166 lines
19 KiB
XML
166 lines
19 KiB
XML
<document id="C38B78D5DC09685E4F24916073819D0D" ID-DOI="10.5281/zenodo.191594" ID-GBIF-Dataset="6f65b06c-5600-4172-9ee2-854f09ef4455" ID-ISSN="1175-5326" ID-Zenodo-Dep="191594" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461129936904" checkinUser="plazi" docAuthor="Haddad, Charles R., Lyle, Robin, Bosselaers, Jan & Ramirez, Martin J." docDate="2009" docId="038087A2DA5B4506FF4FFBE76765FA75" docLanguage="en" docName="zt02296p038.pdf" docOrigin="Zootaxa 2296" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Austrachelas sexoculata Haddad, Lyle, Bosselaers & Ramirez, 2009, n. sp." docType="treatment" docVersion="10" lastPageNumber="27" masterDocId="FFB9FFDADA42451CFFD8FFC5641CFFEC" masterDocTitle="A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="26" updateTime="1698597546718" updateUser="plazi">
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<mods:title id="D935AE16E8AF982306BFB8CDC0D9909C">A revision of the endemic South African spider genus Austrachelas, with its transfer to the Gallieniellidae (Arachnida: Araneae)</mods:title>
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<mods:name id="43C2F24710CB557C5ECC66AD7505DC65" type="personal">
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<mods:namePart id="F2CB2844FE0D54D54D2E2C376260F0B2">Haddad, Charles R.</mods:namePart>
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<mods:namePart id="F3C4A56EEF8CA070D00128FFFDBD63D3">Lyle, Robin</mods:namePart>
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<mods:namePart id="5802469985CA34DF4A0E48C9C88E9318">Bosselaers, Jan</mods:namePart>
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<mods:namePart id="3D94939966C9A33AAF9EC57B5742DB7D">Ramirez, Martin J.</mods:namePart>
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<mods:title id="50648E7862A91AFCEAB020B9BB79163B">Zootaxa</mods:title>
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<mods:part id="BB1EE1EDBFEA7EA919D731F273672BDE">
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<mods:date id="B3BF99B074F74D59AD56ED6832802447">2009</mods:date>
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<mods:number id="D2DF1FD3A564B2358444B46DD75EF217">2296</mods:number>
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<treatment id="038087A2DA5B4506FF4FFBE76765FA75" ID-DOI="http://doi.org/10.5281/zenodo.6225469" ID-GBIF-Taxon="119626930" ID-Zenodo-Dep="6225469" LSID="urn:lsid:plazi:treatment:038087A2DA5B4506FF4FFBE76765FA75" httpUri="http://treatment.plazi.org/id/038087A2DA5B4506FF4FFBE76765FA75" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
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<subSubSection id="C333653FDA5B4505FF4FFBE76545FBB3" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8B9636B4DA5B4505FF4FFBE7661AFBD0" blockId="25.[151,518,1058,1119]" box="[151,518,1058,1084]" pageId="25" pageNumber="26">
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<heading id="D0DE81D8DA5B4505FF4FFBE7661AFBD0" bold="true" box="[151,518,1058,1084]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="B95DEAA6DA5B4505FF4FFBE7661AFBD0" bold="true" box="[151,518,1058,1084]" pageId="25" pageNumber="26">
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<taxonomicName id="4C294D37DA5B4505FF4FFBE765A0FBD0" box="[151,444,1058,1084]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="sexoculata" status="sp. nov.">
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<emphasis id="B95DEAA6DA5B4505FF4FFBE765A0FBD0" bold="true" box="[151,444,1058,1084]" italics="true" pageId="25" pageNumber="26">Austrachelas sexoculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26E57DDDA5B4505FE1CFBE7661AFBD0" box="[452,518,1058,1084]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9636B4DA5B4505FF4FFB806545FBB3" blockId="25.[151,518,1058,1119]" box="[151,345,1093,1119]" pageId="25" pageNumber="26">
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(
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<figureCitation id="13122A31DA5B4505FF47FB8064E5FBB3" box="[159,249,1093,1119]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="25" pageNumber="26">Figs 13</figureCitation>
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,
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<figureCitation id="13122A31DA5B4505FEDEFB80654DFBB3" box="[262,337,1093,1119]" captionStart="FIGURES 76 – 81" captionStartId="26.[151,269,1147,1171]" captionTargetBox="[223,1370,200,1113]" captionTargetId="figure@26.[223,1380,194,1122]" captionTargetPageId="26" captionText="FIGURES 76 – 81. Genitalic morphology of Austrachelas species. (76 – 77) A. sexoculata n. sp.: male palp, ventral view (76), retrolateral view (77); (78 – 81) A. wassenaari n. sp.: female epigyne, ventral view (78), dorsal view (79), male palp, ventral view (80), retrolateral view (81). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191605/files/figure.png" pageId="25" pageNumber="26">76, 77</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA5B4505FF4FFB576142FB40" box="[151,1374,1170,1196]" pageId="25" pageNumber="26" type="diagnosis">
