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<document id="8A7F8989FC5C64E5E37C131DFCE5E033" ID-CLB-Dataset="22343" ID-DOI="10.5852/ejt.2015.167" ID-GBIF-Dataset="72b00f76-b85d-47c0-877f-511110a6a0f4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3805815" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588788296119" checkinUser="carolina" docAuthor="Hughes, Mark, Girmansyah, Deden &amp; Ardi, Wisnu Handoyo" docDate="2015" docId="03FCF055FFEDFA59D89C39EEFC306F0B" docLanguage="en" docName="ejt-2015-167.pdf.imf" docOrigin="European Journal of Taxonomy 167" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia pseudoscottii Girm. 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FFC5882DFFF3FA46D942382BFFB66D01" masterDocTitle="Further discoveries in the ever-expanding genus Begonia (Begoniaceae): fifteen new species from Sumatra" masterLastPageNumber="40" masterPageNumber="1" pageNumber="31" updateTime="1698838759931" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B776DD866AB0976F8FDD90A1DD701E89" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B9206F0A45570A8B80CB55CE93A769D">
<mods:title id="0E3210D51CA32D75A8D571F7BE5EC860">Further discoveries in the ever-expanding genus Begonia (Begoniaceae): fifteen new species from Sumatra</mods:title>
</mods:titleInfo>
<mods:name id="1920E7F53BE0F257265041AA8BA764E2" type="personal">
<mods:role id="2330CDC17F647CA1025E35B00C79C7A1">
<mods:roleTerm id="B366AB284700DFE82BA1F0A7B60A4393">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7DCA48B6B866E35CBE0765CC66BCAAE">Hughes, Mark</mods:namePart>
<mods:affiliation id="4D1C686C8DA85C03322510AAAD324CED">Royal Botanic Garden Edinburgh, 20 a Inverleith Row, EH 3 5 LR, UK. &amp; Corresponding author: m. hughes @ rbge. ac. uk</mods:affiliation>
<mods:nameIdentifier id="BBBA929A078F2AFFD1FC2E68B4CB0A18" type="email">m.hughes@rbge.ac.uk</mods:nameIdentifier>
</mods:name>
<mods:name id="9345260A8EDB691196930D8FEAC197CB" type="personal">
<mods:role id="52FD8F8CDFF1DB3825C1360F047B4B86">
<mods:roleTerm id="01CCDC0B3EC2BE2B7413470665492EBB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F072CD16347A60CF38795C4A79BCA135">Girmansyah, Deden</mods:namePart>
<mods:affiliation id="A1F93ED2B4B5FEF863305E6D539A5433">Herbarium Bogoriense, Botany Division, Research Center for Biology, Indonesian Institute of Sciences, Cibinong Science Center (CSC), Jl. Raya Jakarta-Bogor Km 46, Cibinong, Bogor 16911, Indonesia. &amp; Email: deden _ bo @ yahoo. com</mods:affiliation>
<mods:nameIdentifier id="3FBDE29FC0376D0BB8FEE3CAD0F85726" type="email">deden_bo@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="8FBDAA00AE38D76218DEE525A04384EE" type="personal">
<mods:role id="DCEE5DA480CB929CBFAF6CF4223FBEED">
<mods:roleTerm id="A2CCC4940BEEBC3612919470824B096E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16666FDF423E350B15F095E05C74CB1E">Ardi, Wisnu Handoyo</mods:namePart>
<mods:affiliation id="D890B758AD7B51AABC7177D8F017279E">Center For Plant Conservation-Bogor Botanic Garden, Jl. Ir. H. Juanda No. 13, P. O. Box 309, Bogor 16003, Indonesia. &amp; Email: wisnu. handoyo. ardi @ lipi. go. id</mods:affiliation>
<mods:nameIdentifier id="3613B431F4357691A645D95A2D32AB69" type="email">handoyo.ardi@lipi.go.id</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="5BEC0E8C5161B8C3BE94B30CE18D459B">text</mods:typeOfResource>
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<mods:titleInfo id="47D59FD55DCB449F5E5ACC5196638E9F">
<mods:title id="CC34913A841AE8E2296A65F152A44D84">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="81BF0D74B74F58A220D4A46C55A5EB14">
<mods:date id="3DECCC6C1B0EB17883C7B18F390DB9ED">2015</mods:date>
