" county="Jama Dobravljevac" latitude="43.6347" location="BOSNIA AND HERZEGOVINA" longLatPrecision="6" longitude="17.2328" municipality="Gornji Brisnik" pageId="47" pageNumber="48" specimenCode="NHMW-MO-113642" specimenCount="2">
<collectingCountry id="F30E761BFF95FFEFFF42D049FD92F8B7" box="[189,582,1865,1891]" name="Bosnia and Herzegovina" pageId="47" pageNumber="48">BOSNIA AND HERZEGOVINA</collectingCountry>
•
<specimenCount id="9D1FFD02FF95FFEFFDA0D049FD61F8B7" box="[607,693,1865,1891]" pageId="47" pageNumber="48">2 specs</specimenCount>
(
<figureCitation id="13222A0EFF95FFEFFD3BD049FCCEF8B7" box="[708,794,1865,1891]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="47" pageNumber="48">Fig. 18</figureCitation>
, MicroCT); same collection data as for holotype;
<specimenCode id="DBBF9EF0FF95FFEFFF42D06CFE61F852" box="[189,437,1900,1926]" pageId="47" pageNumber="48">NHMW-MO-113642</specimenCode>
(ex RSC 3760)
</materialsCitation>
•
<materialsCitation id="3B713CD6FF95FFEFFD7AD06CFDC6F87D" collectingDate="2019-08-25" collectionCode="SMF, RSC" collectorName="R. Slapnik &amp; J. Valentincic" country="Bosnia and Herzegovina" county="Jama Dobravljevac" latitude="43.6347" location="Tomislavgrad" longLatPrecision="6" longitude="17.2328" municipality="Gornji Brisnik" pageId="47" pageNumber="48" specimenCode="SMF 349425" specimenCount="2">
<specimenCount id="9D1FFD02FF95FFEFFD7AD06CFD0FF852" box="[645,731,1900,1926]" pageId="47" pageNumber="48">2 specs</specimenCount>
(
<figureCitation id="13222A0EFF95FFEFFD14D06CFC95F852" box="[747,833,1900,1926]" captionStart="Fig" captionStartId="49.[189,232,1856,1882]" captionTargetBox="[189,1398,1262,1816]" captionTargetId="figure-452@49.[189,1398,1262,1816]" captionTargetPageId="49" captionText="Fig. 19. Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (SMF 349425) Scanning Electron Microscopy images. A. Protoconch and upper teleoconch showing microstructure of superficial pitting. B. Close up view of pitting microstructure. C. Last whorl with axial ribbing extending beyond peristome lip. D. Close up view of second whorl showing rows of interrupted dashes of radial pitting. E. Growth lines and radial banding on teleoconch. F. Close up view of growth lines and shell microstructure." figureDoi="http://doi.org/10.5281/zenodo.10847860" httpUri="https://zenodo.org/record/10847860/files/figure.png" pageId="47" pageNumber="48">Fig. 19</figureCitation>
, SEM); same collection data as for holotype;
<specimenCode id="DBBF9EF0FF95FFEFFF42D08FFE83F87E" box="[189,343,1935,1962]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="47" pageNumber="48">SMF 349425</specimenCode>
(ex RSC 3760)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF8AFFF0FF42D609FAD5FE6F" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BA6368BFF8AFFF0FF42D609FC5AFEF7" blockId="48.[189,910,264,291]" box="[189,910,264,291]" pageId="48" pageNumber="49">
<heading id="D0EE81E7FF8AFFF0FF42D609FC5AFEF7" bold="true" box="[189,910,264,291]" fontSize="11" level="3" pageId="48" pageNumber="49" reason="3">
<emphasis id="B96DEA99FF8AFFF0FF42D609FC5AFEF7" bold="true" box="[189,910,264,291]" pageId="48" pageNumber="49">Emended diagnosis considering Micro-CT and SEM Data</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF8AFFF0FF42D637FAD5FE6F" blockId="48.[188,1400,310,443]" pageId="48" pageNumber="49">
Shell ca
<quantity id="4CE19B6EFF8AFFF0FEE0D637FEACFE86" box="[287,376,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="48" pageNumber="49" unit="mm" value="1.4">1.4 mm</quantity>
and 5½–6 whorls; conical with narrow spire; peristome bears slight indentation at upper junction of parietal and palatal sides; low, hump-like callus on the inner columellar side of peristome; teleoconch with dense rows of microscopic, radially aligned, elongated pits; medium-sized columella with either two weak, loosely coiled low incomplete lamellae or a single incomplete lamella.