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<paragraph id="8B9636B4DA5B4505FF4FFB576142FB40" blockId="25.[151,1437,1170,1970]" box="[151,1374,1170,1196]" pageId="25" pageNumber="26">
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<emphasis id="B95DEAA6DA5B4505FF4FFB57650CFB40" bold="true" box="[151,272,1170,1196]" pageId="25" pageNumber="26">Diagnosis</emphasis>
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. This species is unique in the genus by the reduction and absence of the anterior median eyes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA5B4505FF1EFB7D67D0FB16" pageId="25" pageNumber="26" type="etymology">
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<paragraph id="8B9636B4DA5B4505FF1EFB7D67D0FB16" blockId="25.[151,1437,1170,1970]" pageId="25" pageNumber="26">
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<emphasis id="B95DEAA6DA5B4505FF1EFB7D6557FB3E" bold="true" box="[198,331,1208,1234]" pageId="25" pageNumber="26">Etymology</emphasis>
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. This species is named for the Latin “
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<emphasis id="B95DEAA6DA5B4505FCE4FB7C677DFB3E" box="[828,865,1209,1234]" italics="true" pageId="25" pageNumber="26">sex</emphasis>
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” (six) and “
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<emphasis id="B95DEAA6DA5B4505FBDCFB7C6078FB3E" box="[1028,1124,1209,1234]" italics="true" pageId="25" pageNumber="26">ocularis</emphasis>
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” (eyes), and reflects the presence of only six eyes and the absence of the anterior median eyes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA5B4506FF1EFAC365BAFAC9" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
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<paragraph id="8B9636B4DA5B4505FF1EFAC367AEFAAB" blockId="25.[151,1437,1170,1970]" pageId="25" pageNumber="26">
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<emphasis id="B95DEAA6DA5B4505FF1EFAC365C5FACC" bold="true" box="[198,473,1286,1312]" pageId="25" pageNumber="26">Male. Measurements:</emphasis>
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CL 4.43, CW 3.05, FL 0.45, AL 4.35, AW 2.50, SL 2.40, SW 1.60, TL 9.15, ALE–ALE 0.32, PME–PME 0.07, PME–PLE 0.12, PLE–PLE 0.58.
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</paragraph>
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<paragraph id="8B9636B4DA5B4505FF1DFA9165F3FA57" blockId="25.[151,1437,1170,1970]" pageId="25" pageNumber="26">Length of leg segments (sequence from femur to tarsus, and total): I 3.20 + 1.75 + 2.30 + 1.80 + 1.05 = 10.10; II 3.00 + 1.70 + 2.18 + 1.70 + 0.95 = 9.53; III 2.55 + 1.28 + 1.48 + 1.82 + 0.92 = 8.05; IV 3.43 + 1.70 + 2.43 + 3.00 + 1.12 = 11.68.</paragraph>
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<paragraph id="8B9636B4DA5B4505FF1DFA0D6763F85E" blockId="25.[151,1437,1170,1970]" pageId="25" pageNumber="26">
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Carapace deep red, paler along midline (
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<figureCitation id="13122A31DA5B4505FD79FA0D66E4FA0E" box="[673,760,1480,1506]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
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); AME absent, ALE large, separated by distance equal to 1½ times their diameter; clypeus height equal to 1¼ times ALE diameter; PER straight, laterals larger than medians; PME separated by distance equal to ¾ their diameter, PME separated from PLE by distance equal to 1½ times PME diameter. Chelicerae deep red-brown, anterior margin with coarse ridges; promargin with three teeth, proximal and median teeth larger, subequal in size, closer to each other than to distal tooth; retromargin with single small denticle close to fang base. Legs all orange-brown, paler distally. Leg spination: femora: I pl 1 do 1, II pl 1 do 1, III pl 1 do 2 rl 1, IV do 2; patellae: spineless; tibiae: I & II spineless, III pl 2 rl 2 plv 2 rlv 1 vt 2, IV pl 2 rl 2 plv 2 rlv 2 vt 2; metatarsi: I & II spineless, strongly scopulate, III & IV scopulate in distal half, III pl 6 rl 4 plv 1 rlv 1 vt 2, IV pl 5 rl 5 plv 2 rlv 1 vt 1. Abdomen dark grey dorsally (
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<figureCitation id="13122A31DA5B4505FB3BF9386120F8FB" box="[1251,1340,1789,1815]" captionStart="FIGURES 1 – 15" captionStartId="2.[151,269,1353,1377]" captionTargetBox="[151,1435,315,1326]" captionTargetId="figure@2.[151,1436,313,1328]" captionTargetPageId="2" captionText="FIGURES 1 – 15. Digital microscope photographs of the habitus of Austrachelas species: (1) A. bergi n. sp., female; (2) A. incertus Lawrence, 1938, female; (3) same, male; (4) A. kalaharinus n. sp., male; (5) A. merwei n. sp., female; (6) same, male; (7) A. natalensis Lawrence, 1942, female; (8) same, male; (9) A. pondoensis n. sp., female; (10) same, male; (11) A. reavelli n. sp., female; (12) same, male; (13) A. sexoculata n. sp., male; (14) A. wassenaari n. sp., female; (15) same, male. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/191595/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
