treatments-xml/data/03/FC/97/03FC9766FFFBD67DFC25FEB5FEF9786C.xml

<document ID-DOI="10.1590/S1679-62252010000400005" ID-ISSN="1982-0224" ID-Zenodo-Dep="5422215" approvalRequired="87" approvalRequired_for_textStreams="77" approvalRequired_for_treatments="10" checkinTime="1630380595706" checkinUser="carolina" docAuthor="Bertaco, Vinicius A. &amp; Malabarba, Luiz R." docDate="2010" docId="03FC9766FFFBD67DFC25FEB5FEF9786C" docLanguage="en" docName="NeotropIchthyol.2010.63.pdf" docOrigin="Neotropical Ichthyology 8 (4)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252010000400005&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Hemibrycon mikrostiktos Bertaco &amp; Malabarba 2010, new species" docType="treatment" docVersion="2" lastPageId="19" masterDocId="FFC5EF1EFFEAD66EFFB2FFB0FFAA7C11" masterDocTitle="A review of the Cis-Andean species of Hemibrycon Günther (Teleostei: Characiformes: Characidae: Stevardiinae), with description of two new species" masterLastPageNumber="737" masterPageNumber="737" pageId="17" pageNumber="737" updateTime="1630688377022" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A review of the Cis-Andean species of Hemibrycon Günther (Teleostei: Characiformes: Characidae: Stevardiinae), with description of two new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bertaco, Vinicius A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Malabarba, Luiz R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
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<mods:start>737</mods:start>
<mods:end>737</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252010000400005</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">5422215</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03FC9766FFFBD67DFC25FEB5FEF9786C" httpUri="http://treatment.plazi.org/id/03FC9766FFFBD67DFC25FEB5FEF9786C" lastPageId="19" pageId="17">
<subSubSection box="[919,1340,261,285]" pageId="17" type="nomenclature">
<paragraph blockId="17.[919,1340,261,316]" box="[919,1340,261,285]" pageId="17">
<heading box="[919,1340,261,285]" centered="true" fontSize="10" level="2" pageId="17" reason="2">
<emphasis bold="true" box="[919,1340,261,285]" pageId="17">
<taxonomicName authority="Bertaco &amp; Malabarba, 2010" authorityName="Bertaco &amp; Malabarba" authorityYear="2010" box="[919,1195,261,285]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="mikrostiktos" status="new species">
<emphasis bold="true" box="[919,1195,261,285]" italics="true" pageId="17">Hemibrycon mikrostiktos</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1209,1340,261,284]" pageId="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1067,1193,293,316]" pageId="17" type="description">
<paragraph blockId="17.[919,1340,261,316]" box="[1067,1193,293,316]" pageId="17">
<figureCitation box="[1067,1193,293,316]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[136,175,834,857]" captionStartId-1="18.[816,855,1956,1979]" captionTargetBox-0="[136,1459,257,824]" captionTargetBox-1="[816,1458,1432,1942]" captionTargetId-0="figure-468@18.[136,1459,257,824]" captionTargetId-1="figure-470@18.[816,1458,1432,1942]" captionTargetPageId-0="18" captionTargetPageId-1="18" captionText-0="Fig. 16. Hemibrycon mikrostiktos, MUSM 35490, holotype, 44.4 mm SL, upper río Ucayali basin, Peru." captionText-1="Fig. 17. Hemibrycon mikrostiktos, MCP 44521, paratype, female, 35.7 mm SL. Scanning electron micrograph of left side upper and lower jaws. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5422245" figureDoi-1="http://doi.org/10.5281/zenodo.5422247" httpUri-0="https://zenodo.org/record/5422245/files/figure.png" httpUri-1="https://zenodo.org/record/5422247/files/figure.png" pageId="17">
<emphasis bold="true" box="[1067,1193,293,316]" pageId="17">Figs. 16-17</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="17" type="materials_examined">
<paragraph blockId="17.[809,1451,362,441]" pageId="17">
<materialsCitation collectingDate="1999-11-02" collectionCode="MUSM" collectorName="P. de Rham &amp; F. Chang." country="Peru" county="Padre Abad" latitude="-9.042778" location="rio Aguaytia" longLatPrecision="21" longitude="-75.5125" municipality="Aguaytia" pageId="17" specimenCode="MUSM 35490" specimenCount="1" stateProvince="Ucayali" typeStatus="holotype">
<emphasis bold="true" box="[809,909,362,383]" pageId="17">
<typeStatus box="[809,905,362,383]" pageId="17">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[917,1062,362,383]" collectionCode="MUSM" pageId="17">MUSM 35490</specimenCode>
, 
<quantity box="[1074,1162,362,383]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="17" unit="mm" value="44.4">44.4 mm</quantity>
SL, 
<location LSID="urn:lsid:plazi:treatment:03FC9766FFFBD67DFC25FEB5FEF9786C:8E8A70ABFFFBD67FFB0EFEDAFA907D6E" box="[1212,1338,362,383]" country="Peru" county="Padre Abad" latitude="-9.042778" longLatPrecision="21" longitude="-75.5125" municipality="Aguaytia" name="rio Aguaytia" pageId="17" stateProvince="Ucayali">río Aguaytía</location>
, 
<location LSID="urn:lsid:plazi:treatment:03FC9766FFFBD67DFC25FEB5FEF9786C:8E8A70ABFFFBD67FFAF7FEDAFA0D7D6E" box="[1349,1447,362,383]" country="Peru" county="Padre Abad" latitude="-9.042778" longLatPrecision="21" longitude="-75.5125" municipality="Aguaytia" name="rio Negro" pageId="17" stateProvince="Ucayali">río Negro</location>
, upper 
<collectingRegion box="[874,989,391,412]" country="Peru" name="Ucayali" pageId="17">río Ucayali</collectingRegion>
basin, 
<collectingMunicipality box="[1065,1158,391,412]" pageId="17">Aguaytía</collectingMunicipality>
, 
<collectingCounty box="[1172,1293,391,412]" pageId="17">Padre Abad</collectingCounty>
, 
<collectingRegion box="[1306,1386,391,412]" country="Peru" name="Ucayali" pageId="17">Ucayali</collectingRegion>
, 
<collectingCountry box="[1399,1447,391,412]" name="Peru" pageId="17">Peru</collectingCountry>
, 
<geoCoordinate box="[809,922,420,441]" degrees="09" direction="south" minutes="02" orientation="latitude" pageId="17" precision="15" seconds="34" value="-9.042778">09º02’34”S</geoCoordinate>
<geoCoordinate box="[927,1049,420,441]" degrees="75" direction="west" minutes="30" orientation="longitude" pageId="17" precision="15" seconds="45" value="-75.5125">75º30’45”W</geoCoordinate>
, 
<date box="[1059,1174,420,441]" pageId="17" value="1999-11-02">
<collectingDate box="[1059,1174,420,441]" pageId="17" value="1999-11-02">2 Nov 1999</collectingDate>
</date>
, 
<collectorName box="[1184,1292,420,441]" pageId="17">P. de Rham</collectorName>
&amp; 
<collectorName box="[1322,1415,420,441]" pageId="17">F. Chang.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="17.[809,1451,486,564]" pageId="17">
<materialsCitation accessionNumber="MCP 44521" box="[809,1451,486,507]" collectionCode="MCP, MUSM" pageId="17" specimenCode="10" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[809,919,486,507]" pageId="17">
<typeStatus box="[809,915,486,507]" pageId="17">Paratypes</typeStatus>
.