</paragraph>
</subSubSection>
<caption id="DF666603FF8AFFF0FF42D041FE9AF83C" ID-DOI="http://doi.org/10.5281/zenodo.10847858" ID-Zenodo-Dep="10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="48" pageNumber="49" startId="48.[189,232,1857,1883]" targetBox="[186,1395,563,1819]" targetPageId="48">
<paragraph id="8BA6368BFF8AFFF0FF42D041FE9AF83C" blockId="48.[189,1399,1857,2024]" pageId="48" pageNumber="49">
<emphasis id="B96DEA99FF8AFFF0FF42D041FEC9F88F" bold="true" box="[189,285,1857,1883]" pageId="48" pageNumber="49">Fig. 18.</emphasis>
3D visualizations of Micro-CT data of
<taxonomicName id="4C194D08FF8AFFF0FCFFD042FAA5F888" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[768,1393,1857,1884]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="48" pageNumber="49" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF8AFFF0FCFFD042FC1BF88F" box="[768,975,1857,1883]" italics="true" pageId="48" pageNumber="49">Zospeum simplex</emphasis>
<bibRefCitation id="EF884B7AFF8AFFF0FC25D041FAA5F888" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[986,1393,1857,1884]" pageId="48" pageNumber="49" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina.
<emphasis id="B96DEA99FF8AFFF0FF42D088FF26F876" bold="true" box="[189,242,1928,1954]" pageId="48" pageNumber="49">A–F</emphasis>
. Specimen 1 (spm1).
<emphasis id="B96DEA99FF8AFFF0FE07D088FDD9F876" bold="true" box="[504,525,1928,1954]" pageId="48" pageNumber="49">A</emphasis>
. Aperture view.
<emphasis id="B96DEA99FF8AFFF0FD2BD088FD3CF876" bold="true" box="[724,744,1928,1954]" pageId="48" pageNumber="49">B</emphasis>
. Aperture facing right view.
<emphasis id="B96DEA99FF8AFFF0FBBDD088FB82F876" bold="true" box="[1090,1110,1928,1954]" pageId="48" pageNumber="49">C</emphasis>
. Apical view.
<emphasis id="B96DEA99FF8AFFF0FAFCD088FACCF876" bold="true" box="[1283,1304,1928,1954]" pageId="48" pageNumber="49">D</emphasis>
. Dorsal view.
<emphasis id="B96DEA99FF8AFFF0FEFAD0ABFECDF811" bold="true" box="[261,281,1963,1989]" pageId="48" pageNumber="49">E</emphasis>
. Aperture facing left view.
<emphasis id="B96DEA99FF8AFFF0FD97D0ABFDAEF811" bold="true" box="[616,634,1963,1989]" pageId="48" pageNumber="49">F</emphasis>
. Ventral view.
<emphasis id="B96DEA99FF8AFFF0FCCDD0ABFCBFF811" bold="true" box="[818,875,1963,1989]" pageId="48" pageNumber="49">G–L</emphasis>
. Specimen 2 (spm2). Same perspectives as Specimen 1.