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), cream laterally and ventrally; dorsum and venter lacking scutum or other sclerites. Palpal tibia with spoon-shaped prolateral ventral apophysis and simple triangular retrolateral apophysis; median tegular apophysis broad, hook-like; embolus arc-shaped, with small tooth-like projections prolaterally and retrolaterally on distal margin of embolus (
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<figureCitation id="13122A31DA5B4505FE5BF85D6610F85E" box="[387,524,1944,1970]" captionStart="FIGURES 76 – 81" captionStartId="26.[151,269,1147,1171]" captionTargetBox="[223,1370,200,1113]" captionTargetId="figure@26.[223,1380,194,1122]" captionTargetPageId="26" captionText="FIGURES 76 – 81. Genitalic morphology of Austrachelas species. (76 – 77) A. sexoculata n. sp.: male palp, ventral view (76), retrolateral view (77); (78 – 81) A. wassenaari n. sp.: female epigyne, ventral view (78), dorsal view (79), male palp, ventral view (80), retrolateral view (81). Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/191605/files/figure.png" pageId="25" pageNumber="26">Figs 76, 77</figureCitation>
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); palpal spination: femora do 2.
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</paragraph>
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<caption id="DF56663CDA584506FF4FFBBE673BFB34" httpUri="https://zenodo.org/record/191605/files/figure.png" pageId="26" pageNumber="27" targetBox="[223,1370,200,1113]" targetPageId="26">
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<paragraph id="8B9636B4DA584506FF4FFBBE673BFB34" blockId="26.[151,1436,1147,1240]" pageId="26" pageNumber="27">
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<emphasis id="B95DEAA6DA584506FF4FFBBE6542FB7F" bold="true" box="[151,350,1147,1171]" pageId="26" pageNumber="27">FIGURES 76–81.</emphasis>
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Genitalic morphology of
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<taxonomicName id="4C294D37DA584506FDADFBB9671CFB7F" box="[629,768,1148,1171]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
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<emphasis id="B95DEAA6DA584506FDADFBB9671CFB7F" box="[629,768,1148,1171]" italics="true" pageId="26" pageNumber="27">Austrachelas</emphasis>
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</taxonomicName>
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species. (76–77)
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<taxonomicName id="4C294D37DA584506FC65FBB96052FB7F" box="[957,1102,1148,1171]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="sexoculata" status="sp. nov.">
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<emphasis id="B95DEAA6DA584506FC65FBB96052FB7F" box="[957,1102,1148,1171]" italics="true" pageId="26" pageNumber="27">A. sexoculata</emphasis>
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</taxonomicName>
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<emphasis id="B95DEAA6DA584506FB8DFBBE608DFB7F" bold="true" box="[1109,1169,1147,1171]" pageId="26" pageNumber="27">
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<taxonomicNameLabel id="A26E57DDDA584506FB8DFBBE608DFB7F" box="[1109,1169,1147,1171]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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: male palp, ventral view (76), retrolateral view (77); (78–81)
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<taxonomicName id="4C294D37DA584506FDC2FB5A66ADFB5A" box="[538,689,1183,1206]" class="Arachnida" family="Gallieniellidae" genus="Austrachelas" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="wassenaari" status="sp. nov.">
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<emphasis id="B95DEAA6DA584506FDC2FB5A66ADFB5A" box="[538,689,1183,1206]" italics="true" pageId="26" pageNumber="27">A. wassenaari</emphasis>
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</taxonomicName>
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<emphasis id="B95DEAA6DA584506FD6EFB5B66EEFB5A" bold="true" box="[694,754,1182,1206]" pageId="26" pageNumber="27">
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<taxonomicNameLabel id="A26E57DDDA584506FD6EFB5B66EEFB5A" box="[694,754,1182,1206]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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: female epigyne, ventral view (78), dorsal view (79), male palp, ventral view (80), retrolateral view (81). Scale bars = 0.1mm.