</emphasis>
<accessionNumber box="[927,1050,486,507]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP44521" pageId="17">
<collectionCode box="[927,978,486,507]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="17">MCP</collectionCode>
44521
</accessionNumber>
, 
<specimenCode box="[1062,1086,486,507]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="17">10</specimenCode>
(3 c&amp;s), 
<quantity box="[1185,1325,486,507]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.24" metricValueMin="2.76" pageId="17" unit="mm" value="35.0" valueMax="42.4" valueMin="27.6">27.6-42.4 mm</quantity>
SL; 
<collectionCode box="[1377,1451,486,507]" pageId="17">MUSM</collectionCode>
</materialsCitation>
<materialsCitation box="[809,1450,515,536]" collectionCode="UFRGS" pageId="17" specimenCode="19" specimenCount="1" typeStatus="paratype">
15845, 
<specimenCode box="[879,903,515,536]" collectionCode="MUSM" pageId="17">19</specimenCode>
measured and counted of 187, 
<quantity box="[1196,1329,515,536]" metricMagnitude="-2" metricUnit="m" metricValue="3.345" metricValueMax="4.24" metricValueMin="2.45" pageId="17" unit="mm" value="33.45" valueMax="42.4" valueMin="24.5">24.5-42.4 mm</quantity>
SL; 
<collectionCode box="[1373,1450,515,536]" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="17">UFRGS</collectionCode>
</materialsCitation>
<materialsCitation box="[809,1372,543,564]" collectionCode="UFRGS" pageId="17" specimenCode="10, 30.9" specimenCount="1" typeStatus="paratype">
12003, 
<specimenCode box="[882,908,543,564]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="17">10</specimenCode>
, 
<specimenCode box="[918,961,543,564]" collectionCode="UFRGS" country="Brazil" httpUri="http://grbio.org/cool/3q02-t412" name="Universidade Federale do Rio Grande do Sul" pageId="17">30.9</specimenCode>
-36.0 mm SL, collected with the 
<typeStatus box="[1283,1372,543,564]" pageId="17">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="17" type="diagnosis">
<paragraph blockId="17.[809,1451,607,886]" pageId="17">
<emphasis bold="true" box="[809,926,607,630]" pageId="17">Diagnosis.</emphasis>
<taxonomicName authorityName="Bertaco &amp; Malabarba" authorityYear="2010" box="[931,1199,607,630]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[931,1199,607,630]" italics="true" pageId="17">Hemibrycon mikrostiktos</emphasis>
</taxonomicName>
is readily distinguished from all congeners by the size of the humeral spot (2-3 
<emphasis box="[1382,1404,639,662]" italics="true" pageId="17">vs</emphasis>
. 4-9 horizontal series of scales), number of scale rows below of the lateral line (3-4 
<emphasis box="[1008,1030,703,726]" italics="true" pageId="17">vs</emphasis>
. 4-8), and total number of pelvic-fin rays (7 
<emphasis box="[841,863,735,758]" italics="true" pageId="17">vs</emphasis>
. 8). Furthermore, 
<taxonomicName authorityName="Bertaco &amp; Malabarba" authorityYear="2010" box="[1074,1244,735,758]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[1074,1093,735,758]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[1111,1244,735,758]" italics="true" pageId="17">mikrostiktos</emphasis>
</taxonomicName>
differs from most congeners by the number of branched anal-fin rays (18-21 
<emphasis box="[809,832,799,822]" italics="true" pageId="17">vs</emphasis>
. 21-34, and 
<quantity box="[970,1025,799,822]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="17" unit="in" value="17.0">17 in</quantity>
<taxonomicName authorityName="Eigenmann" authorityYear="1922" box="[1032,1143,799,822]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="tridens">
<emphasis box="[1032,1051,799,822]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[1065,1143,799,822]" italics="true" pageId="17">tridens</emphasis>
</taxonomicName>
, 
<figureCitation box="[1157,1224,799,822]" captionStart="Fig" captionStartId="3.[129,168,1924,1947]" captionTargetBox="[133,1445,1416,1903]" captionTargetPageId="3" captionText="Fig. 2. Tukey box plots of number of branched anal-fin rays in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." figureDoi="http://doi.org/10.5281/zenodo.5422219" httpUri="https://zenodo.org/record/5422219/files/figure.png" pageId="17">Fig. 2</figureCitation>
), except 
<taxonomicName authorityName=", Sep" authorityYear="1921" box="[1324,1405,799,822]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="beni">
<emphasis box="[1324,1343,799,822]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[1357,1405,799,822]" italics="true" pageId="17">beni</emphasis>
</taxonomicName>
and 
<taxonomicName box="[809,920,831,854]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="helleri">
<emphasis box="[809,828,831,854]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[845,920,831,854]" italics="true" pageId="17">helleri</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName authorityName=", Sep" authorityYear="1921" box="[1113,1198,831,854]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="beni">
<emphasis box="[1113,1132,831,854]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[1149,1198,831,854]" italics="true" pageId="17">beni</emphasis>
</taxonomicName>
and 
<taxonomicName box="[1258,1369,831,854]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="helleri">
<emphasis box="[1258,1277,831,854]" italics="true" pageId="17">H</emphasis>
. 
<emphasis box="[1294,1369,831,854]" italics="true" pageId="17">helleri</emphasis>
</taxonomicName>
by the number of lateral line scales (40-42 
<emphasis box="[1168,1189,863,886]" italics="true" pageId="17">vs</emphasis>
. 44-53 and 42-44, 
<figureCitation box="[1378,1437,863,886]" captionStart="Fig" captionStartId="3.[809,848,1924,1947]" captionText="Fig. 3. Tukey box plots of number of lateral line scales in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." pageId="17">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" pageId="17" type="description">
<paragraph blockId="17.[809,1451,934,2044]" pageId="17">
<emphasis bold="true" box="[809,944,934,957]" pageId="17">Description.</emphasis>
Morphometric data for 
<taxonomicName authorityName="Bertaco &amp; Malabarba" authorityYear="2010" box="[1189,1451,934,957]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="17" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[1189,1451,934,957]" italics="true" pageId="17">Hemibrycon mikrostiktos</emphasis>
</taxonomicName>
summarized in 
<tableCitation box="[970,1050,965,988]" captionStart="Table 4" captionStartId="19.[131,192,260,283]" captionText="Table 4. Morphometric data of holotype (H) of Hemibrycon metae (CAS 123727), and non-types from río Meta basin, Colombia (IAvHP 2973,3122,3322,3628,3632), andfrom ríoApurebasin,Venezuela(INHS27766,31851,61270;MCNG 98,5646,6759,7923; NRM23991, 23993), all from ríoOrinoco basin;and holotype(MUSM 35490) and paratypes (MCP44521, MUSM 15845, UFRGS 12003, all unsexed)ofHemibrycon mikrostiktos from the upperrío Ucayali basin,Peru.m, males;f, females.SD =Standard deviation." pageId="17">Table 4</tableCitation>
. Largest specimen 
<quantity box="[1251,1345,965,988]" metricMagnitude="-2" metricUnit="m" metricValue="4.4399999999999995" pageId="17" unit="mm" value="44.4">44.4 mm</quantity>
SL. Body compressed and moderately elongate; greatest body depth at vertical through middle length of pectoral fins. Dorsal profile of head nearly straight. Dorsal body profile convex from occipital bony to dorsal-fin origin; posteroventrally slanted at dorsal-fin base; straight from last dorsal-fin ray to adiposefin origin. Ventral profile of head convex. Ventral body profile convex from pectoral-fin origin to pelvic-fin origin, and straight to slightly convex to anal-fin origin. Body profile along analfin base posterodorsally slanted. Caudal peduncle short, nearly straight to slightly concave along dorsal and ventral margins.
</paragraph>
<paragraph blockId="17.[809,1451,934,2044]" pageId="17">
Snout rounded from margin of upper lip to vertical through anterior nostrils. Head relatively small. Mouth terminal, mouth slit nearly at horizontal through below middle of eye. Maxilla long and slightly curved, aligned at angle of approximately 45 
<superScript attach="none" box="[1443,1451,1444,1458]" fontSize="6" pageId="17">o</superScript>
to longitudinal body axis, and its posterior tip extending below orbit, reaching nearly to vertical through center of eye.
</paragraph>
<paragraph blockId="17.[809,1451,934,2044]" pageId="17">
Premaxilla with two tooth rows; outer row with 4-6, tricuspid teeth with central cusp slightly longer; inner row with 4 pentacuspid teeth, gradually decreasing in length from first to third teeth and last tooth smallest; central cusp twice or three times longer and broader than other cusps. Maxilla toothed with 7-11 tri- to pentacuspid teeth, except for last three or four conical teeth, with central cusp longer. Three anteriormost dentary teeth larger, with 5 cusps, followed by medium sized tooth with 3 cusps, and 12-13 teeth with 1-3 cusps or conical; central cusp in all teeth two to three times longer and broader than other cusps. Cusp tips slightly curved posteriorly and lingually (
<figureCitation box="[1161,1240,1893,1916]" captionStart="Fig" captionStartId="18.[816,855,1956,1979]" captionTargetBox="[816,1458,1432,1942]" captionTargetId="figure-470@18.[816,1458,1432,1942]" captionTargetPageId="18" captionText="Fig. 17. Hemibrycon mikrostiktos, MCP 44521, paratype, female, 35.7 mm SL. Scanning electron micrograph of left side upper and lower jaws. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5422247" httpUri="https://zenodo.org/record/5422247/files/figure.png" pageId="17">Fig. 17</figureCitation>
).