</paragraph>
</caption>
<subSubSection id="C3036500FF8BFFF1FF42D609FA8CFC77" pageId="49" pageNumber="50" type="description">
<paragraph id="8BA6368BFF8BFFF1FF42D609FC87FEF7" blockId="49.[189,851,264,291]" box="[189,851,264,291]" pageId="49" pageNumber="50">
<heading id="D0EE81E7FF8BFFF1FF42D609FC87FEF7" bold="true" box="[189,851,264,291]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B96DEA99FF8BFFF1FF42D609FC87FEF7" bold="true" box="[189,851,264,291]" pageId="49" pageNumber="50">Enhanced description with Micro-CT and SEM Data</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF8BFFF1FF42D638FBEAFEA0" blockId="49.[189,1399,311,373]" pageId="49" pageNumber="50">
<smallCapsWord id="8D40A057FF8BFFF1FF42D638FEBAFE84" baselines="331,331" box="[189,366,312,338]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="49" pageNumber="50">MEASUREMENTS</smallCapsWord>
(N = 2). sh:
<quantity id="4CE19B6EFF8BFFF1FDF6D637FD63FE86" box="[521,695,311,338]" metricMagnitude="-3" metricUnit="m" metricValue="1.4049999999999998" metricValueMax="1.41" metricValueMin="1.4" pageId="49" pageNumber="50" unit="mm" value="1.4049999999999998" valueMax="1.41" valueMin="1.4">1.40–1.41 mm</quantity>
; sw:
<quantity id="4CE19B6EFF8BFFF1FD0AD637FC77FE86" box="[757,931,311,338]" metricMagnitude="-4" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="49" pageNumber="50" unit="mm" value="0.945" valueMax="0.96" valueMin="0.93">0.93–0.96 mm</quantity>
; ah:
<quantity id="4CE19B6EFF8BFFF1FC23D637FB9CFE86" box="[988,1096,311,338]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="49" pageNumber="50" unit="mm" value="0.57">0.57 mm</quantity>
; aw:
<quantity id="4CE19B6EFF8BFFF1FB78D637FAE1FE86" box="[1159,1333,311,338]" metricMagnitude="-4" metricUnit="m" metricValue="6.15" metricValueMax="6.2" metricValueMin="6.1" pageId="49" pageNumber="50" unit="mm" value="0.615" valueMax="0.62" valueMin="0.61">0.61–0.62 mm</quantity>
; hlw:
<quantity id="4CE19B6EFF8BFFF1FF41D65AFEBFFEA1" box="[190,363,346,373]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.4" metricValueMin="8.299999999999999" pageId="49" pageNumber="50" unit="mm" value="0.835" valueMax="0.84" valueMin="0.83">0.83–0.84 mm</quantity>
; SA: 56.46–59.45 deg., NHMW-MO-113642 (ex
<collectionCode id="ED08AE4EFF8BFFF1FC4CD65AFC3EFEA0" box="[947,1002,346,372]" pageId="49" pageNumber="50">RSC</collectionCode>
3760).
</paragraph>
<paragraph id="8BA6368BFF8BFFF1FF42D69CFA8CFC77" blockId="49.[187,1399,410,931]" pageId="49" pageNumber="50">
Shell ca
<quantity id="4CE19B6EFF8BFFF1FEDCD69CFE55FE62" box="[291,385,412,438]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="49" pageNumber="50" unit="mm" value="1.4">1.4 mm</quantity>
, conical, with 5½–6 regularly coiled, convex whorls with height of last whorl slightly greater than half of shell height; spire angle narrow; protoconch bulbous with microstructure of dense pitting and some irregular raised bands extending to the suture; the second and third whorls bear rows of interrupted dashes of radial pitting; teleoconch sculpture includes blunt irregular growth lines and weak radial banding on all whorls; density of blunt striations varies, showing a tendency towards costateness at the suture zones; high magnification of shell surface (5 μm) shows tiny pinpoint perforations; aperture reniform, wider than high; peristome slightly oblique on columellar side with a low, hump-like thickening on the inner columellar side (
<figureCitation id="13222A0EFF8BFFF1FDEFD592FDAFFD78" box="[528,635,658,684]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18A</figureCitation>
), palatal side narrowly reflected with low ribbing extending some distance beyond the lip; parietal shield well defined, long and angular, with a slight indentation at upper parietal and palatal junction; upper rim of peristome recedes ca
<superScript id="7C6C9BC3FF8BFFF1FC47D5D8FC14FD32" attach="right" box="[952,960,728,742]" fontSize="6" pageId="49" pageNumber="50">1</superScript>
/
<subScript id="179D34CEFF8BFFF1FC37D5E3FC05FD25" attach="left" box="[968,977,739,753]" fontSize="6" pageId="49" pageNumber="50">5</subScript>
the width of the penultimate whorl (aperture facing left) (
<figureCitation id="13222A0EFF8BFFF1FE3FD5FCFDFCFCC2" box="[448,552,764,790]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18E</figureCitation>
); columella centrally aligned and slender; a low lamella is present under penultimate whorl (aperture facing right) (
<figureCitation id="13222A0EFF8BFFF1FD56D41FFCC5FCED" box="[681,785,799,825]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18B</figureCitation>
) which then turns into either two incomplete loosely coiled low lamellae (dorsal view) (
<figureCitation id="13222A0EFF8BFFF1FDAAD442FD6BFC88" box="[597,703,834,860]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18D</figureCitation>
) or constitutes one well pronounced one (
<figureCitation id="13222A0EFF8BFFF1FB54D442FADAFC88" box="[1195,1294,834,860]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18J</figureCitation>
); ventral side with small, umbilical depression with no puckering; columellar side of peristome arches slightly over and oblique to umbilical depression, alignment of last 1/4 whorl somewhat compact (
<figureCitation id="13222A0EFF8BFFF1FB1CD488FA9FFC77" box="[1251,1355,904,931]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="49" pageNumber="50">Fig. 18F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3036500FF8BFFF2FF42D4CAFE15FC23" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BA6368BFF8BFFF1FF42D4CAFEFBFC30" blockId="49.[189,303,970,996]" box="[189,303,970,996]" pageId="49" pageNumber="50">
<heading id="D0EE81E7FF8BFFF1FF42D4CAFEFBFC30" bold="true" box="[189,303,970,996]" fontSize="11" level="3" pageId="49" pageNumber="50" reason="3">
<emphasis id="B96DEA99FF8BFFF1FF42D4CAFEFBFC30" bold="true" box="[189,303,970,996]" pageId="49" pageNumber="50">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA6368BFF8BFFF1FF42D4F8FAA2FB74" blockId="49.[189,1399,1015,1184]" pageId="49" pageNumber="50">
Differs from
<taxonomicName id="4C194D08FF8BFFF1FEA5D4F9FD99FBC6" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[346,589,1016,1042]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF8BFFF1FEA5D4F9FD99FBC6" box="[346,589,1016,1042]" italics="true" pageId="49" pageNumber="50">Z. troglobalcanicum</emphasis>
</taxonomicName>
in that it bears at least one additional or ½ whorl; the upper parietal and palatal junction has a slight indentation in contrast to a well-defined notch; protoconch is more bulbous and somewhat elevated; the shell spire angle is narrower than that of
<taxonomicName id="4C194D08FF8BFFF1FBA7D340FA9EFB8D" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[1112,1354,1087,1113]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF8BFFF1FBA7D340FA9EFB8D" box="[1112,1354,1087,1113]" italics="true" pageId="49" pageNumber="50">Z. troglobalcanicum</emphasis>
</taxonomicName>
but within the spectrum of the narrowest spire measurements of two shells of
<taxonomicName id="4C194D08FF8BFFF1FBBDD363FEB4FB4B" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="constrictum" status="sp. nov.">
<emphasis id="B96DEA99FF8BFFF1FBBDD363FB24FBA8" box="[1090,1264,1123,1148]" italics="true" pageId="49" pageNumber="50">Z. constrictum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF8BFFF1FE98D386FE15FB74" box="[359,449,1158,1184]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
(57.5–59.2 deg.); width of the shell is narrower than that of
<taxonomicName id="4C194D08FF8BFFF1FB82D386FABAFB4B" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[1149,1390,1157,1183]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF8BFFF1FB82D386FABAFB4B" box="[1149,1390,1157,1183]" italics="true" pageId="49" pageNumber="50">Z. troglobalcanicum</emphasis>
</taxonomicName>
;
</paragraph>
<caption id="DF666603FF8BFFF1FF42D040FAE8F833" ID-DOI="http://doi.org/10.5281/zenodo.10847860" ID-Zenodo-Dep="10847860" httpUri="https://zenodo.org/record/10847860/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1856,1882]" targetBox="[189,1398,1262,1816]" targetPageId="49">
<paragraph id="8BA6368BFF8BFFF1FF42D040FAE8F833" blockId="49.[188,1399,1856,2024]" pageId="49" pageNumber="50">
<emphasis id="B96DEA99FF8BFFF1FF42D040FEF6F88E" bold="true" box="[189,290,1856,1882]" pageId="49" pageNumber="50">Fig. 19.</emphasis>
<taxonomicName id="4C194D08FF8BFFF1FECED041FC6BF88F" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[305,959,1856,1883]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF8BFFF1FECED041FDD1F88E" box="[305,517,1856,1882]" italics="true" pageId="49" pageNumber="50">Zospeum simplex</emphasis>
<bibRefCitation id="EF884B7AFF8BFFF1FDEBD040FC6BF88F" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[532,959,1856,1883]" pageId="49" pageNumber="50" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
, (SMF 349425) Scanning Electron Microscopy images.