<mods:detail id="48E0BFBC0AA4708F1B4285145B745CC2" type="pubDate">
<mods:number id="2225B7DD113DA925338FA3E372201C4D">2015-12-21</mods:number>
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<mods:detail id="573F6DD32E4A93799EA7EEE8E9EF1030" type="issue">
<mods:number id="A57C38BA7004E8AFF47EAC0B9614F786">167</mods:number>
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<mods:start id="995E10EB826570BFF023F3446DF8F98D">1</mods:start>
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<mods:classification id="1C907F5B386AA86DB601F10C2081F7FC">journal article</mods:classification>
<mods:identifier id="A295CB69648631FFA6207CC7A9DFD32F" type="CLB-Dataset">22343</mods:identifier>
<mods:identifier id="B30CBEFC7C0CE23AC44A8642738F7545" type="DOI">10.5852/ejt.2015.167</mods:identifier>
<mods:identifier id="9B29F50629B617C447D3DD0BD0393437" type="GBIF-Dataset">72b00f76-b85d-47c0-877f-511110a6a0f4</mods:identifier>
<mods:identifier id="4A2C9D9FBC814868EE26684FB7681F8E" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="5B75F8B5FF65C5E18A3EB5D52AAB7B5C" type="Zenodo-Dep">3805815</mods:identifier>
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<treatment id="03FCF055FFEDFA59D89C39EEFC306F0B" ID-DOI="http://doi.org/10.5281/zenodo.3815871" ID-GBIF-Taxon="163970623" ID-Zenodo-Dep="3815871" LSID="urn:lsid:plazi:treatment:03FCF055FFEDFA59D89C39EEFC306F0B" httpUri="http://treatment.plazi.org/id/03FCF055FFEDFA59D89C39EEFC306F0B" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<subSubSection id="C34F12C8FFEDFA58D89C39EEFBE36CDE" box="[478,1109,453,480]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BEA4143FFEDFA58D89C39EEFBE36CDE" blockId="30.[478,1109,453,552]" box="[478,1109,453,480]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D89C39EEFBE36CDE" box="[478,1109,453,480]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="4C553AC0FFEDFA58D89C39EEFC826CDE" ID-CoL="87Z4L" authority="Girm." authorityName="Girm." authorityYear="2015" box="[478,820,453,480]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="pseudoscottii" status="sp. nov.">
<emphasis id="B9219D51FFEDFA58D89C39EEFD506CE1" bold="true" box="[478,742,453,480]" italics="true" pageId="30" pageNumber="31">Begonia pseudoscottii</emphasis>
Girm.
</taxonomicName>
<taxonomicNameLabel id="A212202AFFEDFA58DA7939EDFC236CE1" box="[827,917,454,480]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
§
<taxonomicName id="4C553AC0FFEDFA58DAF039EEFBE36CDE" authorityName="Warb. (Warburg" authorityYear="1894" baseAuthorityName="Hassk." box="[946,1109,453,479]" class="Magnoliopsida" family="Begoniaceae" genus="Sphenanthera" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
<emphasis id="B9219D51FFEDFA58DAF039EEFBE36CDE" box="[946,1109,453,479]" italics="true" pageId="30" pageNumber="31">Sphenanthera</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFEDFA58DB0539C2FCE26F29" pageId="30" pageNumber="31" type="description">
<paragraph id="8BEA4143FFEDFA58DB0539C2FC5B6F05" blockId="30.[478,1109,453,552]" box="[583,1005,489,516]" pageId="30" pageNumber="31">urn:lsid:ipni.org:names:77151650-1</paragraph>
<paragraph id="8BEA4143FFEDFA58DB9D3A25FCE26F29" blockId="30.[478,1109,453,552]" box="[735,852,525,552]" pageId="30" pageNumber="31">
<figureCitation id="136E5DC6FFEDFA58DB9D3A25FC9C6F29" box="[735,810,526,552]" captionStart="Fig" captionStartId="8.[189,232,1965,1991]" captionTargetBox="[189,1398,586,1923]" captionTargetId="figure@8.[189,1398,585,1923]" captionTargetPageId="8" captionText="Fig. 2. Map of Sumatra showing the full known species distributions based on collection localities. Each point may represent more than one collection." figureDoi="http://doi.org/10.5281/zenodo.3805823" httpUri="https://zenodo.org/record/3805823/files/figure.png" pageId="30" pageNumber="31">Figs 2</figureCitation>
,
<figureCitation id="136E5DC6FFEDFA58DA753A26FCE26F29" box="[823,852,525,552]" captionStart="Fig" captionStartId="31.[189,232,1896,1922]" captionTargetBox="[189,1398,818,1855]" captionTargetId="figure@31.[189,1398,818,1855]" captionTargetPageId="31" captionText="Fig. 15. Begonia pseudoscottii Girm sp. nov. A. Habit. B. Inflorescence at the base of the plant, with mature fruit below and male and female flowers open at the top. C. Male flower. All photographs from a plant cultivated in Cibodas Botanic Garden (accession number C20090980/IY80, derived from material collected in Gunung Leuser National Park). Scale bars on insets = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3805869" httpUri="https://zenodo.org/record/3805869/files/figure.png" pageId="30" pageNumber="31">15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFEDFA58D9FF3A7BFDF66E51" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BEA4143FFEDFA58D9FF3A7BFE806F6B" blockId="30.[189,310,592,618]" box="[189,310,592,618]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D9FF3A7BFE806F6B" bold="true" box="[189,310,592,618]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9219D51FFEDFA58D9FF3A7BFE806F6B" bold="true" box="[189,310,592,618]" pageId="30" pageNumber="31">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3AABFDF66E51" blockId="30.[189,1399,639,848]" pageId="30" pageNumber="31">
Differs from
<taxonomicName id="4C553AC0FFEDFA58D8143AABFCB06F9B" authority="Tebbitt (Tebbit 2005)" authorityName="Tebbitt (Tebbitt" authorityYear="2005" box="[342,774,639,666]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="scottii">
<emphasis id="B9219D51FFEDFA58D8143AABFDB16F98" box="[342,519,640,665]" italics="true" pageId="30" pageNumber="31">Begonia scottii</emphasis>
Tebbitt (Tebbit 2005)
</taxonomicName>
in bearing fruit in clusters of 10–15 (not 2 or 4), with the fruit being fleshier and rounder, with ridges rather than small wings. The infructescence in terms of fruit number and shape is more similar to that of
<taxonomicName id="4C553AC0FFEDFA58DA783AE2FB5A6FE3" authority="Blume (Blume 1827)" authorityName="Blume (Blume" authorityYear="1827" box="[826,1260,712,738]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="multangula">
<emphasis id="B9219D51FFEDFA58DA783AE2FC526FE3" box="[826,996,712,738]" italics="true" pageId="30" pageNumber="31">B. multangula</emphasis>
Blume (
<bibRefCitation id="EFC43CB2FFEDFA58DD0C3AE3FB526FE3" author="Blume C. L." box="[1102,1252,712,738]" pageId="30" pageNumber="31" refId="ref17665" refString="Blume C. L. 1827. Enumeratio Plantarum Javae et insularum adjacentium minus cognitarum vel novarum ex herbariis Reinwardtii, Kuhlii, Hasseltii et Blumii curavit Carol. Ludov. Blume. Apud J. W. van Leeuwen, Lugduni Batavorum [Leiden]. http: // dx. doi. org / 10.5962 / bhl. title. 44901" type="book" year="1827">Blume 1827</bibRefCitation>
)
</taxonomicName>
, which has lobed leaves that are either subglabrous or have white hairs (not pink), and apical (not basal) more lax infructesences. The flowers have pink centres to the tepals (uniform white in
<taxonomicName id="4C553AC0FFEDFA58DD113B3AFB0A6E2B" authorityName="Tebbitt (Tebbitt" authorityYear="2005" box="[1107,1212,785,810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="scottii">
<emphasis id="B9219D51FFEDFA58DD113B3AFB0A6E2B" box="[1107,1212,785,810]" italics="true" pageId="30" pageNumber="31">B. scottii</emphasis>
</taxonomicName>
) and the leaves have deep pink hairs (not white).
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFEDFA58D9FF3B5CFEA06EE4" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BEA4143FFEDFA58D9FF3B5CFEF26E90" blockId="30.[189,324,887,913]" box="[189,324,887,913]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D9FF3B5CFEF26E90" bold="true" box="[189,324,887,913]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9219D51FFEDFA58D9FF3B5CFEF26E90" bold="true" box="[189,324,887,913]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3B8CFEA06EE4" blockId="30.[189,1398,935,998]" pageId="30" pageNumber="31">
The epithet refers to the previous confusion of this species with
<taxonomicName id="4C553AC0FFEDFA58DA9F3B83FEB96EE7" authority="(Hughes &amp; Girmansyah 2011 b)" baseAuthorityName="Hughes &amp; Girmansyah" baseAuthorityYear="2011" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="scottii">
<emphasis id="B9219D51FFEDFA58DA9F3B83FBFC6EC0" box="[989,1098,936,961]" italics="true" pageId="30" pageNumber="31">B. scottii</emphasis>
(
<bibRefCitation id="EFC43CB2FFEDFA58DD1F3B8CFEB16EE7" author="Hughes M. &amp; Girmansyah D." pageId="30" pageNumber="31" pagination="83 - 96" refId="ref18049" refString="Hughes M. &amp; Girmansyah D. 2011 b. Searching for Sumatran Begonia described by William Jack: following in the footsteps of a 19 th century Scottish botanist. Gardens' Bulletin Singapore 63 (1 &amp; 2): 83 - 96." type="journal article" year="2011">Hughes &amp; Girmansyah 2011b</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFEDFA58D9FF3C26FE3F6813" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="8BEA4143FFEDFA58D9FF3C26FF4C6926" blockId="30.[189,250,1037,1063]" box="[189,250,1037,1063]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D9FF3C26FF4C6926" bold="true" box="[189,250,1037,1063]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9219D51FFEDFA58D9FF3C26FF4C6926" bold="true" box="[189,250,1037,1063]" pageId="30" pageNumber="31">
<typeStatus id="54EEFFE1FFEDFA58D9FF3C26FF4C6926" box="[189,250,1037,1063]" pageId="30" pageNumber="31">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3C16FB0D6956" blockId="30.[189,1211,1085,1111]" box="[189,1211,1085,1111]" pageId="30" pageNumber="31">
<collectingRegion id="49918FA1FFEDFA58D9FF3C16FEF86956" box="[189,334,1085,1111]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">SUMATRA</collectingRegion>
:
<collectingRegion id="49918FA1FFEDFA58D8153C16FE2E6956" box="[343,408,1085,1111]" country="Indonesia" name="Aceh" pageId="30" pageNumber="31">Aceh</collectingRegion>
, Boer ni Bias,
<quantity id="4CADECA6FFEDFA58DB073C16FD166956" box="[581,672,1085,1111]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="m" value="1300.0">1300 m</quantity>
,
<date id="FFEB6783FFEDFA58DBEE3C16FCF86956" box="[684,846,1085,1111]" pageId="30" pageNumber="31" value="1934-08-31">31 Aug. 1934</date>
,
<emphasis id="B9219D51FFEDFA58DA183C15FB956956" box="[858,1059,1085,1111]" italics="true" pageId="30" pageNumber="31">Van Steenis 6207</emphasis>
(holo-:
<collectionCode id="ED44D986FFEDFA58DDC33C16FB186956" box="[1153,1198,1085,1111]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="30" pageNumber="31" type="Herbarium">BO</collectionCode>
).