</paragraph>
<paragraph blockId="17.[809,1451,934,2044]" pageId="17">Dorsal-fin rays ii,8 (n = 20); first unbranched ray approximately one-half length of second ray. Dorsal-fin origin located posterior to middle of SL and posterior to vertical through pelvic-fin origin. First dorsal-fin pterygiophore</paragraph>
<paragraph blockId="18.[634,1458,181,205]" box="[634,1458,181,205]" pageId="18">
<emphasis box="[634,952,181,202]" italics="true" pageId="18">V. A. Bertaco &amp; L. R. Malabarba</emphasis>
755
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5422245" ID-Zenodo-Dep="5422245" box="[136,1232,834,857]" httpUri="https://zenodo.org/record/5422245/files/figure.png" pageId="18" startId="18.[136,175,834,857]" targetBox="[136,1459,257,824]" targetPageId="18">
<paragraph blockId="18.[136,1232,834,857]" box="[136,1232,834,857]" pageId="18">
<emphasis bold="true" box="[136,218,834,857]" pageId="18">Fig. 16.</emphasis>
<taxonomicName authority=", MUSM" authorityName="MUSM" box="[224,587,834,857]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="18" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[224,492,834,857]" italics="true" pageId="18">Hemibrycon mikrostiktos</emphasis>
, MUSM
</taxonomicName>
35490, holotype, 44.4 mm SL, upper río Ucayali basin, Peru.
</paragraph>
</caption>
<paragraph blockId="18.[136,779,900,1819]" pageId="18">
inserted between the neural spines of eleventh to twelfth vertebrae (n = 3). Profile of distal margin of dorsal fin convex. Adipose-fin located at vertical through insertion of last or posterior to last anal-fin ray. Anal-fin rays iii-iv,18-20 (two with 21, mode = 19, n = 20, 
<figureCitation box="[438,503,1028,1051]" captionStart="Fig" captionStartId="3.[129,168,1924,1947]" captionTargetBox="[133,1445,1416,1903]" captionTargetPageId="3" captionText="Fig. 2. Tukey box plots of number of branched anal-fin rays in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." figureDoi="http://doi.org/10.5281/zenodo.5422219" httpUri="https://zenodo.org/record/5422219/files/figure.png" pageId="18">Fig. 2</figureCitation>
). Anal-fin profile slightly concave in all specimens. First anal-fin pterygiophore inserted between the haemal spine of last precaudal vertebra and the first caudal vertebrae.Anal-fin origin approximately at vertical through insertion in the middle dorsal fin. Pectoral-fin rays i,9-10 (mode = 9, n = 20). Pectoral-fin tip reaching pelvic-fin origin. Pelvic-fin rays i,5,i (n = 15) or i,6 (n = 5). Pelvic-fin origin located 3-4 predorsal scales anterior to vertical through dorsal-fin origin. Caudal fin forked with 19 principal rays (n = 20); lobes similar in size. Basal portion of caudal-fin lobes covered with irregular scales and smaller than those of the body, following by one larger scale in each lobe. Dorsal procurrent rays 12-13 and ventral procurrent rays 12-14 (n = 3).
</paragraph>
<paragraph blockId="18.[136,779,900,1819]" pageId="18">
Scales cycloid, moderately large. Lateral line complete with 40-42 (mode = 41, n = 16, 
<figureCitation box="[407,468,1476,1499]" captionStart="Fig" captionStartId="3.[809,848,1924,1947]" captionText="Fig. 3. Tukey box plots of number of lateral line scales in Hemibrycon species. Mean represented by thick vertical bar, and 25th and 75th percetiles as lateral borders of box plots. Number in parentheses = N." pageId="18">Fig. 3</figureCitation>
).[Scale rows between dorsalfin origin and lateral line 6-7 (mode = 6, n = 18); scale rows between lateral line and pelvic-fin origin 3-4 (mode = 4, n = 18). Predorsal scales 14, arranged in regular series (n = 18). Scales rows around caudal peduncle 14 (n = 18). Axillary scale on pelvic-fin origin extends posteriorly covering 2-3 scales. Scale sheath along anal-fin base with 7-10 scales in single series, extending to base of most anterior branched rays.
</paragraph>
<paragraph blockId="18.[136,779,900,1819]" pageId="18">Precaudal vertebrae 17; caudal vertebrae 23-24; total vertebrae 40-41. Supraneurals 6-7 (n = 3). Gill rakers on upper limb of outer gill arch 5-6, and on lower limb 9-10 (n = 5).</paragraph>
<paragraph blockId="18.[136,778,1860,2043]" lastBlockId="18.[816,1459,900,1147]" pageId="18">
<emphasis bold="true" box="[136,321,1860,1883]" pageId="18">Color in alcohol.</emphasis>
Dorsal portion of head and body with dense concentration of black chromatophores. Dorsal and dorsolateral portion of caudal peduncle near caudal fin base densely pigmented. Dorsolateral portion of body with scattered black chromatophores. Snout and upper portion of maxilla densely pigmented. Infraorbitals and opercle with scattered black chromatophores. Midlateral body silvery and dark pigmented. One small black humeral spot, located over third to fourth lateral line scales and extending over 1-2 horizontal series of scales, sometimes including lateral line. Base of caudal fin and middle rays black pigmented. Abdominal region almost devoid of black chromatophores. Dorsal, adipose, anal and caudal fin with dark pigmentation diffuse. Pectoral and pelvic fins hyaline (
<figureCitation box="[1247,1325,1124,1147]" captionStart="Fig" captionStartId="18.[136,175,834,857]" captionTargetBox="[136,1459,257,824]" captionTargetId="figure-468@18.[136,1459,257,824]" captionTargetPageId="18" captionText="Fig. 16. Hemibrycon mikrostiktos, MUSM 35490, holotype, 44.4 mm SL, upper río Ucayali basin, Peru." figureDoi="http://doi.org/10.5281/zenodo.5422245" httpUri="https://zenodo.org/record/5422245/files/figure.png" pageId="18">Fig. 16</figureCitation>
).
</paragraph>
<paragraph blockId="18.[816,1459,1188,1275]" pageId="18">
<emphasis bold="true" box="[816,1046,1188,1211]" pageId="18">Sexual dimorphism.</emphasis>
None of the 
<typeStatus box="[1183,1229,1188,1211]" pageId="18">type</typeStatus>
specimens examined had hooks on fins or any other apparent sexually dimorphic features. Gill glands were not found on first gill arch.
