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<mods:title id="F2EB5700E0712A3F9BBDE0B4B09759B0">A description of the first complete specimen of Diogenes guttatus Henderson, 1888 (Decapoda: Anomura: Paguroidea: Diogenidae)</mods:title>
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<taxonomicName id="4C554D65774B7A0EFEC6C5F81D9BFB5E" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[264,699,1200,1226]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="1" pageNumber="2" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774B7A0EFEC6C5F81EC1FB5E" bold="true" box="[264,481,1200,1226]" italics="true" pageId="1" pageNumber="2">Diogenes guttatus</emphasis>
<bibRefCitation id="EFC44B17774B7A0EFE26C5F81D9BFB5E" author="Henderson" box="[488,699,1200,1226]" lastPageId="1" lastPageNumber="8" pageId="1" pageNumber="7" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger, (Zoology) Volume 27. Her Majesty's Stationary Office, Edinburgh etc., 221 pp." type="book chapter" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
(
<figureCitation id="136E2A63774B7A0EFD04C5F81C3DFB5E" box="[714,797,1200,1226]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
, 
<figureCitation id="136E2A63774B7A0EFCE2C5F81C1CFB5E" box="[812,828,1200,1226]" captionStart="FIGURE 2" captionStartId="3.[264,367,1598,1622]" captionTargetBox="[284,1289,836,1571]" captionTargetId="figure@3.[264,1323,812,1596]" captionTargetPageId="3" captionText="FIGURE 2. Diogenes guttatus Henderson, 1888, female, sl = 2.7 mm, off Thursday Island (AM 67380). A, chela and carpus of left cheliped (outer face); B, chela and carpus of right cheliped (outer face). Scale equals 1 mm." httpUri="https://zenodo.org/record/157219/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
)
</emphasis>
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</paragraph>
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<subSubSection id="C34F656D774B7A0EFEC6C4441CBCFA37" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BEA36E6774B7A0EFEC6C4441DA1FAF7" blockId="1.[264,1323,1292,1443]" pageId="1" pageNumber="2">
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<taxonomicName id="4C554D65774B7A0EFEC6C4441D99FAB7" authority="Henderson, 1888: 54" authorityName="Henderson" authorityPageNumber="54" authorityYear="1888" box="[264,697,1292,1315]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="1" pageNumber="2" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774B7A0EFEC6C4441EEBFAB7" box="[264,459,1292,1315]" italics="true" pageId="1" pageNumber="2">Diogenes guttatus</emphasis>
<treatmentCitation id="0AF410F7774B7A0EFE1AC4441D99FAB7" author="Henderson" box="[468,697,1292,1315]" page="54" pageId="1" pageNumber="2" year="1888">
<bibRefCitation id="EFC44B17774B7A0EFE1AC4441DB0FAB7" author="Henderson" box="[468,656,1292,1315]" lastPageId="1" lastPageNumber="8" pageId="1" pageNumber="7" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger, (Zoology) Volume 27. Her Majesty's Stationary Office, Edinburgh etc., 221 pp." type="book chapter" year="1888">Henderson, 1888</bibRefCitation>
: 54
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</taxonomicName>
, pl. 6, fig. 4, 4a; 
<treatmentCitation id="0AF410F7774B7A0EFCB4C4441B63FAB7" author="Alcock" box="[890,1091,1292,1315]" page="166" pageId="1" pageNumber="2" year="1905">
<bibRefCitation id="EFC44B17774B7A0EFCB4C4441B2FFAB7" author="Alcock" box="[890,1039,1292,1315]" pageId="1" pageNumber="7" refString="Alcock, A. (1905) Anomura. Fascicle I. Pagurides. - Catalogue of the Indian decapod Crustacea in the collections of the Indian Museum, Volume 2. Indian Museum Calcutta, 197 pp." type="book" year="1905">Alcock, 1905</bibRefCitation>
: 166
</treatmentCitation>
(list); 
<treatmentCitation id="0AF410F7774B7A0EFB40C4441E40FAD7" author="Gordan" page="317" pageId="1" pageNumber="2" year="1956">
<bibRefCitation id="EFC44B17774B7A0EFB40C4441A07FAB7" author="Gordan" box="[1166,1319,1292,1315]" pageId="1" pageNumber="7" refString="Gordan, J. (1956) A bibliography of pagurid crabs, exclusive of Alcock, 1905. Bulletin of the American Museum of Natural History, 108, 253 - 352." type="journal article" year="1956">
<collectingCountry id="F3427676774B7A0EFB40C4441BC1FAB7" box="[1166,1249,1292,1315]" name="Jordan" pageId="1" pageNumber="2">Gordan</collectingCountry>
, 1956
</bibRefCitation>
: 317
</treatmentCitation>
(lit.); 
<treatmentCitation id="0AF410F7774B7A0EFE63C4641DD4FAD7" author="Morgan" box="[429,756,1324,1347]" page="664" pageId="1" pageNumber="2" year="1991">
<bibRefCitation id="EFC44B17774B7A0EFE63C4641D9AFAD7" author="Morgan" box="[429,698,1324,1347]" pageId="1" pageNumber="8" refString="Morgan, G. J. &amp; Forest, J. (1991) Seven new species of hermit crabs from northern and western Australia (Decapoda, Anomura, Diogenidae). Bulletin du Museum national d'Histoire naturelle, (4) A, 12, 649 - 689." type="journal article" year="1991">Morgan &amp; Forest, 1991</bibRefCitation>
: 664
</treatmentCitation>
; 
<treatmentCitation id="0AF410F7774B7A0EFCCAC4641B39FAD7" author="McLaughlin" box="[772,1049,1324,1347]" page="411" pageId="1" pageNumber="2" year="2002">
<bibRefCitation id="EFC44B17774B7A0EFCCAC4641CC1FAD7" author="McLaughlin" box="[772,993,1324,1347]" pageId="1" pageNumber="8" refString="McLaughlin, P. A. (2002 b) A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Crustacea of the Andaman Sea 29 November - 20 December, 1998, Phuket Marine Biological Center. Phuket Marine Biological Center Special Publication, 23, 385 - 460." type="journal article" year="2002" yearSuffix="b">McLaughlin, 2002b</bibRefCitation>
: 411
</treatmentCitation>
, fig. 1A, B; 
<treatmentCitation id="0AF410F7774B7A0EFB61C4641D16FAF7" author="Siddiqui" page="961" pageId="1" pageNumber="2" year="2003">
<bibRefCitation id="EFC44B17774B7A0EFB61C4641D22FAF7" author="Siddiqui" pageId="1" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin, 2003</bibRefCitation>
: 961
</treatmentCitation>
, fig. 4.