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</paragraph>
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</caption>
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<paragraph id="8B9636B4DA584506FF1EFACF65BAFAC9" blockId="26.[151,1436,1290,1433]" box="[198,422,1290,1317]" pageId="26" pageNumber="27">
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<emphasis id="B95DEAA6DA584506FF1EFACF653DFAC8" bold="true" box="[198,289,1290,1316]" pageId="26" pageNumber="27">Female</emphasis>
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: unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C333653FDA584506FF1EFAF465C0FA9E" pageId="26" pageNumber="27" type="materials_examined">
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<paragraph id="8B9636B4DA584506FF1EFAF465C0FA9E" blockId="26.[151,1436,1290,1433]" pageId="26" pageNumber="27">
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<emphasis id="B95DEAA6DA584506FF1EFAF46569FAA7" bold="true" box="[198,373,1329,1355]" pageId="26" pageNumber="27">
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<typeStatus id="54928816DA584506FF1EFAF4651FFAA7" box="[198,259,1329,1355]" pageId="26" pageNumber="27">Type</typeStatus>
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material
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</emphasis>
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:
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<typeStatus id="54928816DA584506FE5CFAF765EDFAA0" box="[388,497,1330,1356]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
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3:
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<collectingCountry id="F33E7624DA584506FDC4FAF766E9FAA0" box="[540,757,1330,1356]" name="South Africa" pageId="26" pageNumber="27">SOUTH AFRICA</collectingCountry>
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:
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<emphasis id="B95DEAA6DA584506FCD8FAF7600CFAA7" box="[768,1040,1330,1355]" italics="true" pageId="26" pageNumber="27">Eastern Cape Province</emphasis>
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, East London [
|
||
<geoCoordinate id="EE1D5073DA584506FB19FAF7613DFAA0" box="[1217,1313,1330,1356]" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="925" value="-33.033333">33°02'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1D5073DA584506FAF5FAF7618CFAA0" box="[1325,1424,1330,1356]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="925" value="27.916666">27°55'E</geoCoordinate>
|
||
],
|
||
<date id="FF971074DA584506FF4FFA9D64F0FA9E" box="[151,236,1368,1394]" pageId="26" pageNumber="27" value="1916-05">V.1916</date>
|
||
, Dr Rattray (
|
||
<collectionCode id="ED38AE71DA584506FE5DFA9D65CCFA9E" box="[389,464,1368,1394]" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="26" pageNumber="27">AMG</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C333653FDA584506FF1EFABB6765FA75" box="[198,889,1406,1433]" pageId="26" pageNumber="27" type="distribution">
|
||
<paragraph id="8B9636B4DA584506FF1EFABB6765FA75" blockId="26.[151,1436,1290,1433]" box="[198,889,1406,1433]" pageId="26" pageNumber="27">
|
||
<emphasis id="B95DEAA6DA584506FF1EFABB6543FA74" bold="true" box="[198,351,1406,1432]" pageId="26" pageNumber="27">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="54928816DA584506FDAAFABA66B8FA75" box="[626,676,1407,1433]" pageId="26" pageNumber="27">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="13122A31DA584506FCCBFABA6777FA75" box="[787,875,1407,1433]" captionStart="FIGURE 82" captionStartId="28.[151,255,1122,1146]" captionTargetBox="[274,1329,188,1095]" captionTargetId="figure@28.[268,1335,181,1109]" captionTargetPageId="28" captionText="FIGURE 82. Distribution of the genus Austrachelas, endemic to South Africa." httpUri="https://zenodo.org/record/191606/files/figure.png" pageId="26" pageNumber="27">Fig. 82</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |