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<document id="598855671073F5A0E15CC5EB6DD4EA6B" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFA7F76401B015964FE2C9AD" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada dubitabile Köhler 2010, n.sp." docType="treatment" docUuid="8BBD9360-B48A-485F-ABD2-9649567681B9" docUuidSource="ZooBank" docVersion="3" lastPageNumber="273" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="271" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="7DC263219827F155B2E1A96FFD7FF36E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E7583845FD4008A2CB966FE32E84693F">
<mods:title id="FB807C37DD763470D2BFD21DB32A0F1B">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="D06F4212AC41994ECC1F65A0852E82E0" type="personal">
<mods:role id="7B457A22607ED17B66A3FDACC179A7E9">
<mods:roleTerm id="F1DE48DCB4F087EDB0093B7EB071D9D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1581195EE6613F2904E541FE79605D12">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="8A523FB49B92E6FCD7B62327949B8F82">text</mods:typeOfResource>
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<mods:title id="8F4F70DA266D4A828A494A5CDD605537">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="6F868965244D9EDCD5401D5F19876FA8">
<mods:date id="86281BF189975C4F037F5427BCDAA2FB">2010</mods:date>
<mods:detail id="72C7E5854220DE55ED8C625D9071F68E" type="pubDate">
<mods:number id="68A85C2B3BC05D0F9342DA77434D94AD">2010-11-24</mods:number>
</mods:detail>
<mods:detail id="342826CA761496A671362CEF9F02D850" type="volume">
<mods:number id="C78AF450C92B1E65376F17FC70F73DE4">62</mods:number>
</mods:detail>
<mods:detail id="A3BAAD0473BFBF3BB4896C956864ABC9" type="issue">
<mods:number id="B8D4CEEC3ACABF5143BAC7F236B1AB79">2</mods:number>
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<mods:start id="F2E6EBE06AF2BCE644CD229FAD3F7F12">217</mods:start>
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<mods:location id="F5AD12153563DDCA20D83C3A093ACDB7">
<mods:url id="3BDA362B9D34A180A3928049EB047F00">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
</mods:location>
<mods:classification id="37139651FFAD316DBA8F273398890A6E">journal article</mods:classification>
<mods:identifier id="7BC605AE3D71E359B1F5EE6DF28F6978" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="87A6C83B56DF9C119BA97AE3567205CD" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="FEAB3FDE21BDEC99E7EF66E253AFE043" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="2468D6729F8DA88F6BC51F97E3404BA3" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FFA7F76401B015964FE2C9AD" LSID="urn:lsid:zoobank.org:act:8BBD9360-B48A-485F-ABD2-9649567681B9" httpUri="http://treatment.plazi.org/id/03F987C5FFA7F76401B015964FE2C9AD" lastPageId="56" lastPageNumber="273" pageId="54" pageNumber="271">
<subSubSection id="C34A6558FFA7F76A01B015964828CB98" box="[965,1341,1490,1517]" pageId="54" pageNumber="271" type="nomenclature">
<paragraph id="8BEF36D3FFA7F76A01B015964828CB98" blockId="54.[965,1341,1490,1517]" box="[965,1341,1490,1517]" pageId="54" pageNumber="271">
<heading id="D0A781BFFFA7F76A01B015964828CB98" box="[965,1341,1490,1517]" centered="true" fontSize="11" level="2" pageId="54" pageNumber="271" reason="2">
<emphasis id="B924EAC1FFA7F76A01B015964828CB98" bold="true" box="[965,1341,1490,1517]" pageId="54" pageNumber="271">
<taxonomicName id="4C504D50FFA7F76A01B0159649ECCB99" ID-CoL="D7QS" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[965,1273,1491,1517]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="54" pageNumber="271" phylum="Mollusca" rank="species" species="dubitabile" status="sp. nov.">
<emphasis id="B924EAC1FFA7F76A01B0159649ECCB99" bold="true" box="[965,1273,1491,1517]" italics="true" pageId="54" pageNumber="271">Amplirhagada dubitabile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA7F76A077415974828CB98" box="[1281,1341,1490,1516]" pageId="54" pageNumber="271" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFA7F76A0132164A4805C897" pageId="54" pageNumber="271" type="materials_examined">
<paragraph id="8BEF36D3FFA7F76A0132164A481CC80A" blockId="54.[839,1467,1551,1662]" pageId="54" pageNumber="271">
<materialsCitation id="3B383C8EFFA7F76A0132164A4810C80A" collectingDate="1988-07-08" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="Kimberley" latitude="-16.06" location="Doubtful Bay" longLatPrecision="21" longitude="124.3675" municipality="Bonaparte Archipelago" pageId="54" pageNumber="271" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFA7F76A0132164A4EF3C853" bold="true" box="[839,998,1551,1575]" pageId="54" pageNumber="271">