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</paragraph>
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Not 
<taxonomicName id="4C554D65774B7A0EFEF9C4241ED8FA17" box="[311,504,1388,1411]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="1" pageNumber="2" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774B7A0EFEF9C4241ED8FA17" box="[311,504,1388,1411]" italics="true" pageId="1" pageNumber="2">Diogenes guttatus</emphasis>
</taxonomicName>
. – 
<bibRefCitation id="EFC44B17774B7A0EFDD5C4241C0EFA17" author="Tirmizi" box="[539,814,1388,1411]" pageId="1" pageNumber="8" refString="Tirmizi, N. M. &amp; Siddiqui, F. A. (1981) An illustrated key to the identification of northern Arabian Sea pagurids. Institute of Marine Biology, 1, 1 - 31." type="journal article" year="1981">Tirmizi &amp; Siddiqui, 1981</bibRefCitation>
: fig. 17; 
<treatmentCitation id="0AF410F7774B7A0EFC43C4241BE8FA17" author="Tirmizi" box="[909,1224,1388,1411]" page="57" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="EFC44B17774B7A0EFC43C4241B80FA17" author="Tirmizi" box="[909,1184,1388,1411]" pageId="1" pageNumber="8" refString="Tirmizi, N. M. &amp; Siddiqui, F. A. (1982) The marine fauna of Pakistan: 1 Hermit crabs (Crustacea, Anomura). University Grants Commission, Karachi, 103 pp." type="book" year="1982">Tirmizi &amp; Siddiqui, 1982</bibRefCitation>
: 57
</treatmentCitation>
, figs. 30, 31 (= 
<taxonomicName id="4C554D65774B7A0EFEB6C4C41CB0FA37" authority="Siddiqui &amp; McLaughlin, 2003" authorityName="Siddiqui &amp; McLaughlin" authorityYear="2003" box="[376,912,1420,1443]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="1" pageNumber="2" phylum="Chlorophyta" rank="species" species="tirmiziae">
<emphasis id="B921EAF4774B7A0EFEB6C4C41D62FA37" box="[376,578,1420,1443]" italics="true" pageId="1" pageNumber="2">Diogenes tirmiziae</emphasis>
<bibRefCitation id="EFC44B17774B7A0EFD87C4C41CB0FA37" author="Siddiqui" box="[585,912,1420,1443]" pageId="1" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin, 2003</bibRefCitation>
</taxonomicName>
).
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<emphasis id="B921EAF4774B7A0EFEC6C49C1C03FA7A" bold="true" box="[264,803,1492,1518]" pageId="1" pageNumber="2">
Material examined. Queensland, 
<collectingCountry id="F3427676774B7A0EFD60C49C1C03FA7A" box="[686,803,1492,1518]" name="Australia" pageId="1" pageNumber="2">Australia</collectingCountry>
</emphasis>
. 
<typeStatus id="54EE8844774B7A0EFCFBC49C1C82FA7A" box="[821,930,1492,1518]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
male (sl = 
<quantity id="4CAD9B03774B7A0EFBFEC49C1BB1FA7A" box="[1072,1169,1492,1518]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="2" unit="mm" value="3.2">3.2 mm</quantity>
), 
<emphasis id="B921EAF4774B7A0EFB69C49C1A0BFA79" box="[1191,1323,1492,1517]" italics="true" pageId="1" pageNumber="2">Challenger</emphasis>
station 187, off Cape York, 
<geoCoordinate id="EE615021774B7A0EFD95C4B41D9EF982" box="[603,702,1532,1558]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-10.6">10°36’S</geoCoordinate>
, 
<geoCoordinate id="EE615021774B7A0EFD03C4B41C60F982" box="[717,832,1532,1558]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="141.91667">141°55’E</geoCoordinate>
, 
<quantity id="4CAD9B03774B7A0EFC81C4B41CAFF982" box="[847,911,1532,1558]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="2" unit="m" value="11.0">11 m</quantity>
, 
<date id="FFEB1026774B7A0EFC50C4B41B08F982" box="[926,1064,1532,1558]" pageId="1" pageNumber="2" value="1874-09-09">9 Sep 1874</date>
(NHM 1888.33.1); 
<specimenCount id="9D53FD6F774B7A0EFAD2C4B41E78F9AA" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(sl = 
<quantity id="4CAD9B03774B7A0EFE52C76C1ED9F9AA" box="[412,505,1572,1598]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="2" unit="mm" value="2.7">2.7 mm</quantity>
), off Thursday Island, 
<geoCoordinate id="EE615021774B7A0EFCCDC76C1C45F9AA" box="[771,869,1572,1598]" direction="south" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="-10.583333">10°35’S</geoCoordinate>
, 
<geoCoordinate id="EE615021774B7A0EFCBCC76C1CC3F9AA" box="[882,995,1572,1598]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="142.21666">142°13’E</geoCoordinate>
(AM P 67380).