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3C54FE026998" blockId="30.[189,436,1151,1177]" box="[189,436,1151,1177]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D9FF3C54FE026998" bold="true" box="[189,436,1151,1177]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9219D51FFEDFA58D9FF3C54FE026998" bold="true" box="[189,436,1151,1177]" pageId="30" pageNumber="31">Additional material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3C84FE3F6813" blockId="30.[189,1399,1199,1298]" pageId="30" pageNumber="31">
<collectingRegion id="49918FA1FFEDFA58D9FF3C84FEF869C8" box="[189,334,1199,1225]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">SUMATRA</collectingRegion>
:
<collectingRegion id="49918FA1FFEDFA58D8193C84FDBB69C8" box="[347,525,1199,1225]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="31">North Sumatra</collectingRegion>
, Gunong Batu Lopang,
<quantity id="4CADECA6FFEDFA58DA6F3C84FCCE69C8" box="[813,888,1199,1225]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="10.0">10 km</quantity>
ESE of Lake Prapat,
<quantity id="4CADECA6FFEDFA58DDC23C84FA9169C8" box="[1152,1319,1199,1226]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="30" pageNumber="31" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400–1500 m</quantity>
,
<date id="FFEB6783FFEDFA58DC773C84FF4C69EC" pageId="30" pageNumber="31" value="1972-07-08">8 Jul. 1972</date>
,
<emphasis id="B9219D51FFEDFA58D8473CF8FD1769EC" box="[261,673,1235,1262]" italics="true" pageId="30" pageNumber="31">de Wilde &amp; de Wilde-Duyfies 13531</emphasis>
(
<collectionCode id="ED44D986FFEDFA58DBF23CF8FD6A69EC" box="[688,732,1235,1261]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="30" pageNumber="31" type="Herbarium">BO</collectionCode>
,
<collectionCode id="ED44D986FFEDFA58DBA53CFFFCB669EF" box="[743,768,1236,1262]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="30" pageNumber="31" type="Herbarium">K</collectionCode>
);
<collectingRegion id="49918FA1FFEDFA58DA503CF8FCE569EC" box="[786,851,1235,1261]" country="Indonesia" name="Aceh" pageId="30" pageNumber="31">Aceh</collectingRegion>
, Gunung Kemiri,
<quantity id="4CADECA6FFEDFA58DD603CF8FBCA69EF" box="[1058,1148,1235,1262]" metricMagnitude="3" metricUnit="m" metricValue="1.696" pageId="30" pageNumber="31" unit="m" value="1696.0">1696 m</quantity>
,
<date id="FFEB6783FFEDFA58DDC53CF8FA9569EC" box="[1159,1315,1235,1262]" pageId="30" pageNumber="31" value="2008-03-11">11 Mar. 2008</date>
,
<emphasis id="B9219D51FFEDFA58DC6F3CF8FEE36810" italics="true" pageId="30" pageNumber="31">
Wilkie et al.
<collectionCode id="ED44D986FFEDFA58D9BD3CDCFE9D6810" box="[255,299,1271,1297]" country="Thailand" name="Paleontological Collections" pageId="30" pageNumber="31">PW</collectionCode>
678
</emphasis>
(
<collectionCode id="ED44D986FFEDFA58D8243CD3FECE6813" box="[358,376,1272,1298]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="30" pageNumber="31" type="Herbarium">E</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFEDFA58D9FF3D12FE1F6AEA" pageId="30" pageNumber="31" type="description">
<paragraph id="8BEA4143FFEDFA58D9FF3D12FEF86852" blockId="30.[189,334,1337,1363]" box="[189,334,1337,1363]" pageId="30" pageNumber="31">
<heading id="D0A2F62FFFEDFA58D9FF3D12FEF86852" bold="true" box="[189,334,1337,1363]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B9219D51FFEDFA58D9FF3D12FEF86852" bold="true" box="[189,334,1337,1363]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFEDFA58D9FF3D41FE1F6AEA" blockId="30.[189,1399,1385,2027]" pageId="30" pageNumber="31">
Erect robust terrestrial herb
<quantity id="4CADECA6FFEDFA58DB4C3D42FD336885" box="[526,645,1385,1412]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="30" pageNumber="31" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">30–40 cm</quantity>
tall. Stem fleshy, repent at the base, rooting at the nodes, with simple white hairs, becoming glabrous with age. Stipules ovate-lanceolate, reddish,
<emphasis id="B9219D51FFEDFA58DDD73DA5FB1F68A6" box="[1173,1193,1422,1447]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
15 ×
<quantity id="4CADECA6FFEDFA58DDAF3DA5FA8268A9" box="[1261,1332,1422,1448]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="31" unit="mm" value="9.0">9 mm</quantity>
, apex acuminate, keel with sparse short hairs. Leaves: petiole
<quantity id="4CADECA6FFEDFA58DA093D9AFC7668CD" box="[843,960,1457,1484]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" metricValueMax="2.6" metricValueMin="1.6" pageId="30" pageNumber="31" unit="cm" value="21.0" valueMax="26.0" valueMin="16.0">16–26 cm</quantity>