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" pageId="18" type="distribution">
<paragraph blockId="18.[816,1458,1316,1371]" pageId="18">
<emphasis bold="true" box="[816,964,1316,1339]" pageId="18">Distribution.</emphasis>
<taxonomicName box="[973,1248,1316,1339]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="18" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[973,1248,1316,1339]" italics="true" pageId="18">Hemibrycon mikrostiktos</emphasis>
</taxonomicName>
is known from río Aguaytía, upper 
<collectingRegion box="[992,1110,1348,1371]" country="Peru" name="Ucayali" pageId="18">río Ucayali</collectingRegion>
drainage, 
<collectingRegion box="[1219,1305,1348,1371]" country="Peru" name="Ucayali" pageId="18">Ucayali</collectingRegion>
, 
<collectingCountry box="[1314,1363,1348,1371]" name="Peru" pageId="18">Peru</collectingCountry>
(
<figureCitation box="[1376,1441,1348,1371]" captionStart="Fig" captionStartId="4.[136,175,1739,1762]" captionTargetBox="[145,760,1305,1720]" captionTargetId="figure-936@4.[136,778,1291,1720]" captionTargetPageId="4" captionText="Fig. 4. Map of central and northern South America showing geographic distribution of Hemibrycon beni (1, solid hexagon), H. divisorensis (2, open triangle), H. helleri (3, ellipse), H. huambonicus (4, solid triangles), H. jelskii (5, solid dots), H. inambari (6, rectangles), H. metae (7, solid squares), H. mikrostiktos (8, open square), H. polyodon (9, inverted triangle), H. surinamensis (10, solid diamonds), H. taeniurus (11,Trinidad Island), H.tridens (12, open circle).Some symbols represent more than one lot or locality of specimens. 1-12 = type locality of each species." figureDoi="http://doi.org/10.5281/zenodo.5422221" httpUri="https://zenodo.org/record/5422221/files/figure.png" pageId="18">Fig. 4</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5422247" ID-Zenodo-Dep="5422247" httpUri="https://zenodo.org/record/5422247/files/figure.png" pageId="18" startId="18.[816,855,1956,1979]" targetBox="[816,1458,1432,1942]" targetPageId="18">
<paragraph blockId="18.[816,1458,1956,2043]" pageId="18">
<emphasis bold="true" box="[816,904,1956,1979]" pageId="18">Fig. 17.</emphasis>
<taxonomicName authority=", MCP" authorityName="MCP" box="[914,1264,1956,1979]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="18" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[914,1190,1956,1979]" italics="true" pageId="18">Hemibrycon mikrostiktos</emphasis>
, MCP
</taxonomicName>
44521, paratype, female, 35.7 mm SL. Scanning electron micrograph of left side upper and lower jaws. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph blockId="19.[129,1045,180,205]" box="[129,1045,180,205]" pageId="19">
756 A review of the Cis-Andean species of 
<taxonomicName baseAuthorityName="Malabarba &amp; Weitzman" baseAuthorityYear="2003" box="[925,1045,180,201]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="genus">
<emphasis box="[925,1045,180,201]" italics="true" pageId="19">Hemibrycon</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<caption pageId="19" startId="19.[131,192,260,283]">
<paragraph blockId="19.[131,1454,260,379]" pageId="19">
<emphasis bold="true" box="[131,217,260,283]" pageId="19">Table 4.</emphasis>
Morphometric data of holotype (H) of 
<taxonomicName box="[619,815,260,283]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="species" species="metae">
<emphasis box="[619,815,260,283]" italics="true" pageId="19">Hemibrycon metae</emphasis>
</taxonomicName>
(CAS 123727), and non-types from río Meta basin, Colombia (IAvHP 2973, 3122, 3322, 3628, 3632), and from ríoApure basin,Venezuela (INHS 27766, 31851, 61270; MCNG 98, 5646, 6759, 7923; NRM 23991, 23993), all from río Orinoco basin; and holotype (MUSM 35490) and paratypes (MCP 44521, MUSM 15845, UFRGS 12003, all unsexed) of 
<taxonomicName box="[354,607,356,379]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[354,607,356,379]" italics="true" pageId="19">Hemibrycon mikrostiktos</emphasis>
</taxonomicName>
from the upper río Ucayali basin, Peru. m, males; f, females. SD = Standard deviation.
</paragraph>
</caption>
<paragraph pageId="19">
<table box="[134,1444,395,1006]" gridcols="15" gridrows="25" pageId="19">
<tr box="[134,1444,395,414]" gridrow="0" pageId="19" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-14="1" rowspan-2="1" rowspan-3="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<td box="[718,857,395,414]" colspan="3" colspanRight="2" gridcol="4" gridrow="0" pageId="19">
<taxonomicName box="[758,835,395,414]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="species" species="metae">
<emphasis box="[758,773,395,414]" italics="true" pageId="19">H</emphasis>
. 
<emphasis box="[784,835,395,414]" italics="true" pageId="19">metae</emphasis>
</taxonomicName>
</td>
<td box="[1217,1393,395,414]" colspan="2" colspanRight="1" gridcol="12" gridrow="0" pageId="19">
<taxonomicName box="[1217,1347,395,414]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[1217,1232,395,414]" italics="true" pageId="19">H</emphasis>
. 
<emphasis box="[1243,1347,395,414]" italics="true" pageId="19">mikrostiktos</emphasis>
</taxonomicName>
</td>
</tr>
<tr box="[134,1444,421,440]" gridrow="1" pageId="19" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-14="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
<td box="[603,767,421,440]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="19">Río Meta basin</td>
<td box="[886,1050,421,440]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="19">Río Apure basin</td>
<td box="[1180,1393,421,440]" colspan="3" colspanRight="2" gridcol="11" gridrow="1" pageId="19">Río Ucayali basin</td>
</tr>
<tr box="[134,1444,446,465]" gridrow="2" pageId="19" rowspan-0="1">
<td box="[496,533,446,465]" gridcol="1" gridrow="2" pageId="19">H</td>
<td box="[553,575,446,465]" gridcol="2" gridrow="2" pageId="19">n</td>
<td box="[603,685,446,465]" gridcol="3" gridrow="2" pageId="19">Range</td>
<td box="[718,767,446,465]" gridcol="4" gridrow="2" pageId="19">Mean</td>
<td box="[790,818,446,465]" gridcol="5" gridrow="2" pageId="19">SD</td>
<td box="[836,857,446,465]" gridcol="6" gridrow="2" pageId="19">n</td>
<td box="[886,968,446,465]" gridcol="7" gridrow="2" pageId="19">Range</td>
<td box="[1001,1050,446,465]" gridcol="8" gridrow="2" pageId="19">Mean</td>
<td box="[1073,1101,446,465]" gridcol="9" gridrow="2" pageId="19">SD</td>
<td box="[1122,1159,446,465]" gridcol="10" gridrow="2" pageId="19">H</td>