</paragraph>
<paragraph id="8BEA36E6774B7A0EFEF6C7041D6CF8C2" blockId="1.[264,1323,1492,1878]" pageId="1" pageNumber="2">
<emphasis id="B921EAF4774B7A0EFEF6C7041ECDF9F2" bold="true" box="[312,493,1612,1638]" pageId="1" pageNumber="2">Redescription.</emphasis>
Shield (
<figureCitation id="136E2A63774B7A0EFD90C7041D81F9F2" box="[606,673,1612,1638]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
A) slightly broader than long, weakly convex; central dorsal surface smooth or with scattered tiny spinules; lateral marginal areas generally unarmed or with numerous small spines and spinules, each also with protuberant spinulose ridge in proximal half. Rostrum obsolete; lateral projections prominently produced, each with 1or more terminal spines or spinules. Dorsal margins of branchiostegites each with 5 or 6 acute spines. Intercalary rostriform process simple, tapering to acute tip, not reaching to apices of acicular spines.
</paragraph>
<caption id="DF2A666E77487A0DFEC6C4F21C4FF9C8" httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="2" pageNumber="3" targetBox="[266,1321,328,1465]" targetPageId="2">
<paragraph id="8BEA36E677487A0DFEC6C4F21C4FF9C8" blockId="2.[264,1323,1466,1628]" pageId="2" pageNumber="3">
<emphasis id="B921EAF477487A0DFEC6C4F21EA3FA46" bold="true" box="[264,387,1466,1490]" pageId="2" pageNumber="3">FIGURE 1</emphasis>
. 
<taxonomicName id="4C554D6577487A0DFE58C4F21C36FA45" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[406,790,1466,1489]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="2" pageNumber="3" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF477487A0DFE58C4F21D77FA45" box="[406,599,1466,1489]" italics="true" pageId="2" pageNumber="3">Diogenes guttatus</emphasis>
Henderson, 1888
</taxonomicName>
, A–D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm.
</paragraph>
</caption>
<paragraph id="8BEA36E677487A0DFEF6C7F01C34F8B6" blockId="2.[264,1323,1720,1906]" pageId="2" pageNumber="3">Ocular peduncles 0.8 to almost full length of shield, dorsal surfaces each with or without row of sparse setae; corneal diameter 0.2–0.3 peduncular length; ocular acicles with 2– 5 spines on distal margin, innermost longest.</paragraph>
<paragraph id="8BEA36E677487A0CFEF6C6781EEAFECA" blockId="2.[264,1323,1720,1906]" lastBlockId="3.[264,1324,284,750]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Antennular peduncles overreaching distal corneal margins by 0.8 to entire length of ultimate segments; ultimate and penultimate segments each with few scattered, long setae; basal segments each with row of small spines on distolateral margin, continued onto ventrodistal margin.</paragraph>
<paragraph id="8BEA36E677497A0CFEF6C0241B34FD7A" blockId="3.[264,1324,284,750]" pageId="3" pageNumber="4">Antennal peduncles overreaching distal corneal margins by 0.4–0.6 length of ultimate segments; fifth segments each with 2 rows of long setae on ventral margin; fourth and third segments each with few scattered long setae; second segments each with dorsolateral distal angle produced into small to prominent spine, dorsomesial distal angle unarmed or with small spine, distolateral margin with row of spines ventrally, continued onto ventrodistal margin; first segments each with row of spines on distolateral margin ventrally, continued on ventrodistal margin. Antennal acicle short, broad, generally subquadrate, outermost spine reaching to or beyond midlength of penultimate peduncular segment, anterior margin with 2 or 3 additional prominent spines. Antennal flagellum shorter than twice carapace length, articles each with pair of long ventral setae.</paragraph>
<caption id="DF2A666E77497A0CFEC6C7761D47F90F" httpUri="https://zenodo.org/record/157219/files/figure.png" pageId="3" pageNumber="4" targetBox="[284,1289,836,1571]" targetPageId="3">
<paragraph id="8BEA36E677497A0CFEC6C7761D47F90F" blockId="3.[264,1323,1598,1691]" pageId="3" pageNumber="4">
<emphasis id="B921EAF477497A0CFEC6C7761EA5F9C2" bold="true" box="[264,389,1598,1622]" pageId="3" pageNumber="4">FIGURE 2</emphasis>
. 
<taxonomicName id="4C554D6577497A0CFE5BC7771C3DF9C1" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" box="[405,797,1598,1622]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="3" pageNumber="4" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF477497A0CFE5BC7771D79F9C2" box="[405,601,1599,1622]" italics="true" pageId="3" pageNumber="4">Diogenes guttatus</emphasis>
Henderson, 1888
</taxonomicName>
, female, sl = 2.7 mm, off Thursday Island (AM 67380). A, chela and carpus of left cheliped (outer face); B, chela and carpus of right cheliped (outer face). Scale equals 1 mm.