<emphasis id="B96DEA99FF8BFFF1FE4DD063FE13F8A9" bold="true" box="[434,455,1891,1917]" pageId="49" pageNumber="50">A</emphasis>
. Protoconch and upper teleoconch showing microstructure of superficial pitting.
<emphasis id="B96DEA99FF8BFFF1FF42D087FF05F875" bold="true" box="[189,209,1927,1953]" pageId="49" pageNumber="50">B</emphasis>
. Close up view of pitting microstructure.
<emphasis id="B96DEA99FF8BFFF1FD4DD087FD13F875" bold="true" box="[690,711,1927,1953]" pageId="49" pageNumber="50">C</emphasis>
. Last whorl with axial ribbing extending beyond peristome lip.
<emphasis id="B96DEA99FF8BFFF1FF14D0AAFED4F810" bold="true" box="[235,256,1962,1988]" pageId="49" pageNumber="50">D</emphasis>
. Close up view of second whorl showing rows of interrupted dashes of radial pitting.
<emphasis id="B96DEA99FF8BFFF1FB04D0AAFADAF810" bold="true" box="[1275,1294,1962,1988]" pageId="49" pageNumber="50">E</emphasis>
. Growth lines and radial banding on teleoconch.
<emphasis id="B96DEA99FF8BFFF1FD6FD0CDFD76F833" bold="true" box="[656,674,1997,2023]" pageId="49" pageNumber="50">F</emphasis>
. Close up view of growth lines and shell microstructure.
</paragraph>
</caption>
<paragraph id="8BA6368BFF88FFF2FF42D609FD01FE92" blockId="50.[189,1399,265,327]" pageId="50" pageNumber="51">the low, hump-like thickening of callus (not a denticle) a short distance inside the columellar side of the peristome is so far distinctive for this species.</paragraph>
<paragraph id="8BA6368BFF88FFF2FF42D66EFBBFFE26" blockId="50.[189,1399,366,498]" pageId="50" pageNumber="51">
When compared to its molecularly closest congener,
<taxonomicName id="4C194D08FF88FFF2FC9CD66EFEB4FE7F" authority="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityName="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityYear="2024" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="kolbae" status="sp. nov.">
<emphasis id="B96DEA99FF88FFF2FC9CD66EFC0EFE5C" box="[867,986,366,392]" italics="true" pageId="50" pageNumber="51">Z. kolbae</emphasis>
Jochum, Inäbnit, Kneubühler &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF88FFF2FE92D691FE1BFE7F" box="[365,463,401,427]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C194D08FF88FFF2FE1AD692FBC2FE7F" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[485,1046,401,427]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FE1AD692FDB1FE7F" box="[485,613,401,427]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
<bibRefCitation id="EF884B7AFF88FFF2FD8CD691FBC2FE7F" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[627,1046,401,427]" pageId="50" pageNumber="51" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
shows a significantly lower apertural height of an almost equal width with that of
<taxonomicName id="4C194D08FF88FFF2FCBFD6B5FC7BFE1A" authorityName="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityYear="2024" box="[832,943,436,462]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="kolbae">
<emphasis id="B96DEA99FF88FFF2FCBFD6B5FC7BFE1A" box="[832,943,436,462]" italics="true" pageId="50" pageNumber="51">Z. kolbae</emphasis>
</taxonomicName>
as well as a narrower spire angle than that of
<taxonomicName id="4C194D08FF88FFF2FEEFD6D8FEAAFE25" authorityName="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityYear="2024" box="[272,382,471,497]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="kolbae">
<emphasis id="B96DEA99FF88FFF2FEEFD6D8FEAAFE25" box="[272,382,471,497]" italics="true" pageId="50" pageNumber="51">Z. kolbae</emphasis>
</taxonomicName>
. Shell height shows more range in
<taxonomicName id="4C194D08FF88FFF2FCE3D6D8FC5FFE25" authorityName="Jochum, Inabnit, Kneubuhler &amp; Ruthensteiner" authorityYear="2024" box="[796,907,471,497]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="kolbae">
<emphasis id="B96DEA99FF88FFF2FCE3D6D8FC5FFE25" box="[796,907,471,497]" italics="true" pageId="50" pageNumber="51">Z. kolbae</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C194D08FF88FFF2FC15D6D8FBB0FE25" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[1002,1124,471,497]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FC15D6D8FBB0FE25" box="[1002,1124,471,497]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA6368BFF88FFF2FF42D519FD9FFCFE" blockId="50.[186,1399,537,810]" pageId="50" pageNumber="51">
Though
<taxonomicName id="4C194D08FF88FFF2FEE2D519FCC1FDE7" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[285,789,537,563]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FEE2D519FE47FDE7" box="[285,403,537,563]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
<bibRefCitation id="EF884B7AFF88FFF2FE67D519FCC1FDE7" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[408,789,537,563]" pageId="50" pageNumber="51" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
resembles
<taxonomicName id="4C194D08FF88FFF2FC70D519FAA3FDE7" authority="Jochum &amp; Ruthensteiner" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" box="[911,1399,537,563]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="amplioscutum" status="sp. nov.">
<emphasis id="B96DEA99FF88FFF2FC70D519FC4BFDE6" box="[911,927,537,562]" italics="true" pageId="50" pageNumber="51">Z</emphasis>
.