<td box="[1180,1201,446,465]" gridcol="11" gridrow="2" pageId="19">n</td>
<td box="[1217,1311,446,465]" gridcol="12" gridrow="2" pageId="19">Range</td>
<td box="[1344,1393,446,465]" gridcol="13" gridrow="2" pageId="19">Mean</td>
<td box="[1417,1444,446,465]" gridcol="14" gridrow="2" pageId="19">SD</td>
</tr>
<tr box="[134,1444,472,491]" gridrow="3" pageId="19">
<td box="[134,388,472,491]" gridcol="0" gridrow="3" pageId="19">Standard length (mm)</td>
<td box="[496,533,472,491]" gridcol="1" gridrow="3" pageId="19">77.2</td>
<td box="[553,575,472,491]" gridcol="2" gridrow="3" pageId="19">43</td>
<td box="[603,685,472,491]" gridcol="3" gridrow="3" pageId="19">51.3-93.1</td>
<td box="[718,767,472,491]" gridcol="4" gridrow="3" pageId="19">67.5</td>
<td box="[790,818,472,491]" gridcol="5" gridrow="3" pageId="19">-</td>
<td box="[836,857,472,491]" gridcol="6" gridrow="3" pageId="19">97</td>
<td box="[886,968,472,491]" gridcol="7" gridrow="3" pageId="19">32.4-76.7</td>
<td box="[1001,1050,472,491]" gridcol="8" gridrow="3" pageId="19">44.5</td>
<td box="[1073,1101,472,491]" gridcol="9" gridrow="3" pageId="19">-</td>
<td box="[1122,1159,472,491]" gridcol="10" gridrow="3" pageId="19">44.4</td>
<td box="[1180,1201,472,491]" gridcol="11" gridrow="3" pageId="19">12</td>
<td box="[1217,1311,472,491]" gridcol="12" gridrow="3" pageId="19">32.7-44.4</td>
<td box="[1344,1393,472,491]" gridcol="13" gridrow="3" pageId="19">36.8</td>
<td box="[1417,1444,472,491]" gridcol="14" gridrow="3" pageId="19">-</td>
</tr>
<tr box="[134,1444,497,516]" gridrow="4" pageId="19" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-8="1" rowspan-9="1">
<td box="[603,968,497,516]" colspan="5" colspanRight="4" gridcol="3" gridrow="4" pageId="19">Percents of Standard length</td>
</tr>
<tr box="[134,1444,521,540]" gridrow="5" pageId="19">
<td box="[134,388,521,540]" gridcol="0" gridrow="5" pageId="19">Predorsal distance</td>
<td box="[496,533,521,540]" gridcol="1" gridrow="5" pageId="19">50.8</td>
<td box="[553,575,521,540]" gridcol="2" gridrow="5" pageId="19">43</td>
<td box="[603,685,521,540]" gridcol="3" gridrow="5" pageId="19">47.4-53.3</td>
<td box="[718,767,521,540]" gridcol="4" gridrow="5" pageId="19">50.6</td>
<td box="[790,818,521,540]" gridcol="5" gridrow="5" pageId="19">1.4</td>
<td box="[836,857,521,540]" gridcol="6" gridrow="5" pageId="19">97</td>
<td box="[886,968,521,540]" gridcol="7" gridrow="5" pageId="19">48.4-56.2</td>
<td box="[1001,1050,521,540]" gridcol="8" gridrow="5" pageId="19">52.5</td>
<td box="[1073,1101,521,540]" gridcol="9" gridrow="5" pageId="19">1.4</td>
<td box="[1122,1159,521,540]" gridcol="10" gridrow="5" pageId="19">52.8</td>
<td box="[1180,1201,521,540]" gridcol="11" gridrow="5" pageId="19">12</td>
<td box="[1217,1311,521,540]" gridcol="12" gridrow="5" pageId="19">52.1-55.2</td>
<td box="[1344,1393,521,540]" gridcol="13" gridrow="5" pageId="19">53.6</td>
<td box="[1417,1444,521,540]" gridcol="14" gridrow="5" pageId="19">1.0</td>
</tr>
<tr box="[134,1444,546,565]" gridrow="6" pageId="19">
<td box="[134,388,546,565]" gridcol="0" gridrow="6" pageId="19">Prepelvic distance</td>
<td box="[496,533,546,565]" gridcol="1" gridrow="6" pageId="19">42.2</td>
<td box="[553,575,546,565]" gridcol="2" gridrow="6" pageId="19">43</td>
<td box="[603,685,546,565]" gridcol="3" gridrow="6" pageId="19">39.8-45.3</td>
<td box="[718,767,546,565]" gridcol="4" gridrow="6" pageId="19">42.0</td>
<td box="[790,818,546,565]" gridcol="5" gridrow="6" pageId="19">1.3</td>
<td box="[836,857,546,565]" gridcol="6" gridrow="6" pageId="19">97</td>
<td box="[886,968,546,565]" gridcol="7" gridrow="6" pageId="19">37.9-49.3</td>
<td box="[1001,1050,546,565]" gridcol="8" gridrow="6" pageId="19">43.9</td>
<td box="[1073,1101,546,565]" gridcol="9" gridrow="6" pageId="19">1.5</td>
<td box="[1122,1159,546,565]" gridcol="10" gridrow="6" pageId="19">43.9</td>
<td box="[1180,1201,546,565]" gridcol="11" gridrow="6" pageId="19">12</td>
<td box="[1217,1311,546,565]" gridcol="12" gridrow="6" pageId="19">43.6-45.7</td>
<td box="[1344,1393,546,565]" gridcol="13" gridrow="6" pageId="19">44.8</td>
<td box="[1417,1444,546,565]" gridcol="14" gridrow="6" pageId="19">0.7</td>
</tr>
<tr box="[134,1444,570,589]" gridrow="7" pageId="19">
<td box="[134,388,570,589]" gridcol="0" gridrow="7" pageId="19">Prepectoral distance</td>
<td box="[496,533,570,589]" gridcol="1" gridrow="7" pageId="19">22.5</td>
<td box="[553,575,570,589]" gridcol="2" gridrow="7" pageId="19">43</td>
<td box="[603,685,570,589]" gridcol="3" gridrow="7" pageId="19">21.0-24.6</td>
<td box="[718,767,570,589]" gridcol="4" gridrow="7" pageId="19">22.7</td>
<td box="[790,818,570,589]" gridcol="5" gridrow="7" pageId="19">0.7</td>
<td box="[836,857,570,589]" gridcol="6" gridrow="7" pageId="19">97</td>
<td box="[886,968,570,589]" gridcol="7" gridrow="7" pageId="19">21.7-26.2</td>
<td box="[1001,1050,570,589]" gridcol="8" gridrow="7" pageId="19">23.6</td>
<td box="[1073,1101,570,589]" gridcol="9" gridrow="7" pageId="19">0.9</td>
<td box="[1122,1159,570,589]" gridcol="10" gridrow="7" pageId="19">23.1</td>
<td box="[1180,1201,570,589]" gridcol="11" gridrow="7" pageId="19">12</td>
<td box="[1217,1311,570,589]" gridcol="12" gridrow="7" pageId="19">23.1-25.2</td>
<td box="[1344,1393,570,589]" gridcol="13" gridrow="7" pageId="19">24.1</td>
<td box="[1417,1444,570,589]" gridcol="14" gridrow="7" pageId="19">0.6</td>
</tr>
<tr box="[134,1444,595,614]" gridrow="8" pageId="19">
<td box="[134,388,595,614]" gridcol="0" gridrow="8" pageId="19">Preanal distance</td>
<td box="[496,533,595,614]" gridcol="1" gridrow="8" pageId="19">58.4</td>
<td box="[553,575,595,614]" gridcol="2" gridrow="8" pageId="19">43</td>
<td box="[603,685,595,614]" gridcol="3" gridrow="8" pageId="19">53.3-60.7</td>
<td box="[718,767,595,614]" gridcol="4" gridrow="8" pageId="19">57.1</td>
<td box="[790,818,595,614]" gridcol="5" gridrow="8" pageId="19">1.9</td>
<td box="[836,857,595,614]" gridcol="6" gridrow="8" pageId="19">97</td>
<td box="[886,968,595,614]" gridcol="7" gridrow="8" pageId="19">54.7-62.1</td>
<td box="[1001,1050,595,614]" gridcol="8" gridrow="8" pageId="19">58.1</td>
<td box="[1073,1101,595,614]" gridcol="9" gridrow="8" pageId="19">1.6</td>
<td box="[1122,1159,595,614]" gridcol="10" gridrow="8" pageId="19">58.9</td>
<td box="[1180,1201,595,614]" gridcol="11" gridrow="8" pageId="19">12</td>
<td box="[1217,1311,595,614]" gridcol="12" gridrow="8" pageId="19">56.5-60.5</td>
<td box="[1344,1393,595,614]" gridcol="13" gridrow="8" pageId="19">59.1</td>