</paragraph>
</caption>
<paragraph id="8BEA36E677497A0BFEF6C7BF1D96FB92" blockId="3.[264,1324,1783,1889]" lastBlockId="4.[264,1324,284,1870]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Left cheliped (
<figureCitation id="136E2A6377497A0CFE29C7BF1D0EF885" box="[487,558,1783,1809]" captionStart="FIGURE 2" captionStartId="3.[264,367,1598,1622]" captionTargetBox="[284,1289,836,1571]" captionTargetId="figure@3.[264,1323,812,1596]" captionTargetPageId="3" captionText="FIGURE 2. Diogenes guttatus Henderson, 1888, female, sl = 2.7 mm, off Thursday Island (AM 67380). A, chela and carpus of left cheliped (outer face); B, chela and carpus of right cheliped (outer face). Scale equals 1 mm." httpUri="https://zenodo.org/record/157219/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
A) with slight to prominent hiatus between dactyl and fixed finger. Dactyl equal to length of palm or slightly shorter; with row of small to moderately large spines on upper margin, adjacent shorter row of smaller spines weakly to well developed; outer surface with numerous spinules and flattened tubercles; inner surface with scattered, moderately short setae; cutting edge with row of small, calcareous teeth; dactyl overlapping fixed finger in 
<typeStatus id="54EE8844774E7A0BFD85C0241D95FE12" box="[587,693,364,390]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, but overlapped by fixed finger in female. Palm with irregular single to double row of small to moderately large spines on upper margin; convex outer surface with covering of circular, mushroom­shaped, flattened tubercles and few spinules, often tear­drop in shape, lower margin with row of spines, 3 moderately stout, short (
<typeStatus id="54EE8844774E7A0BFE9DC3441E9DFDB2" box="[339,445,524,550]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
) or several moderate to very prominent spines (female) near lower proximal angle, continued on proximal marginal as weak (
<typeStatus id="54EE8844774E7A0BFCB6C37C1CC2FDDA" box="[888,994,564,590]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
) or prominent (female) row of spines curving onto outer face; inner face with scattered low protuberances. Carpus with irregular single or double row of small to moderately large spines on upper margin, outer surface with covering of spinules, denser in lower half, lower margin with 2–5 small to large spines distally, lower surface with numerous spinulose tubercles, distal margin with row of very small, tuberculate spinules; inner surface with scattered spinules. Merus with row of spinulose protuberances and long setae on dorsal margin; mesial face with scattered spinules dorsally and tuberculate spines ventrally, ventromesial margin with irregular row of tuberculate spines; lateral face with scattered spinules, ventrolateral margin with irregular double row of spines; ventral surface with scattered small spines and spinules. Ischium with short row of small tubercles on laterodistal margin ventrally; ventromesial margin with few tubercles.
</paragraph>
<paragraph id="8BEA36E6774E7A0BFEF6C55C1BA4F96A" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">
Right cheliped (
<figureCitation id="136E2A63774E7A0BFE3BC55C1D1CFBBA" box="[501,572,1044,1070]" captionStart="FIGURE 2" captionStartId="3.[264,367,1598,1622]" captionTargetBox="[284,1289,836,1571]" captionTargetId="figure@3.[264,1323,812,1596]" captionTargetPageId="3" captionText="FIGURE 2. Diogenes guttatus Henderson, 1888, female, sl = 2.7 mm, off Thursday Island (AM 67380). A, chela and carpus of left cheliped (outer face); B, chela and carpus of right cheliped (outer face). Scale equals 1 mm." httpUri="https://zenodo.org/record/157219/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
B) appreciably shorter than left; with prominent hiatus between dactyl and fixed finger. Dactyl approximately 1.5 length of palm, distinctly overlapped by fixed finger; upper margin with 2 irregular rows of quite small spines; outer surface with irregular row of small spines and few additional scattered spinules; inner face with double row of small spines in proximal 0.6, not extending to tip; ventral surface unarmed; cutting edge with row of small calcareous teeth; all surfaces with moderately abundant, sparse tufts of long setae, not concealing armature. Palm slightly more than 0.5 length of carpus; upper margin with irregular double row of small spines; outer face with scattered small spines and spinules accompanied by sparse tufts of long setae; lower margin with row of small spines; inner face also with scattered spinules and setae. Carpus with irregular double row of small spines on upper margin partially obscured by long setae; outer face with weakly defined row of small spines and additional scattered spinules accompanied by sparse tufts of long setae; lower margin with few spinulose protuberances; inner surface also with scattered spinules, mesiodistal margin with row of small spines, extending onto lower distal margin as row of smaller spinules. Merus with row of low protuberances and tufts of long setae on dorsal margin; ventromesial margin with row of small spines and tufts of long setae; lateral face with few spinules in ventral 0.5, ventrolateral margin not distinctly delimited but with few spinules in proximal 0.5. Ischium with row of small spinules on ventromesial margin not concealed by sparse tufts of long setae.
</paragraph>
<paragraph id="8BEA36E6774E7A0BFEF6C6441A0BF8DA" blockId="4.[264,1324,284,1870]" pageId="4" pageNumber="5">
Second (
<figureCitation id="136E2A63774E7A0BFE6CC6441EC9F8B2" box="[418,489,1804,1830]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
B, C) and third pereopods with dactyls slightly to distinctly curved, longer than propodi; all surfaces, mesial faces particularly, with moderately long setae.
</paragraph>
<paragraph id="8BEA36E6774F7A0AFEC6C0541D50FE6A" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
Propodi unarmed but with setae dorsally and ventrally. Carpi of second pereopods each with row of quite small spines on dorsal margin; carpi of third pereopods each with row of very small spinules on dorsal margin. Meri each with low protuberances and setae on dorsal surface, ventral margins with fewer setae. Ischia unarmed, but with setae dorsally and ventrally. Sternite of third pereopods (
<figureCitation id="136E2A63774F7A0AFD03C0F41C2EFE42" box="[717,782,444,470]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
D) generally subquadrate, with median tuft of setae on either side of midline.
