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<document id="907BD1EC5C5A4DE62C3F84066F433507" ID-DOI="10.5281/zenodo.186907" ID-GBIF-Dataset="81506f05-81d1-4f04-9ff4-a40d5f257671" ID-ISSN="1175-5326" ID-Zenodo-Dep="186907" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059273533" checkinUser="plazi" docAuthor="Ba, Yi-Bin &amp; Ren, Guo-Dong" docDate="2009" docId="038087B8FFAB327485CCA0AC6499FC10" docLanguage="en" docName="zt02064p038.pdf" docOrigin="Zootaxa 2064" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Promethis" docType="key" docVersion="9" lastPageNumber="28" masterDocId="FFB9FFC0FFAB3275855BA62D6102FFDF" masterDocTitle="Taxonomy of Promethis Pascoe (Coleoptera, Tenebrionidae) on Hainan Island, China" masterLastPageNumber="38" masterPageNumber="27" pageNumber="27" updateTime="1698581798099" updateUser="plazi">
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<mods:title id="AEC855D202445CC9888E22222238675A">Taxonomy of Promethis Pascoe (Coleoptera, Tenebrionidae) on Hainan Island, China</mods:title>
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<mods:namePart id="98554E8EF80752C3751EA671E7D0A1CA">Ba, Yi-Bin</mods:namePart>
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<mods:namePart id="416D20B1B44A902C409CCEE2B2257641">Ren, Guo-Dong</mods:namePart>
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<mods:date id="0D0AA0D40699BA59523768BFB4953470">2009</mods:date>
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Key to species of
<taxonomicName id="4C294D2DFFAB32758436A0AF60EAF943" box="[365,488,1666,1692]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B95DEABCFFAB32758436A0AF60EAF943" bold="true" box="[365,488,1666,1692]" italics="true" pageId="0" pageNumber="27">Promethis</emphasis>
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in Hainan Island,
<collectingCountry id="F33E763EFFAB3275878AA0AC621CF944" box="[721,798,1665,1691]" name="China" pageId="0" pageNumber="27">China</collectingCountry>
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<keyLead id="30D8959BFFAB327585CCA0E66499F93D" box="[151,1435,1739,1762]" pageId="0" pageNumber="27">1 Anal segment without deep groove apically................................................................................................................. 2</keyLead>
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- Anal segment with deep groove apically; frontoclypeal suture shallowly concave; protibiae incurved, inner margin with one obvious tooth centrally; head densely punctate; pronotum similarly densely punctate; elytral intervals con- vex; length 18.028.0 mm .........................................................................................................................
<taxonomicName id="4C294D2DFFAB32758054A1066499F89D" box="[1295,1435,1835,1858]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="rectangula">
<emphasis id="B95DEABCFFAB32758054A1066499F89D" box="[1295,1435,1835,1858]" italics="true" pageId="0" pageNumber="27">P. rectangula</emphasis>
</taxonomicName>
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<paragraph id="8B9636AEFFAB327585CCA1666499F8BD" blockId="0.[151,1436,1739,2018]" box="[151,1435,1867,1890]" pageId="0" pageNumber="27">
<keyLead id="30D8959BFFAB327585CCA1666499F8BD" box="[151,1435,1867,1890]" pageId="0" pageNumber="27">2 Anterior margin of profemora with dense hairs in male.............................................................................................. 3</keyLead>
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<paragraph id="8B9636AEFFAB327585CCA1466499F85D" blockId="0.[151,1436,1739,2018]" box="[151,1435,1899,1922]" pageId="0" pageNumber="27">
<keyLead id="30D8959BFFAB327585CCA1466499F85D" box="[151,1435,1899,1922]" pageId="0" pageNumber="27">- Anterior margin of profemora without dense hairs in male.......................................................................................... 4</keyLead>
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</keyStep>
<paragraph id="8B9636AEFFAB327585CCA1A6649EF81D" blockId="0.[151,1436,1739,2018]" pageId="0" pageNumber="27">
<keyLead id="30D8959BFFAB327585CCA1A6649EF81D" pageId="0" pageNumber="27">
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3 Elytral rows of punctures feeble, intervals with fine and sparse granules; protibiae strongly incurved, inner margin with one obvious tooth at basal 1/
<quantity id="4CD19B4BFFAB32758775A186635AF81D" box="[558,600,1963,1986]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="0" pageNumber="27" unit="in" value="3.0">3 in</quantity>
male; length 28.036.0 mm
<emphasis id="B95DEABCFFAB327586D1A186649EF81D" box="[906,1436,1963,1986]" italics="true" pageId="0" pageNumber="27">
..........................................................