<td box="[1417,1444,595,614]" gridcol="14" gridrow="8" pageId="19">1.1</td>
</tr>
<tr box="[134,1444,619,638]" gridrow="9" pageId="19">
<td box="[134,388,619,638]" gridcol="0" gridrow="9" pageId="19">Depth at dorsal-fin origin (m)</td>
<td box="[496,533,619,638]" gridcol="1" gridrow="9" pageId="19">-</td>
<td box="[553,575,619,638]" gridcol="2" gridrow="9" pageId="19">19</td>
<td box="[603,685,619,638]" gridcol="3" gridrow="9" pageId="19">26.4-31.5</td>
<td box="[718,767,619,638]" gridcol="4" gridrow="9" pageId="19">29.9</td>
<td box="[790,818,619,638]" gridcol="5" gridrow="9" pageId="19">1.2</td>
<td box="[836,857,619,638]" gridcol="6" gridrow="9" pageId="19">27</td>
<td box="[886,968,619,638]" gridcol="7" gridrow="9" pageId="19">25.6-33.4</td>
<td box="[1001,1050,619,638]" gridcol="8" gridrow="9" pageId="19">28.4</td>
<td box="[1073,1101,619,638]" gridcol="9" gridrow="9" pageId="19">1.8</td>
<td box="[1122,1159,619,638]" gridcol="10" gridrow="9" pageId="19">-</td>
<td box="[1180,1201,619,638]" gridcol="11" gridrow="9" pageId="19">-</td>
<td box="[1217,1311,619,638]" gridcol="12" gridrow="9" pageId="19">-</td>
<td box="[1344,1393,619,638]" gridcol="13" gridrow="9" pageId="19">-</td>
<td box="[1417,1444,619,638]" gridcol="14" gridrow="9" pageId="19">-</td>
</tr>
<tr box="[134,1444,644,663]" gridrow="10" pageId="19">
<td box="[134,388,644,663]" gridcol="0" gridrow="10" pageId="19">Depth at dorsal-fin origin (f)</td>
<td box="[496,533,644,663]" gridcol="1" gridrow="10" pageId="19">33.5</td>
<td box="[553,575,644,663]" gridcol="2" gridrow="10" pageId="19">24</td>
<td box="[603,685,644,663]" gridcol="3" gridrow="10" pageId="19">29.4-37.0</td>
<td box="[718,767,644,663]" gridcol="4" gridrow="10" pageId="19">32.5</td>
<td box="[790,818,644,663]" gridcol="5" gridrow="10" pageId="19">1.7</td>
<td box="[836,857,644,663]" gridcol="6" gridrow="10" pageId="19">70</td>
<td box="[886,968,644,663]" gridcol="7" gridrow="10" pageId="19">25.6-36.5</td>
<td box="[1001,1050,644,663]" gridcol="8" gridrow="10" pageId="19">31.5</td>
<td box="[1073,1101,644,663]" gridcol="9" gridrow="10" pageId="19">2.2</td>
<td box="[1122,1159,644,663]" gridcol="10" gridrow="10" pageId="19">32.4</td>
<td box="[1180,1201,644,663]" gridcol="11" gridrow="10" pageId="19">12</td>
<td box="[1217,1311,644,663]" gridcol="12" gridrow="10" pageId="19">30.0-33.6</td>
<td box="[1344,1393,644,663]" gridcol="13" gridrow="10" pageId="19">31.6</td>
<td box="[1417,1444,644,663]" gridcol="14" gridrow="10" pageId="19">1.3</td>
</tr>
<tr box="[134,1444,668,687]" gridrow="11" pageId="19">
<td box="[134,388,668,687]" gridcol="0" gridrow="11" pageId="19">Caudal peduncle depth</td>
<td box="[496,533,668,687]" gridcol="1" gridrow="11" pageId="19">12.4</td>
<td box="[553,575,668,687]" gridcol="2" gridrow="11" pageId="19">43</td>
<td box="[603,685,668,687]" gridcol="3" gridrow="11" pageId="19">10.6-12.5</td>
<td box="[718,767,668,687]" gridcol="4" gridrow="11" pageId="19">11.5</td>
<td box="[790,818,668,687]" gridcol="5" gridrow="11" pageId="19">0.5</td>
<td box="[836,857,668,687]" gridcol="6" gridrow="11" pageId="19">97</td>
<td box="[886,968,668,687]" gridcol="7" gridrow="11" pageId="19">9.5-13.0</td>
<td box="[1001,1050,668,687]" gridcol="8" gridrow="11" pageId="19">10.8</td>
<td box="[1073,1101,668,687]" gridcol="9" gridrow="11" pageId="19">0.6</td>
<td box="[1122,1159,668,687]" gridcol="10" gridrow="11" pageId="19">12.4</td>
<td box="[1180,1201,668,687]" gridcol="11" gridrow="11" pageId="19">12</td>
<td box="[1217,1311,668,687]" gridcol="12" gridrow="11" pageId="19">11.0-13.6</td>
<td box="[1344,1393,668,687]" gridcol="13" gridrow="11" pageId="19">12.2</td>
<td box="[1417,1444,668,687]" gridcol="14" gridrow="11" pageId="19">0.8</td>
</tr>
<tr box="[134,1444,693,712]" gridrow="12" pageId="19">
<td box="[134,388,693,712]" gridcol="0" gridrow="12" pageId="19">Caudal peduncle length</td>
<td box="[496,533,693,712]" gridcol="1" gridrow="12" pageId="19">13.5</td>
<td box="[553,575,693,712]" gridcol="2" gridrow="12" pageId="19">43</td>
<td box="[603,685,693,712]" gridcol="3" gridrow="12" pageId="19">10.7-15.3</td>
<td box="[718,767,693,712]" gridcol="4" gridrow="12" pageId="19">13.2</td>
<td box="[790,818,693,712]" gridcol="5" gridrow="12" pageId="19">1.0</td>
<td box="[836,857,693,712]" gridcol="6" gridrow="12" pageId="19">97</td>
<td box="[886,968,693,712]" gridcol="7" gridrow="12" pageId="19">11.1-14.9</td>
<td box="[1001,1050,693,712]" gridcol="8" gridrow="12" pageId="19">12.9</td>
<td box="[1073,1101,693,712]" gridcol="9" gridrow="12" pageId="19">0.8</td>
<td box="[1122,1159,693,712]" gridcol="10" gridrow="12" pageId="19">15.9</td>
<td box="[1180,1201,693,712]" gridcol="11" gridrow="12" pageId="19">12</td>
<td box="[1217,1311,693,712]" gridcol="12" gridrow="12" pageId="19">13.6-19.3</td>
<td box="[1344,1393,693,712]" gridcol="13" gridrow="12" pageId="19">16.8</td>
<td box="[1417,1444,693,712]" gridcol="14" gridrow="12" pageId="19">1.4</td>
</tr>
<tr box="[134,1444,717,736]" gridrow="13" pageId="19">
<td box="[134,388,717,736]" gridcol="0" gridrow="13" pageId="19">Anal-fin base</td>
<td box="[496,533,717,736]" gridcol="1" gridrow="13" pageId="19">33.2</td>
<td box="[553,575,717,736]" gridcol="2" gridrow="13" pageId="19">43</td>
<td box="[603,685,717,736]" gridcol="3" gridrow="13" pageId="19">31.4-37.7</td>
<td box="[718,767,717,736]" gridcol="4" gridrow="13" pageId="19">34.6</td>
<td box="[790,818,717,736]" gridcol="5" gridrow="13" pageId="19">1.3</td>
<td box="[836,857,717,736]" gridcol="6" gridrow="13" pageId="19">97</td>
<td box="[886,968,717,736]" gridcol="7" gridrow="13" pageId="19">30.4-36.4</td>
<td box="[1001,1050,717,736]" gridcol="8" gridrow="13" pageId="19">33.2</td>
<td box="[1073,1101,717,736]" gridcol="9" gridrow="13" pageId="19">1.1</td>
<td box="[1122,1159,717,736]" gridcol="10" gridrow="13" pageId="19">28.2</td>
<td box="[1180,1201,717,736]" gridcol="11" gridrow="13" pageId="19">12</td>
<td box="[1217,1311,717,736]" gridcol="12" gridrow="13" pageId="19">26.6-30.4</td>
<td box="[1344,1393,717,736]" gridcol="13" gridrow="13" pageId="19">28.0</td>
<td box="[1417,1444,717,736]" gridcol="14" gridrow="13" pageId="19">1.1</td>
</tr>
<tr box="[134,1444,742,761]" gridrow="14" pageId="19">
<td box="[134,388,742,761]" gridcol="0" gridrow="14" pageId="19">Dorsal-fin length</td>
<td box="[496,533,742,761]" gridcol="1" gridrow="14" pageId="19">22.6</td>
<td box="[553,575,742,761]" gridcol="2" gridrow="14" pageId="19">43</td>