long, terete, with dense white to pale pink hairs; lamina ovate, asymmetric, 17–23 ×
<quantity id="4CADECA6FFEDFA58DB9A3DFDFCF868F1" box="[728,846,1494,1520]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="30" pageNumber="31" unit="cm" value="16.0" valueMax="18.0" valueMin="14.0">14–18 cm</quantity>
, upper surface with short dense hairs, hairs deep pink when young becoming paler with age, basifixed, base cordate with lobes sometimes overlapping slightly, venation palmate-pinnate, margin sometimes slightly undulate, serrulate, teeth tipped by short glandular hairs, apex acute. Inflorescence axillary, usually borne near the base of the stem, total length
<emphasis id="B9219D51FFEDFA58D9FF3E43FF676B80" box="[189,209,1640,1665]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
<quantity id="4CADECA6FFEDFA58D9993E4DFEAD6B80" box="[219,283,1638,1665]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="cm" value="5.0">5 cm</quantity>
, a congested cyme, bisexual, with 10–25 flowers, male and female flowers open at the same time. Male flowers: pedicels with sparse minute hairs,
<quantity id="4CADECA6FFEDFA58DA763EA0FC006BA4" box="[820,950,1675,1701]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="30" pageNumber="31" unit="cm" value="3.2" valueMax="3.3" valueMin="3.1">3.1–3.3 cm</quantity>
long; tepals 4, subequal, elliptic, outer pair adaxially hairy, pink with a white margin, 1.3 ×
<quantity id="4CADECA6FFEDFA58DA743E84FC116BCB" box="[822,935,1711,1738]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="30" pageNumber="31" unit="cm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 cm</quantity>
; inner pair, glabrous, paler pink with a white margin, 1.2 ×
<quantity id="4CADECA6FFEDFA58D8F03EF8FE476BEF" box="[434,497,1747,1774]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="cm" value="1.0">1 cm</quantity>
; androecium yellow, symmetric, loosely globose; stamens 60–70, filaments unequal, shorter in the basal stamens, anthers club-shaped, dehiscing through slits
<emphasis id="B9219D51FFEDFA58DDC23ED3FB226A10" box="[1152,1172,1784,1809]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
¾ the length of the anther, slits lateral, connective extended. Female flowers: pedicels
<quantity id="4CADECA6FFEDFA58DA843F37FB836A37" box="[966,1077,1820,1846]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="30" pageNumber="31" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long; ovary red, triangularglobose, sparsely hairy to glabrous,
<emphasis id="B9219D51FFEDFA58DB233F6AFDC36A5B" box="[609,629,1857,1882]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
10 ×
<quantity id="4CADECA6FFEDFA58DBF53F6BFCBB6A5B" box="[695,781,1856,1882]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="mm" value="10.0">10 mm</quantity>
; capsule 3-locular, placentae bifid; tepals 5, obovateoblong, outer 3
<emphasis id="B9219D51FFEDFA58D83E3F4EFE266A7F" box="[380,400,1893,1918]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
15 ×
<quantity id="4CADECA6FFEDFA58D8983F4FFDF96A7E" box="[474,591,1892,1919]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="30" pageNumber="31" unit="mm" value="10.5" valueMax="13.0" valueMin="8.0">8–13 mm</quantity>
, pink with a white margin, sparsely adaxially hairy, inner 2
<emphasis id="B9219D51FFEDFA58DC603F4EFA806A7F" box="[1314,1334,1893,1918]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
15 ×
<quantity id="4CADECA6FFEDFA58D9FF3FA3FEBE6AA2" box="[189,264,1928,1955]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="mm" value="5.0">5 mm</quantity>
, white, glabrous; styles 3, greenish yellow, twice spirally twisted. Fruit
<quantity id="4CADECA6FFEDFA58DD3A3FA3FB526AA3" box="[1144,1252,1928,1955]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="30" pageNumber="31" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
number, on stiff pedicels
<emphasis id="B9219D51FFEDFA58D8193F86FED96AC7" box="[347,367,1965,1990]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
<quantity id="4CADECA6FFEDFA58D8353F86FE076AC6" box="[375,433,1965,1991]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="30" pageNumber="31" unit="cm" value="2.0">2 cm</quantity>
long, purple, berry-like,
<emphasis id="B9219D51FFEDFA58DB9B3F86FD5A6AC7" box="[729,748,1965,1990]" italics="true" pageId="30" pageNumber="31">c.</emphasis>
12 ×
<quantity id="4CADECA6FFEDFA58DA703F87FC3E6AC6" box="[818,904,1964,1991]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="30" pageNumber="31" unit="mm" value="15.0">15 mm</quantity>
, triangular-globose, fleshy, 3-ridged; apex truncate to rounded.