</paragraph>
<paragraph id="8BEA36E6774F7A0AFEF6C3441D59FD0A" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">Female with paired gonopores; unpaired left pleopods biramous; second with endopod 2­segmented, exopod incompletely 3­segmented; third and fourth with endopods also 2­ segmented, exopods incompletely 5­segmented; fifth pleopod with endopod vestigial, exopod incompletely 5­segmented.</paragraph>
<paragraph id="8BEA36E6774F7A0AFEF6C3E41C2BFC22" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
Telson with small median cleft; left lobe of 
<typeStatus id="54EE8844774F7A0AFCFBC3E41CBBFD52" box="[821,923,684,710]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
(
<figureCitation id="136E2A63774F7A0AFC65C3E41CCDFD52" box="[939,1005,684,710]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
E) with 4 moderately large spines on terminal margin, 1more elongate spine at outer angle and 4 large spines on posterior 0.5 of lateral margin; right lobe with 3 small and 3 larger spines on terminal margin, outer 2 actually on rounded outer angle. Telson of female (
<figureCitation id="136E2A63774F7A0AFC70C26C1B21FCAA" box="[958,1025,804,830]" captionStart="FIGURE 1" captionStartId="2.[264,367,1466,1490]" captionTargetBox="[266,1321,328,1465]" captionTargetId="figure@2.[264,1323,308,1471]" captionTargetPageId="2" captionText="FIGURE 1. Diogenes guttatus Henderson, 1888, A – D, F, female, sl = 2.7 mm, off Thursday Island (AM 67380); E, male, holotype sl = 3.1 mm, Challenger stn 187 (NHM 1888.33.1). A, shield and cephalic appendages (aesthetascs omitted); B, right second pereopod (lateral view); C, dactyl of right second pereopod (mesial view); D, anterior lobe of sternite of third pereopods; E, F, telson (E after Siddiqui &amp; McLaughlin, 2003). Scales equal 1 mm." httpUri="https://zenodo.org/record/157218/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
F) with row of small and large spines on terminal margin of left lobe extending nearly full length of lateral margin; right lobe with several small spines accompanied by few larger spines on terminal margin, reaching only onto rounded proximal angle.
</paragraph>
<paragraph id="8BEA36E6774F7A0AFEF6C28C1D69FC4A" blockId="5.[264,1325,284,1870]" box="[312,585,964,990]" pageId="5" pageNumber="6">
<emphasis id="B921EAF4774F7A0AFEF6C28C1E9FFC4A" bold="true" box="[312,447,964,990]" pageId="5" pageNumber="6">Coloration</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34F656D774F7A0AFEF6C2A41D02FB92" box="[312,546,1004,1030]" pageId="5" pageNumber="6" type="description">
<paragraph id="8BEA36E6774F7A0AFEF6C2A41D02FB92" blockId="5.[264,1325,284,1870]" box="[312,546,1004,1030]" pageId="5" pageNumber="6">
<emphasis id="B921EAF4774F7A0AFEF6C2A41EB8FB92" bold="true" box="[312,408,1004,1030]" pageId="5" pageNumber="6">Habitat</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34F656D774F7A0AFEF6C55C1D09FBC2" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BEA36E6774F7A0AFEF6C55C1D09FBC2" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
<emphasis id="B921EAF4774F7A0AFEF6C55C1EF1FBBA" bold="true" box="[312,465,1044,1070]" pageId="5" pageNumber="6">Distribution</emphasis>
. Off Cape York Peninsula, Queensland, 
<collectingCountry id="F3427676774F7A0AFC7AC55C1B03FBBA" box="[948,1059,1044,1070]" name="Australia" pageId="5" pageNumber="6">Australia</collectingCountry>
; off Phuket, 
<collectingCountry id="F3427676774F7A0AFB75C55C1A07FBBA" box="[1211,1319,1044,1070]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
, Andaman Sea; 
<quantity id="4CAD9B03774F7A0AFE75C5741D06FBC2" box="[443,550,1084,1110]" metricMagnitude="1" metricUnit="m" metricValue="3.6" metricValueMax="6.1" metricValueMin="1.1" pageId="5" pageNumber="6" unit="m" value="36.0" valueMax="61.0" valueMin="11.0">
11– 
<specimenCount id="9D53FD6F774F7A0AFE28C5741D06FBC2" box="[486,550,1084,1110]" pageId="5" pageNumber="6" type="generic">61 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F656D774F7A09FEF6C52C1C36FB32" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BEA36E6774F7A0AFEF6C52C1CF7FA2A" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
<emphasis id="B921EAF4774F7A0AFEF6C52C1E8AFBEA" bold="true" box="[312,426,1124,1150]" pageId="5" pageNumber="6">Remarks</emphasis>
. As indicated in the redescription, 
<taxonomicName id="4C554D65774F7A0AFC82C52C1CEFFBE9" box="[844,975,1124,1149]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="5" pageNumber="6" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774F7A0AFC82C52C1CEFFBE9" box="[844,975,1124,1149]" italics="true" pageId="5" pageNumber="6">D. guttatus</emphasis>
</taxonomicName>
, like several other species of the genus is subject to appreciable morphological variation in the strength of the armature of the chelipeds (e.g., 