<taxonomicName id="4C294D2DFFAB32758056A186649EF81D" box="[1293,1436,1963,1986]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="vietnamica">P. vietnamica</taxonomicName>
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<keyLead id="30D8959BFFAB327585CCA1E66499F83D" box="[151,1435,1995,2018]" pageId="0" pageNumber="27">- Elytral rows of punctures strong; protibiae weakly incurved, inner margin obtusely extruded at basal 1/3, but with-</keyLead>
</paragraph>
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out obvious tooth; length 24.025.0 mm.
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................................................................................................
<taxonomicName id="4C294D2DFFAA3274805FA6B5649EFF70" box="[1284,1436,152,175]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="guangxiana">P. guangxiana</taxonomicName>
</emphasis>
<keyLead id="30D8959BFFAA327485CCA6956499FF10" box="[151,1435,184,207]" pageId="1" pageNumber="28">
<keyStep id="30DD2E0BFFAA327485CCA6956499FF10" box="[151,1435,184,207]" pageId="1" pageNumber="28">4 Anterior margin of profemora obviously bordered from base to middle in male, the border without tooth apically .. 5</keyStep>
</keyLead>
</paragraph>
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<keyLead id="30D8959BFFAA327485CCA6F5649EFEF0" pageId="1" pageNumber="28">
- Anterior margin of profemora obviously bordered from base beyond middle in male, and the border with one sharp tooth apically; anterior angles of pronotum roundly obtuse; elytral rows of punctures feeble, punctures on rows weaker or absent apically, intervals flat; length 20.030.0 mm..........................................................
<taxonomicName id="4C294D2DFFAA32748184A735649EFEF0" box="[1247,1436,280,303]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="punctatostriata">
<emphasis id="B95DEABCFFAA32748184A735649EFEF0" box="[1247,1436,280,303]" italics="true" pageId="1" pageNumber="28">P. punctatostriata</emphasis>
</taxonomicName>
</keyLead>
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<keyStep id="30DD2E0BFFAA327485CCA715649EFEB0" pageId="1" pageNumber="28">
<paragraph id="8B9636AEFFAA327485CCA7156499FE90" blockId="1.[151,1436,152,975]" box="[151,1435,312,335]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA7156499FE90" box="[151,1435,312,335]" pageId="1" pageNumber="28">5 Mentum with dense and erected yellow mane.............................................................................................................. 6</keyLead>
</paragraph>
<paragraph id="8B9636AEFFAA327485CCA775649EFEB0" blockId="1.[151,1436,152,975]" box="[151,1436,344,367]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA775649EFEB0" box="[151,1436,344,367]" pageId="1" pageNumber="28">- Mentum without manes, at most with sparse hairs.......................................................................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E0BFFAA327485CCA755649EFE30" pageId="1" pageNumber="28">
<paragraph id="8B9636AEFFAA327485CCA755649EFE70" blockId="1.[151,1436,152,975]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA755649EFE70" pageId="1" pageNumber="28">
6 Elytra with rows of punctures; intervals flat, finely punctate, each puncture with one obvious seta; metatibiae obtusely expanded interiorly near the middle; length 18.023.5 mm
<emphasis id="B95DEABCFFAA327486CCA7B5649EFE70" box="[919,1436,408,431]" italics="true" pageId="1" pageNumber="28">
.......................................................
<taxonomicName id="4C294D2DFFAA32748051A7B5649EFE70" box="[1290,1436,408,431]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="evanescens">P. evanescens</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636AEFFAA327485CCA795649EFE30" blockId="1.[151,1436,152,975]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA795649EFE30" pageId="1" pageNumber="28">
- Elytra with striae of punctures; intervals convex, finely punctate, but puncture without seta; metatibiae straight, only weakly incurved apically; length 24.028.0 mm
<emphasis id="B95DEABCFFAA327487A5A7F5649EFE30" box="[766,1436,472,495]" italics="true" pageId="1" pageNumber="28">
.................................................................................
<taxonomicName id="4C294D2DFFAA32748041A7F5649EFE30" box="[1306,1436,472,495]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="oshimana">P. oshimana</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DD2E0BFFAA327485CCA7D56499FDB0" pageId="1" pageNumber="28">
<paragraph id="8B9636AEFFAA327485CCA7D5649EFD90" blockId="1.[151,1436,152,975]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA7D5649EFD90" pageId="1" pageNumber="28">
7 Protibiae strongly incurved in apical 1/4, outer margin strongly dilated and bordered apically; pronotum square, disc coarsely and densely punctate, intervals of punctures less than punctures diameter; elytral striae deeply concave, intervals somewhat convex, and with sparse white hairs; length 11.016.0 mm
<emphasis id="B95DEABCFFAA32748154A415649EFD90" box="[1039,1436,568,591]" italics="true" pageId="1" pageNumber="28">
.................................