<td box="[603,685,742,761]" gridcol="3" gridrow="14" pageId="19">21.6-26.6</td>
<td box="[718,767,742,761]" gridcol="4" gridrow="14" pageId="19">24.0</td>
<td box="[790,818,742,761]" gridcol="5" gridrow="14" pageId="19">1.1</td>
<td box="[836,857,742,761]" gridcol="6" gridrow="14" pageId="19">97</td>
<td box="[886,968,742,761]" gridcol="7" gridrow="14" pageId="19">19.0-27.6</td>
<td box="[1001,1050,742,761]" gridcol="8" gridrow="14" pageId="19">24.0</td>
<td box="[1073,1101,742,761]" gridcol="9" gridrow="14" pageId="19">1.1</td>
<td box="[1122,1159,742,761]" gridcol="10" gridrow="14" pageId="19">23.4</td>
<td box="[1180,1201,742,761]" gridcol="11" gridrow="14" pageId="19">12</td>
<td box="[1217,1311,742,761]" gridcol="12" gridrow="14" pageId="19">22.8-25.6</td>
<td box="[1344,1393,742,761]" gridcol="13" gridrow="14" pageId="19">23.7</td>
<td box="[1417,1444,742,761]" gridcol="14" gridrow="14" pageId="19">0.7</td>
</tr>
<tr box="[134,1444,766,785]" gridrow="15" pageId="19">
<td box="[134,388,766,785]" gridcol="0" gridrow="15" pageId="19">Pelvic-fin length (m)</td>
<td box="[496,533,766,785]" gridcol="1" gridrow="15" pageId="19">-</td>
<td box="[553,575,766,785]" gridcol="2" gridrow="15" pageId="19">19</td>
<td box="[603,685,766,785]" gridcol="3" gridrow="15" pageId="19">13.4-16.5</td>
<td box="[718,767,766,785]" gridcol="4" gridrow="15" pageId="19">14.7</td>
<td box="[790,818,766,785]" gridcol="5" gridrow="15" pageId="19">0.9</td>
<td box="[836,857,766,785]" gridcol="6" gridrow="15" pageId="19">27</td>
<td box="[886,968,766,785]" gridcol="7" gridrow="15" pageId="19">13.0-15.6</td>
<td box="[1001,1050,766,785]" gridcol="8" gridrow="15" pageId="19">14.4</td>
<td box="[1073,1101,766,785]" gridcol="9" gridrow="15" pageId="19">0.7</td>
<td box="[1122,1159,766,785]" gridcol="10" gridrow="15" pageId="19">-</td>
<td box="[1180,1201,766,785]" gridcol="11" gridrow="15" pageId="19">-</td>
<td box="[1217,1311,766,785]" gridcol="12" gridrow="15" pageId="19">-</td>
<td box="[1344,1393,766,785]" gridcol="13" gridrow="15" pageId="19">-</td>
<td box="[1417,1444,766,785]" gridcol="14" gridrow="15" pageId="19">-</td>
</tr>
<tr box="[134,1444,791,810]" gridrow="16" pageId="19">
<td box="[134,388,791,810]" gridcol="0" gridrow="16" pageId="19">Pelvic-fin length (f)</td>
<td box="[496,533,791,810]" gridcol="1" gridrow="16" pageId="19">14.0</td>
<td box="[553,575,791,810]" gridcol="2" gridrow="16" pageId="19">24</td>
<td box="[603,685,791,810]" gridcol="3" gridrow="16" pageId="19">13.0-15.2</td>
<td box="[718,767,791,810]" gridcol="4" gridrow="16" pageId="19">14.1</td>
<td box="[790,818,791,810]" gridcol="5" gridrow="16" pageId="19">0.6</td>
<td box="[836,857,791,810]" gridcol="6" gridrow="16" pageId="19">70</td>
<td box="[886,968,791,810]" gridcol="7" gridrow="16" pageId="19">12.7-16.3</td>
<td box="[1001,1050,791,810]" gridcol="8" gridrow="16" pageId="19">14.1</td>
<td box="[1073,1101,791,810]" gridcol="9" gridrow="16" pageId="19">0.7</td>
<td box="[1122,1159,791,810]" gridcol="10" gridrow="16" pageId="19">13.6</td>
<td box="[1180,1201,791,810]" gridcol="11" gridrow="16" pageId="19">12</td>
<td box="[1217,1311,791,810]" gridcol="12" gridrow="16" pageId="19">13.0-14.6</td>
<td box="[1344,1393,791,810]" gridcol="13" gridrow="16" pageId="19">13.8</td>
<td box="[1417,1444,791,810]" gridcol="14" gridrow="16" pageId="19">0.5</td>
</tr>
<tr box="[134,1444,815,834]" gridrow="17" pageId="19">
<td box="[134,388,815,834]" gridcol="0" gridrow="17" pageId="19">Pectoral-fin length (m)</td>
<td box="[496,533,815,834]" gridcol="1" gridrow="17" pageId="19">-</td>
<td box="[553,575,815,834]" gridcol="2" gridrow="17" pageId="19">19</td>
<td box="[603,685,815,834]" gridcol="3" gridrow="17" pageId="19">18.8-25.8</td>
<td box="[718,767,815,834]" gridcol="4" gridrow="17" pageId="19">21.5</td>
<td box="[790,818,815,834]" gridcol="5" gridrow="17" pageId="19">1.6</td>
<td box="[836,857,815,834]" gridcol="6" gridrow="17" pageId="19">27</td>
<td box="[886,968,815,834]" gridcol="7" gridrow="17" pageId="19">19.5-22.0</td>
<td box="[1001,1050,815,834]" gridcol="8" gridrow="17" pageId="19">21.0</td>
<td box="[1073,1101,815,834]" gridcol="9" gridrow="17" pageId="19">0.6</td>
<td box="[1122,1159,815,834]" gridcol="10" gridrow="17" pageId="19">-</td>
<td box="[1180,1201,815,834]" gridcol="11" gridrow="17" pageId="19">-</td>
<td box="[1217,1311,815,834]" gridcol="12" gridrow="17" pageId="19">-</td>
<td box="[1344,1393,815,834]" gridcol="13" gridrow="17" pageId="19">-</td>
<td box="[1417,1444,815,834]" gridcol="14" gridrow="17" pageId="19">-</td>
</tr>
<tr box="[134,1444,840,859]" gridrow="18" pageId="19">
<td box="[134,388,840,859]" gridcol="0" gridrow="18" pageId="19">Pectoral-fin length (f)</td>
<td box="[496,533,840,859]" gridcol="1" gridrow="18" pageId="19">21.2</td>
<td box="[553,575,840,859]" gridcol="2" gridrow="18" pageId="19">24</td>
<td box="[603,685,840,859]" gridcol="3" gridrow="18" pageId="19">19.5-23.6</td>
<td box="[718,767,840,859]" gridcol="4" gridrow="18" pageId="19">21.1</td>
<td box="[790,818,840,859]" gridcol="5" gridrow="18" pageId="19">1.0</td>
<td box="[836,857,840,859]" gridcol="6" gridrow="18" pageId="19">70</td>
<td box="[886,968,840,859]" gridcol="7" gridrow="18" pageId="19">18.6-22.8</td>
<td box="[1001,1050,840,859]" gridcol="8" gridrow="18" pageId="19">21.3</td>
<td box="[1073,1101,840,859]" gridcol="9" gridrow="18" pageId="19">0.8</td>
<td box="[1122,1159,840,859]" gridcol="10" gridrow="18" pageId="19">22.1</td>
<td box="[1180,1201,840,859]" gridcol="11" gridrow="18" pageId="19">12</td>
<td box="[1217,1311,840,859]" gridcol="12" gridrow="18" pageId="19">20.7-23.7</td>
<td box="[1344,1393,840,859]" gridcol="13" gridrow="18" pageId="19">22.1</td>
<td box="[1417,1444,840,859]" gridcol="14" gridrow="18" pageId="19">0.7</td>
</tr>
<tr box="[134,1444,864,883]" gridrow="19" pageId="19">
<td box="[134,388,864,883]" gridcol="0" gridrow="19" pageId="19">Head length</td>
<td box="[496,533,864,883]" gridcol="1" gridrow="19" pageId="19">21.4</td>
<td box="[553,575,864,883]" gridcol="2" gridrow="19" pageId="19">43</td>
<td box="[603,685,864,883]" gridcol="3" gridrow="19" pageId="19">20.4-23.6</td>
<td box="[718,767,864,883]" gridcol="4" gridrow="19" pageId="19">21.6</td>
<td box="[790,818,864,883]" gridcol="5" gridrow="19" pageId="19">0.7</td>
<td box="[836,857,864,883]" gridcol="6" gridrow="19" pageId="19">97</td>
<td box="[886,968,864,883]" gridcol="7" gridrow="19" pageId="19">21.4-25.0</td>
<td box="[1001,1050,864,883]" gridcol="8" gridrow="19" pageId="19">23.2</td>