<bibRefCitation id="EFC44B17774F7A0AFDC3C5FC1C33FB5A" author="Forest" box="[525,787,1204,1230]" pageId="5" pageNumber="7" refString="Forest, J. &amp; Guinot, D. (1956) Sur une collection de Crustaces Decapodes et Stomatopodes des mers tunisiennes. Bulletin Station Oceanographique Salammbo, 53, 24 - 44." type="journal article" year="1956">Forest &amp; Guinot 1956</bibRefCitation>
, 
<bibRefCitation id="EFC44B17774F7A0AFCEEC5FC1B0EFB5A" author="Rahayu" box="[800,1070,1204,1230]" pageId="5" pageNumber="8" refString="Rahayu, D. L. &amp; Forest, J. (1995) Le genre Diogenes (Decapoda, Anomura, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin du Museum national d'Histoire naturelle, (4) A, 16, 383 - 415. [Dated 1994]." type="journal article" year="1995">Rahayu &amp; Forest 1995</bibRefCitation>
, 
<bibRefCitation id="EFC44B17774F7A0AFBF5C5FC1A01FB5A" author="McLaughlin" box="[1083,1313,1204,1230]" pageId="5" pageNumber="8" refString="McLaughlin, P. A. (2002 a) Diogenes pallescens Whitelegge, D. gardineri Alcock and D. serenei Forest (Decapoda; Anomura; Diogenidae): morphological variants or distinct species? Raffles Bulletin of Zoology, 50, 81 - 94." type="journal article" year="2002" yearSuffix="a">McLaughlin 2002a</bibRefCitation>
). Whether the differences seen in this armature of the left cheliped between the female from off Thursday Island and the 
<typeStatus id="54EE8844774F7A0AFD9EC44C1D96FA8A" box="[592,694,1284,1310]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
indicates sexual dimorphism cannot be determined at present. Given the relatively similar sizes of the two specimens, neither can it be ruled out. The female reported here is considerably larger than either the male or female 
<collectingCountry id="F3427676774F7A0AFB69C41C1BFDFAFA" box="[1191,1245,1364,1390]" name="Thailand" pageId="5" pageNumber="6">Thai</collectingCountry>
specimens reported by 
<bibRefCitation id="EFC44B17774F7A0AFE2DC4341DFFFA02" author="McLaughlin" box="[483,735,1404,1430]" pageId="5" pageNumber="8" refString="McLaughlin, P. A. (2002 b) A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Crustacea of the Andaman Sea 29 November - 20 December, 1998, Phuket Marine Biological Center. Phuket Marine Biological Center Special Publication, 23, 385 - 460." type="journal article" year="2002" yearSuffix="b">McLaughlin (2002b)</bibRefCitation>
. Nonetheless, there is general agreement on the armature of the right chelipeds among those three specimens.
</paragraph>
<paragraph id="8BEA36E6774F7A0AFEF6C4841C33F8DA" blockId="5.[264,1325,284,1870]" pageId="5" pageNumber="6">
In spination of the pereopodal carpi, the female from off Thursday Island, like the 
<typeStatus id="54EE8844774F7A0AFEC6C4BC1E51F99A" box="[264,369,1524,1550]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
, has a row of spines or spinules on each pereopodal carpus, although they are slightly smaller in the female. This is in contrast to the two or three spines and few spinules reported by 
<bibRefCitation id="EFC44B17774F7A0AFDCAC70C1DDCF9CA" author="McLaughlin" box="[516,764,1604,1630]" pageId="5" pageNumber="8" refString="McLaughlin, P. A. (2002 b) A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Crustacea of the Andaman Sea 29 November - 20 December, 1998, Phuket Marine Biological Center. Phuket Marine Biological Center Special Publication, 23, 385 - 460." type="journal article" year="2002" yearSuffix="b">McLaughlin (2002b)</bibRefCitation>
for the 
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specimens. However, as demonstrated by 
<bibRefCitation id="EFC44B17774F7A0AFE4AC7241D85F912" author="Rahayu" box="[388,677,1644,1670]" pageId="5" pageNumber="8" refString="Rahayu, D. L. &amp; Forest, J. (1995) Le genre Diogenes (Decapoda, Anomura, Diogenidae) en Indonesie, avec la description de six especes nouvelles. Bulletin du Museum national d'Histoire naturelle, (4) A, 16, 383 - 415. [Dated 1994]." type="journal article" year="1995">Rahayu &amp; Forest (1995)</bibRefCitation>
and 
<bibRefCitation id="EFC44B17774F7A0AFD2EC7241B76F912" author="Siddiqui" box="[736,1110,1644,1670]" pageId="5" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin (2003)</bibRefCitation>
, variations in carpal spination are not uncommon, even between right and left or second and third pereopods of individuals from a single locality. The telsons described for the 
<collectingCountry id="F3427676774F7A0AFB68C7F41BFCF942" box="[1190,1244,1724,1750]" name="Thailand" pageId="5" pageNumber="6">Thai</collectingCountry>
specimens appear to be intermediate in spination between the 
<typeStatus id="54EE8844774F7A0AFC6BC7AC1B2BF96A" box="[933,1035,1764,1790]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
and the female from off Thursday Island, but both 
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representatives are very similar to the Australian female in the spines of the ocular and antennal acicles.