<emphasis id="B96DEA99FF88FFF2FC50D519FB81FDE7" box="[943,1109,537,563]" italics="true" pageId="50" pageNumber="51">amplioscutum</emphasis>
Jochum &amp; Ruthensteiner
</taxonomicName>
<taxonomicNameLabel id="A25E57E2FF88FFF2FF45D53CFECCFD82" box="[186,280,572,598]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
, it differs by its reduced extension onto the body whorl and less heavily callused parietal shield.
<taxonomicName id="4C194D08FF88FFF2FF43D560FEE2FDAD" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[188,310,607,633]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FF43D560FEE2FDAD" box="[188,310,607,633]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
</taxonomicName>
is larger, has a narrower spire angle and a markedly less textured teleoconch in comparison to that of
<taxonomicName id="4C194D08FF88FFF2FED3D583FE25FD48" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" box="[300,497,642,668]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="amplioscutum">
<emphasis id="B96DEA99FF88FFF2FED3D583FE25FD48" box="[300,497,642,668]" italics="true" pageId="50" pageNumber="51">Z. amplioscutum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C194D08FF88FFF2FDFFD583FD18FD48" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[512,716,642,668]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FDFFD583FD18FD48" box="[512,716,642,668]" italics="true" pageId="50" pageNumber="51">Zospeum simplex</emphasis>
</taxonomicName>
does not have a second lamella on the columella (aperture facing right) (
<figureCitation id="13222A0EFF88FFF2FE9AD5A5FE05FD14" box="[357,465,677,704]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="50" pageNumber="51">Fig. 18B</figureCitation>
) but rather, weak ripples of a potential double one (
<figureCitation id="13222A0EFF88FFF2FBABD5A5FB16FD14" box="[1108,1218,677,704]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="50" pageNumber="51">Fig. 18D</figureCitation>
) or a stronger, incomplete single one in dorsal view (
<figureCitation id="13222A0EFF88FFF2FD7FD5C9FD37FD37" box="[640,739,713,739]" captionStart="Fig" captionStartId="48.[189,232,1857,1883]" captionTargetBox="[186,1395,563,1819]" captionTargetId="figure-152@48.[186,1395,563,1819]" captionTargetPageId="48" captionText="Fig. 18. 3D visualizations of Micro-CT data of Zospeum simplex Inäbnit, Jochum &amp; Neubert, 2021, (NMHW-MO-113642 ex RSC 3760) from Dobravljevac jama, Gornji Brišnik, Bosnia and Herzegovina. A–F. Specimen 1 (spm1). A. Aperture view. B. Aperture facing right view. C. Apical view. D. Dorsal view. E. Aperture facing left view. F. Ventral view. G–L. Specimen 2 (spm2). Same perspectives as Specimen 1." figureDoi="http://doi.org/10.5281/zenodo.10847858" httpUri="https://zenodo.org/record/10847858/files/figure.png" pageId="50" pageNumber="51">Fig. 18J</figureCitation>
). The narrow columella (
<figureCitation id="13222A0EFF88FFF2FBF2D5C9FB59FD37" box="[1037,1165,713,739]" captionStart="Fig" captionStartId="32.[189,232,1787,1813]" captionTargetBox="[187,1396,810,1746]" captionTargetId="figure-200@32.[187,1396,810,1746]" captionTargetPageId="32" captionText="Fig. 9. Zospeum amplioscutum Jochum &amp; Ruthensteiner sp. nov., (NHMW Mol.Coll.Edlauer 19.069) with sample labels. A. Light microscopic images of paratype (NHMW Mol.Coll.Edlauer 19.069), apertural and dorsal views.B. Light microscopic images of holotype (NHMW Mol.Coll.Edlauer 19.069), aperture and dorsal views. C–H. 3D visualizations of X-ray Micro-CT data of holotype (NHMW Mol. Coll.Edlauer 19.069). C. Aperture view. D. Aperture facing right view. E. Apical view. F. Dorsal view. G. Aperture facing left view. H. Ventral view showing coiling projecting high up on shell and peristome oblique to umbilicus." figureDoi="http://doi.org/10.5281/zenodo.10847828" httpUri="https://zenodo.org/record/10847828/files/figure.png" pageId="50" pageNumber="51">Fig. 9C–D</figureCitation>