<td box="[1073,1101,864,883]" gridcol="9" gridrow="19" pageId="19">0.8</td>
<td box="[1122,1159,864,883]" gridcol="10" gridrow="19" pageId="19">23.8</td>
<td box="[1180,1201,864,883]" gridcol="11" gridrow="19" pageId="19">12</td>
<td box="[1217,1311,864,883]" gridcol="12" gridrow="19" pageId="19">23.7-25.8</td>
<td box="[1344,1393,864,883]" gridcol="13" gridrow="19" pageId="19">24.8</td>
<td box="[1417,1444,864,883]" gridcol="14" gridrow="19" pageId="19">0.7</td>
</tr>
<tr box="[134,1444,889,908]" gridrow="20" pageId="19" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-2="1" rowspan-8="1" rowspan-9="1">
<td box="[603,968,889,908]" colspan="5" colspanRight="4" gridcol="3" gridrow="20" pageId="19">Percents of Head length</td>
</tr>
<tr box="[134,1444,913,932]" gridrow="21" pageId="19">
<td box="[134,388,913,932]" gridcol="0" gridrow="21" pageId="19">Snout length</td>
<td box="[496,533,913,932]" gridcol="1" gridrow="21" pageId="19">24.9</td>
<td box="[553,575,913,932]" gridcol="2" gridrow="21" pageId="19">43</td>
<td box="[603,685,913,932]" gridcol="3" gridrow="21" pageId="19">20.2-24.4</td>
<td box="[718,767,913,932]" gridcol="4" gridrow="21" pageId="19">22.0</td>
<td box="[790,818,913,932]" gridcol="5" gridrow="21" pageId="19">0.9</td>
<td box="[836,857,913,932]" gridcol="6" gridrow="21" pageId="19">97</td>
<td box="[886,968,913,932]" gridcol="7" gridrow="21" pageId="19">18.3-24.3</td>
<td box="[1001,1050,913,932]" gridcol="8" gridrow="21" pageId="19">22.1</td>
<td box="[1073,1101,913,932]" gridcol="9" gridrow="21" pageId="19">1.1</td>
<td box="[1122,1159,913,932]" gridcol="10" gridrow="21" pageId="19">21.3</td>
<td box="[1180,1201,913,932]" gridcol="11" gridrow="21" pageId="19">12</td>
<td box="[1217,1311,913,932]" gridcol="12" gridrow="21" pageId="19">19.3-22.9</td>
<td box="[1344,1393,913,932]" gridcol="13" gridrow="21" pageId="19">20.9</td>
<td box="[1417,1444,913,932]" gridcol="14" gridrow="21" pageId="19">1.0</td>
</tr>
<tr box="[134,1444,938,957]" gridrow="22" pageId="19">
<td box="[134,388,938,957]" gridcol="0" gridrow="22" pageId="19">Upper jaw length</td>
<td box="[496,533,938,957]" gridcol="1" gridrow="22" pageId="19">45.3</td>
<td box="[553,575,938,957]" gridcol="2" gridrow="22" pageId="19">43</td>
<td box="[603,685,938,957]" gridcol="3" gridrow="22" pageId="19">40.9-46.5</td>
<td box="[718,767,938,957]" gridcol="4" gridrow="22" pageId="19">43.6</td>
<td box="[790,818,938,957]" gridcol="5" gridrow="22" pageId="19">1.4</td>
<td box="[836,857,938,957]" gridcol="6" gridrow="22" pageId="19">96</td>
<td box="[886,968,938,957]" gridcol="7" gridrow="22" pageId="19">40.8-50.3</td>
<td box="[1001,1050,938,957]" gridcol="8" gridrow="22" pageId="19">45.4</td>
<td box="[1073,1101,938,957]" gridcol="9" gridrow="22" pageId="19">1.7</td>
<td box="[1122,1159,938,957]" gridcol="10" gridrow="22" pageId="19">50.2</td>
<td box="[1180,1201,938,957]" gridcol="11" gridrow="22" pageId="19">12</td>
<td box="[1217,1311,938,957]" gridcol="12" gridrow="22" pageId="19">44.5-50.2</td>
<td box="[1344,1393,938,957]" gridcol="13" gridrow="22" pageId="19">47.7</td>
<td box="[1417,1444,938,957]" gridcol="14" gridrow="22" pageId="19">1.7</td>
</tr>
<tr box="[134,1444,962,981]" gridrow="23" pageId="19">
<td box="[134,388,962,981]" gridcol="0" gridrow="23" pageId="19">Orbital diameter</td>
<td box="[496,533,962,981]" gridcol="1" gridrow="23" pageId="19">28.7</td>
<td box="[553,575,962,981]" gridcol="2" gridrow="23" pageId="19">43</td>
<td box="[603,685,962,981]" gridcol="3" gridrow="23" pageId="19">26.0-34.5</td>
<td box="[718,767,962,981]" gridcol="4" gridrow="23" pageId="19">31.1</td>
<td box="[790,818,962,981]" gridcol="5" gridrow="23" pageId="19">1.8</td>
<td box="[836,857,962,981]" gridcol="6" gridrow="23" pageId="19">97</td>
<td box="[886,968,962,981]" gridcol="7" gridrow="23" pageId="19">26.8-38.7</td>
<td box="[1001,1050,962,981]" gridcol="8" gridrow="23" pageId="19">34.2</td>
<td box="[1073,1101,962,981]" gridcol="9" gridrow="23" pageId="19">1.8</td>
<td box="[1122,1159,962,981]" gridcol="10" gridrow="23" pageId="19">33.2</td>
<td box="[1180,1201,962,981]" gridcol="11" gridrow="23" pageId="19">12</td>
<td box="[1217,1311,962,981]" gridcol="12" gridrow="23" pageId="19">33.2-37.6</td>
<td box="[1344,1393,962,981]" gridcol="13" gridrow="23" pageId="19">35.7</td>
<td box="[1417,1444,962,981]" gridcol="14" gridrow="23" pageId="19">1.1</td>
</tr>
<tr box="[134,1444,987,1006]" gridrow="24" pageId="19">
<td box="[134,388,987,1006]" gridcol="0" gridrow="24" pageId="19">Interorbital width</td>
<td box="[496,533,987,1006]" gridcol="1" gridrow="24" pageId="19">33.1</td>
<td box="[553,575,987,1006]" gridcol="2" gridrow="24" pageId="19">43</td>
<td box="[603,685,987,1006]" gridcol="3" gridrow="24" pageId="19">30.3-35.9</td>
<td box="[718,767,987,1006]" gridcol="4" gridrow="24" pageId="19">32.7</td>
<td box="[790,818,987,1006]" gridcol="5" gridrow="24" pageId="19">1.2</td>
<td box="[836,857,987,1006]" gridcol="6" gridrow="24" pageId="19">97</td>
<td box="[886,968,987,1006]" gridcol="7" gridrow="24" pageId="19">29.6-34.7</td>
<td box="[1001,1050,987,1006]" gridcol="8" gridrow="24" pageId="19">32.4</td>
<td box="[1073,1101,987,1006]" gridcol="9" gridrow="24" pageId="19">1.1</td>
<td box="[1122,1159,987,1006]" gridcol="10" gridrow="24" pageId="19">34.1</td>
<td box="[1180,1201,987,1006]" gridcol="11" gridrow="24" pageId="19">12</td>
<td box="[1217,1311,987,1006]" gridcol="12" gridrow="24" pageId="19">32.7-35.6</td>
<td box="[1344,1393,987,1006]" gridcol="13" gridrow="24" pageId="19">34.1</td>
<td box="[1417,1444,987,1006]" gridcol="14" gridrow="24" pageId="19">0.8</td>
</tr>
</table>
</paragraph>
<subSubSection pageId="19" type="etymology">
<paragraph blockId="19.[129,771,1062,1149]" pageId="19">
<emphasis bold="true" box="[129,253,1062,1085]" pageId="19">Etymology.</emphasis>
The name 
<taxonomicName box="[364,490,1062,1085]" class="Actinopterygii" family="Characidae" genus="Hemibrycon" kingdom="Animalia" order="Characiformes" pageId="19" phylum="Chordata" rank="species" species="mikrostiktos">
<emphasis box="[364,490,1062,1085]" italics="true" pageId="19">mikrostiktos</emphasis>
</taxonomicName>
is from the Greek, 
<emphasis box="[683,764,1062,1085]" italics="true" pageId="19">mickros</emphasis>
, small, and 
<emphasis box="[242,311,1094,1117]" italics="true" pageId="19">stiktos</emphasis>
, spot, referring to the small humeral spot. A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>