</paragraph>
<paragraph id="8BEA36E6774C7A09FEF6C0541D39FC82" blockId="6.[264,1324,284,1190]" pageId="6" pageNumber="7">
<bibRefCitation id="EFC44B17774C7A09FEF6C0541D0DFEA2" author="McLaughlin" box="[312,557,284,310]" pageId="6" pageNumber="8" refString="McLaughlin, P. A. (2002 b) A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Crustacea of the Andaman Sea 29 November - 20 December, 1998, Phuket Marine Biological Center. Phuket Marine Biological Center Special Publication, 23, 385 - 460." type="journal article" year="2002" yearSuffix="b">McLaughlin (2002b)</bibRefCitation>
remarked on the very distinctive antennal acicles seen in both 
<taxonomicName id="4C554D65774C7A09FAC0C0541E46FEC9" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FAC0C0541A03FEA1" box="[1294,1315,284,309]" italics="true" pageId="6" pageNumber="7">D</emphasis>
. 
<emphasis id="B921EAF4774C7A09FEC6C00C1E46FEC9" box="[264,358,324,349]" italics="true" pageId="6" pageNumber="7">guttatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C554D65774C7A09FE50C00C1D0AFEC9" box="[414,554,324,349]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="tirmiziae">
<emphasis id="B921EAF4774C7A09FE50C00C1D0AFEC9" box="[414,554,324,349]" italics="true" pageId="6" pageNumber="7">D. tirmiziae</emphasis>
</taxonomicName>
. The two species are also similar in the armature of the right cheliped; however, as pointed out by both 
<bibRefCitation id="EFC44B17774C7A09FD15C0241CF1FE12" author="McLaughlin" box="[731,977,364,390]" pageId="6" pageNumber="8" refString="McLaughlin, P. A. (2002 b) A review of the hermit crab (Decapoda: Anomura: Paguridea) fauna of southern Thailand, with particular emphasis on the Andaman Sea, and descriptions of three new species. In: Bruce, N., Berggren, M. &amp; Bussarawit, S. (Eds.), Proceedings of the International Workshop on the Crustacea of the Andaman Sea 29 November - 20 December, 1998, Phuket Marine Biological Center. Phuket Marine Biological Center Special Publication, 23, 385 - 460." type="journal article" year="2002" yearSuffix="b">McLaughlin (2002b)</bibRefCitation>
and 
<bibRefCitation id="EFC44B17774C7A09FBC3C0241E79FE3A" author="Siddiqui" pageId="6" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin (2003)</bibRefCitation>
, 
<taxonomicName id="4C554D65774C7A09FEAAC0DC1ED0FE39" box="[356,496,404,430]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="tirmiziae">
<emphasis id="B921EAF4774C7A09FEAAC0DC1E59FE39" box="[356,377,404,429]" italics="true" pageId="6" pageNumber="7">D</emphasis>
. 
<emphasis id="B921EAF4774C7A09FE46C0DC1ED0FE39" box="[392,496,404,429]" italics="true" pageId="6" pageNumber="7">tirmiziae</emphasis>
</taxonomicName>
can be differentiated by the presence of a row of spines on dorsal margin of each propodus and ventral margin of each merus of the second pereopods. Additionally, although McLaughlin did not describe the basal segments of the antennular peduncles of the 
<collectingCountry id="F3427676774C7A09FE1BC3441D2BFDB2" box="[469,523,524,550]" name="Thailand" pageId="6" pageNumber="7">Thai</collectingCountry>
specimens, she did note the row of spines on the lateral margin of the second segment of each antennal peduncle. 
<bibRefCitation id="EFC44B17774C7A09FC8FC37C1B92FDDA" author="Siddiqui" box="[833,1202,564,590]" pageId="6" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin (2003)</bibRefCitation>
described this margin in 
<taxonomicName id="4C554D65774C7A09FE7DC3141D1FFDE1" box="[435,575,604,629]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="tirmiziae">
<emphasis id="B921EAF4774C7A09FE7DC3141D1FFDE1" box="[435,575,604,629]" italics="true" pageId="6" pageNumber="7">D. tirmiziae</emphasis>
</taxonomicName>
as having only three to five spines. At present, the mushroomshaped tubercles of the palm of the left cheliped of 
<taxonomicName id="4C554D65774C7A09FC4BC3CC1B2AFD09" box="[901,1034,644,670]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FC4BC3CC1CBAFD09" box="[901,922,644,669]" italics="true" pageId="6" pageNumber="7">D</emphasis>
. 
<emphasis id="B921EAF4774C7A09FC62C3CC1B2AFD09" box="[940,1034,644,669]" italics="true" pageId="6" pageNumber="7">guttatus</emphasis>
</taxonomicName>
distinguish this species from all others in the genus. Considering the similarities seen between the female of 
<taxonomicName id="4C554D65774C7A09FAC0C3E41E46FD79" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FAC0C3E41E46FD79" italics="true" pageId="6" pageNumber="7">D. guttatus</emphasis>
</taxonomicName>
from off Thursday Island and the two 
<collectingCountry id="F3427676774C7A09FC8EC39C1C56FD7A" box="[832,886,724,750]" name="Thailand" pageId="6" pageNumber="7">Thai</collectingCountry>
specimens, there can be little doubt that all are conspecific.