) and the elongation of the last whorl further beyond the umbilical depression (ventral side) additionally differentiate
<taxonomicName id="4C194D08FF88FFF2FF42D410FE56FCFD" authorityName="Jochum &amp; Ruthensteiner" authorityYear="2024" box="[189,386,783,809]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="amplioscutum">
<emphasis id="B96DEA99FF88FFF2FF42D410FE56FCFD" box="[189,386,783,809]" italics="true" pageId="50" pageNumber="51">Z. amplioscutum</emphasis>
</taxonomicName>
from
<taxonomicName id="4C194D08FF88FFF2FE35D410FD90FCFD" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[458,580,783,809]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FE35D410FD90FCFD" box="[458,580,783,809]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA6368BFF88FFF2FF42D450FE15FC23" blockId="50.[187,1399,848,1016]" pageId="50" pageNumber="51">
The incidence of the one solid lamella versus the two incomplete low lamellae occurred in one of the two shells scanned. In sync with
<bibRefCitation id="EF884B7AFF88FFF2FDF3D474FD28FC5A" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[524,764,884,910]" pageId="50" pageNumber="51" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">
Inäbnit
<emphasis id="B96DEA99FF88FFF2FD97D474FD48FC5A" box="[616,668,884,910]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. (2021)
</bibRefCitation>
, the shell height:shell width ratio is the most effective way to differentiate
<taxonomicName id="4C194D08FF88FFF2FE55D498FC63FC65" authority="Inabnit, Jochum &amp; Neubert, 2021" authorityName="Inabnit, Jochum &amp; Neubert" authorityYear="2021" box="[426,951,919,945]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="simplex">
<emphasis id="B96DEA99FF88FFF2FE55D498FDF0FC65" box="[426,548,919,945]" italics="true" pageId="50" pageNumber="51">Z. simplex</emphasis>
<bibRefCitation id="EF884B7AFF88FFF2FDD3D497FC63FC65" author="Inabnit T. &amp; Jochum A. &amp; Slapnik R. &amp; Neubert E." box="[556,951,919,945]" pageId="50" pageNumber="51" pagination="175 - 193" refId="ref31366" refString="Inabnit T., Jochum A., Slapnik R. &amp; Neubert E. 2021. New genetic data reveals a new species of Zospeum in Bosnia (Gastropoda, Ellobioidea, Carychiinae). ZooKeys 1071: 175 - 193. https: // doi. org / 10.3897 / zookeys. 1071.66417" type="journal article" year="2021">Inäbnit, Jochum &amp; Neubert, 2021</bibRefCitation>
</taxonomicName>
from morphs of
<taxonomicName id="4C194D08FF88FFF2FB81D498FABBFC65" authorityName="Absolon, 1916 Absolon" authorityYear="1916" box="[1150,1391,919,945]" class="Gastropoda" family="Ellobiidae" genus="Zospeum" kingdom="Animalia" order="Ellobiida" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="troglobalcanicum">
<emphasis id="B96DEA99FF88FFF2FB81D498FABBFC65" box="[1150,1391,919,945]" italics="true" pageId="50" pageNumber="51">Z. troglobalcanicum</emphasis>
</taxonomicName>
. The low, hump-like callus on the inner columellar side of the peristome is consistent in the three shells from the
<typeStatus id="54A28829FF88FFF2FED6D4DEFE88FC2C" box="[297,348,990,1016]" pageId="50" pageNumber="51">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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