<taxonomicName id="4C294D2DFFAA327481B7A415649EFD90" box="[1260,1436,568,591]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="crenatostriata">P. crenatostriata</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636AEFFAA327485CCA4756499FDB0" blockId="1.[151,1436,152,975]" box="[151,1435,600,623]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA4756499FDB0" box="[151,1435,600,623]" pageId="1" pageNumber="28">- Protibiae weakly incurved, apices simple, and outer margin unbordered .................................................................... 8</keyLead>
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</keyStep>
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<keyLead id="30D8959BFFAA327485CCA455649EFD70" pageId="1" pageNumber="28">
8 Inner margin of mesotibiae with obtuse tooth centrally in male; pronotum sparsely punctate on middle and densely so on sides; elytral rows of punctures weak, intervals flat and lackluster; length 23.528.0 mm
<emphasis id="B95DEABCFFAA327481EEA4B5649EFD70" box="[1205,1436,664,687]" italics="true" pageId="1" pageNumber="28">
..
<taxonomicName id="4C294D2DFFAA32748192A4B5649EFD70" box="[1225,1436,664,687]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="valgipes" subSpecies="valgipes">P. valgipes valgipes</taxonomicName>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B9636AEFFAA327485CCA4956499FD10" blockId="1.[151,1436,152,975]" box="[151,1435,696,719]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA4956499FD10" box="[151,1435,696,719]" pageId="1" pageNumber="28">- Mesotibiae simple, inner margin without obtuse tooth................................................................................................. 9</keyLead>
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9 Protibiae strongly incurved in apical 1/
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male; terminal segment of antennae rectangular; pronotal punctures rounded, dense and joined on middle; elytral intervals very convex.......................................................................... 10
</keyLead>
</paragraph>
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<keyLead id="30D8959BFFAA327485CCA5356499FC90" pageId="1" pageNumber="28">
- Protibiae somewhat incurved in apical 1/
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male; terminal segment of antennae nearly oval; pronotal punctures fine but distinct; elytral intervals somewhat convex, and finely punctate; length 16.021.0 mm
<emphasis id="B95DEABCFFAA327481EEA5156499FC90" box="[1205,1435,824,847]" italics="true" pageId="1" pageNumber="28">
..........
<taxonomicName id="4C294D2DFFAA327481A2A5156499FC90" box="[1273,1435,824,847]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="sinuatocollis">P. sinuatocollis</taxonomicName>
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10 Terminal segment of metatarsus length equal to the sum of basal 3 segments; pronotal punctures rounded, dense, and joined; base of 1st abdominal segment without long groove; length 22.0 mm
<emphasis id="B95DEABCFFAA327486A0A5556442FC50" box="[1019,1344,888,911]" italics="true" pageId="1" pageNumber="28">
.........................
<taxonomicName id="4C294D2DFFAA327481F9A5556442FC50" box="[1186,1344,888,911]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="crassihornis" status="sp. nov.">P. crassihornis</taxonomicName>
</emphasis>
<emphasis id="B95DEABCFFAA3274801CA55A649EFC50" bold="true" box="[1351,1436,887,911]" pageId="1" pageNumber="28">
<taxonomicNameLabel id="A26E57C7FFAA3274801CA55A649EFC50" box="[1351,1436,887,911]" pageId="1" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8B9636AEFFAA327485CCA5B56499FC10" blockId="1.[151,1436,152,975]" pageId="1" pageNumber="28">
<keyLead id="30D8959BFFAA327485CCA5B56499FC10" pageId="1" pageNumber="28">
- Terminal segment of metatarsus length equal to the sum of basal 2 segments; pronotal punctures absent on sides; base of 1st abdominal segment with a long groove; length 18.023.0 mm
<emphasis id="B95DEABCFFAA32748688A5956499FC10" box="[979,1435,952,975]" italics="true" pageId="1" pageNumber="28">
................................................
<taxonomicName id="4C294D2DFFAA3274804FA5956499FC10" box="[1300,1435,952,975]" class="Insecta" family="Tenebrionidae" genus="Promethis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="28" phylum="Arthropoda" rank="species" species="pauperula">P. pauperula</taxonomicName>
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