</paragraph>
<paragraph id="8BEA36E6774C7A09FEF6C26C1C36FB32" blockId="6.[264,1324,284,1190]" pageId="6" pageNumber="7">
<bibRefCitation id="EFC44B17774C7A09FEF6C26C1D85FCAA" author="Siddiqui" box="[312,677,804,830]" pageId="6" pageNumber="8" refString="Siddiqui, F. A. &amp; McLaughlin, P. A. (2003) A new species of the hermit crab genus Diogenes (Decapoda: Anomura: Paguroidea: Diogenidae) from Pakistan, with a comparative diagnosis of D. guttatus Henderson, 1888. Proceedings of the Biological Society of Washington, 116, 956 - 966." type="journal article" year="2003">Siddiqui &amp; McLaughlin (2003)</bibRefCitation>
also discussed the similarities between 
<taxonomicName id="4C554D65774C7A09FBBAC26C1A20FCA9" box="[1140,1280,804,829]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="tirmiziae">
<emphasis id="B921EAF4774C7A09FBBAC26C1A20FCA9" box="[1140,1280,804,829]" italics="true" pageId="6" pageNumber="7">D. tirmiziae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C554D65774C7A09FAC0C26C1E46FCF1" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FAC0C26C1A03FCA9" box="[1294,1315,804,829]" italics="true" pageId="6" pageNumber="7">D</emphasis>
. 
<emphasis id="B921EAF4774C7A09FEC6C2041E46FCF1" box="[264,358,844,869]" italics="true" pageId="6" pageNumber="7">guttatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C554D65774C7A09FE6FC2041C77FCF2" authority="Morgan &amp; Forest 1991" authorityName="Morgan &amp; Forest" authorityYear="1991" box="[417,855,844,870]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="dorotheae">
<emphasis id="B921EAF4774C7A09FE6FC2041D1CFCF1" box="[417,572,844,869]" italics="true" pageId="6" pageNumber="7">D. dorotheae</emphasis>
<bibRefCitation id="EFC44B17774C7A09FD8AC2041C77FCF2" author="Morgan" box="[580,855,844,870]" pageId="6" pageNumber="8" refString="Morgan, G. J. &amp; Forest, J. (1991) Seven new species of hermit crabs from northern and western Australia (Decapoda, Anomura, Diogenidae). Bulletin du Museum national d'Histoire naturelle, (4) A, 12, 649 - 689." type="journal article" year="1991">Morgan &amp; Forest 1991</bibRefCitation>
</taxonomicName>
. The latter species is known only from its unique and quite small 
<typeStatus id="54EE8844774C7A09FDF2C23C1D86FC1A" box="[572,678,884,910]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
, collected west of Cape Bossut, northern Western 
<collectingCountry id="F3427676774C7A09FB3CC23C1E68FC22" name="Australia" pageId="6" pageNumber="7">Australia</collectingCountry>
. 
<bibRefCitation id="EFC44B17774C7A09FE94C2D41D5CFC22" author="Morgan" box="[346,636,924,950]" pageId="6" pageNumber="8" refString="Morgan, G. J. &amp; Forest, J. (1991) Seven new species of hermit crabs from northern and western Australia (Decapoda, Anomura, Diogenidae). Bulletin du Museum national d'Histoire naturelle, (4) A, 12, 649 - 689." type="journal article" year="1991">Morgan &amp; Forest (1991)</bibRefCitation>
distinguished 
<taxonomicName id="4C554D65774C7A09FCE8C2D41CE0FC21" box="[806,960,924,949]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="dorotheae">
<emphasis id="B921EAF4774C7A09FCE8C2D41CE0FC21" box="[806,960,924,949]" italics="true" pageId="6" pageNumber="7">D. dorotheae</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C554D65774C7A09FBC6C2D41BAAFC21" box="[1032,1162,924,949]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FBC6C2D41BAAFC21" box="[1032,1162,924,949]" italics="true" pageId="6" pageNumber="7">D. guttatus</emphasis>
</taxonomicName>
by the mushroom­shaped tubercles of the latter’s left chela. 
<taxonomicName id="4C554D65774C7A09FCA1C28C1B41FC49" box="[879,1121,964,989]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="dorotheae">
<emphasis id="B921EAF4774C7A09FCA1C28C1B41FC49" box="[879,1121,964,989]" italics="true" pageId="6" pageNumber="7">Diogenes dorotheae</emphasis>
</taxonomicName>
shares with the 
<typeStatus id="54EE8844774C7A09FEC6C2A41E4EFB92" box="[264,366,1004,1030]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C554D65774C7A09FE59C2A41D3AFB91" box="[407,538,1004,1029]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FE59C2A41D3AFB91" box="[407,538,1004,1029]" italics="true" pageId="6" pageNumber="7">D. guttatus</emphasis>
</taxonomicName>
the prominent spines at the lower proximal angle of the left palm, and with the Australian female, the general armature of the telson. However, 
<bibRefCitation id="EFC44B17774C7A09FB62C55C1E89FBC2" author="Morgan" pageId="6" pageNumber="8" refString="Morgan, G. J. &amp; Forest, J. (1991) Seven new species of hermit crabs from northern and western Australia (Decapoda, Anomura, Diogenidae). Bulletin du Museum national d'Histoire naturelle, (4) A, 12, 649 - 689." type="journal article" year="1991">Morgan &amp; Forest (1991)</bibRefCitation>
described the basal segment of the antennular peduncle as being unarmed. This segment in 
<taxonomicName id="4C554D65774C7A09FE1AC52C1D79FBE9" box="[468,601,1124,1149]" class="Chlorophyceae" family="Coccomyxaceae" genus="Diogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Chlamydomonadales" pageId="6" pageNumber="7" phylum="Chlorophyta" rank="species" species="guttatus">
<emphasis id="B921EAF4774C7A09FE1AC52C1D79FBE9" box="[468,601,1124,1149]" italics="true" pageId="6" pageNumber="7">D. guttatus</emphasis>
</taxonomicName>
is provided with a distolateral row of spines that continues uninterruptedly onto the ventrodistal margin.
</paragraph>
</subSubSection>
</treatment>
</document>