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFECFA59D9FF3920FB0F6C55" pageId="31" pageNumber="32" type="multiple">
<paragraph id="8BEA4143FFECFA59D9FF3920FE586C24" blockId="31.[189,494,267,293]" box="[189,494,267,293]" pageId="31" pageNumber="32">
<heading id="D0A2F62FFFECFA59D9FF3920FE586C24" bold="true" box="[189,494,267,293]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B9219D51FFECFA59D9FF3920FE586C24" bold="true" box="[189,494,267,293]" pageId="31" pageNumber="32">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFECFA59D9FF3911FB0F6C55" blockId="31.[189,1209,313,340]" box="[189,1209,313,340]" pageId="31" pageNumber="32">
In montane rainforest in
<collectingRegion id="49918FA1FFECFA59D89D3912FD3B6C55" box="[479,653,313,340]" country="Indonesia" name="Sumatera Utara" pageId="31" pageNumber="32">North Sumatra</collectingRegion>
and
<collectingRegion id="49918FA1FFECFA59DB863912FCB46C52" box="[708,770,313,339]" country="Indonesia" name="Aceh" pageId="31" pageNumber="32">Aceh</collectingRegion>
at altitudes of
<quantity id="4CADECA6FFECFA59DAF23911FBE46C55" box="[944,1106,314,340]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="31" pageNumber="32" unit="m" value="1500.0" valueMax="1700.0" valueMin="1300.0">1300–1700 m</quantity>
(
<figureCitation id="136E5DC6FFECFA59DD203912FB1C6C55" box="[1122,1194,313,340]" captionStart="Fig" captionStartId="8.[189,232,1965,1991]" captionTargetBox="[189,1398,586,1923]" captionTargetId="figure@8.[189,1398,585,1923]" captionTargetPageId="8" captionText="Fig. 2. Map of Sumatra showing the full known species distributions based on collection localities. Each point may represent more than one collection." figureDoi="http://doi.org/10.5281/zenodo.3805823" httpUri="https://zenodo.org/record/3805823/files/figure.png" pageId="31" pageNumber="32">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F12C8FFECFA59D9FF3950FC306F0B" pageId="31" pageNumber="32" type="conservation">
<paragraph id="8BEA4143FFECFA59D9FF3950FE036C94" blockId="31.[189,437,379,405]" box="[189,437,379,405]" pageId="31" pageNumber="32">
<heading id="D0A2F62FFFECFA59D9FF3950FE036C94" bold="true" box="[189,437,379,405]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B9219D51FFECFA59D9FF3950FE036C94" bold="true" box="[189,437,379,405]" pageId="31" pageNumber="32">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA4143FFECFA59D9FF3981FC306F0B" blockId="31.[189,1398,425,522]" pageId="31" pageNumber="32">
<taxonomicName id="4C553AC0FFECFA59D9FF3981FE0E6CC2" box="[189,440,425,451]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="pseudoscotti">
<emphasis id="B9219D51FFECFA59D9FF3981FE0E6CC2" box="[189,440,425,451]" italics="true" pageId="31" pageNumber="32">Begonia pseudoscotti</emphasis>
</taxonomicName>
does not appear to be common, but does have a fairly large distribution in northern
<collectingRegion id="49918FA1FFECFA59D9FF39E6FE956CE6" box="[189,291,461,487]" country="Indonesia" name="Sumatera Utara" pageId="31" pageNumber="32">Sumatra</collectingRegion>
. The preference for mid to high montane forests which are not under heavy threat means that currently an IUCN category of Least Concern is appropriate.
</paragraph>
</subSubSection>
</treatment>
</document>
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