<typeStatus id="54EB8871FFA7F76A0132164A4E96C853" box="[839,899,1551,1575]" pageId="54" pageNumber="271">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFA7F76A018F164A4954C852" box="[1018,1089,1551,1574]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="54" pageNumber="271">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFA7F76A0628164A483BC852" box="[1117,1326,1551,1574]" country="Australia" name="Western Australia" pageId="54" pageNumber="271">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFA7F76A0748164A48A2C852" box="[1341,1463,1551,1574]" pageId="54" pageNumber="271">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFA7F76A013216684955C830" box="[839,1088,1581,1604]" pageId="54" pageNumber="271">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FFA7F76A063C166849CFC830" LSID="urn:lsid:plazi:treatment:03F987C5FFA7F76401B015964FE2C9AD:8E8F6008FFA7F76A063C166849CFC830" box="[1097,1242,1581,1604]" county="Kimberley" latitude="-16.06" longLatPrecision="21" longitude="124.3675" municipality="Bonaparte Archipelago" name="Doubtful Bay" pageId="54" pageNumber="271" stateProvince="Western Australia">Doubtful Bay</location>
,
<location id="8E8F6008FFA7F76A06911668487DC830" LSID="urn:lsid:plazi:treatment:03F987C5FFA7F76401B015964FE2C9AD:8E8F6008FFA7F76A06911668487DC830" box="[1252,1384,1581,1604]" county="Kimberley" latitude="-16.06" longLatPrecision="21" longitude="124.3675" municipality="Bonaparte Archipelago" name="Steep Island" pageId="54" pageNumber="271" stateProvince="Western Australia">Steep Island</location>
,
<locationDeviation id="9A5D2FF1FFA7F76A070716684EECC815" location="Raft Point" pageId="54" pageNumber="271">
<quantity id="4CA89B36FFA7F76A0707166848AFC830" box="[1394,1466,1581,1604]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="54" pageNumber="271" unit="km" value="2.3">2.3 km</quantity>
NE of Raft Point
</locationDeviation>
,
<geoCoordinate id="EE645014FFA7F76A0677160F496EC815" box="[1026,1147,1610,1633]" degrees="16" direction="south" minutes="03" orientation="latitude" pageId="54" pageNumber="271" precision="15" seconds="36" value="-16.06">16°03'36&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFA7F76A06F4160F4819C815" box="[1153,1292,1610,1633]" degrees="124" direction="east" minutes="22" orientation="longitude" pageId="54" pageNumber="271" precision="15" seconds="03" value="124.3675">124°22'03&quot;E</geoCoordinate>
; KC-002 (leg.
<collectionCode id="ED41AE16FFA7F76A07D1160F48A2C815" box="[1444,1463,1610,1633]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="54" pageNumber="271" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FFA7F76A013216224E89C80A" box="[839,924,1639,1662]" pageId="54" pageNumber="271">Kessner</collectorName>
&amp;
<collectorName id="26A55305FFA7F76A01C816224970C80A" box="[957,1125,1639,1662]" pageId="54" pageNumber="271">A. Longbottom</collectorName>
,
<date id="FFEE1013FFA7F76A061A162249E9C80A" box="[1135,1276,1639,1662]" pageId="54" pageNumber="271" value="1988-07-08">
<collectingDate id="EFAAE9FBFFA7F76A061A162249E9C80A" box="[1135,1276,1639,1662]" pageId="54" pageNumber="271" value="1988-07-08">08 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFA7F76A013216E54805C897" blockId="54.[839,1467,1696,1763]" pageId="54" pageNumber="271">
<emphasis id="B924EAC1FFA7F76A013216E54EC5C8C7" bold="true" box="[839,976,1696,1715]" pageId="54" pageNumber="271">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFA7F76A01AA16E54834C8C7" accessionNumber="S34627" box="[991,1313,1696,1715]" collectionCode="WAM" pageId="54" pageNumber="271" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FFA7F76A01AA16E54926C8C7" box="[991,1075,1696,1715]" pageId="54" pageNumber="271" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFA7F76A064E16E54964C8C7" box="[1083,1137,1696,1715]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="54" pageNumber="271" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA7F76A060C16E549A8C8C7" box="[1145,1213,1696,1715]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34627" pageId="54" pageNumber="271" type="EnaNcbi">S34627</accessionNumber>
(Pl. 2.15)
</materialsCitation>
.
<materialsCitation id="3B383C8EFFA7F76A075916E54819C897" accessionNumber="S41478" collectionCode="WAM" pageId="54" pageNumber="271" specimenCode="FMNH 219047" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EB8871FFA7F76A075916E54896C8C7" box="[1324,1411,1696,1715]" pageId="54" pageNumber="271" type="paratype">Paratypes</typeStatus>
AMS C463762 (3 preserved specimens),
<specimenCode id="DBF69EA8FFA7F76A06D316FD482DC8BF" box="[1190,1336,1720,1739]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="54" pageNumber="271" type="Museum">FMNH 219047</specimenCode>
(8 preserved specimens),
<collectionCode id="ED41AE16FFA7F76A01C116954EFCC897" box="[948,1001,1744,1763]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="54" pageNumber="271" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA7F76A019B16954925C897" box="[1006,1072,1744,1763]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41478" pageId="54" pageNumber="271" type="EnaNcbi">S41478</accessionNumber>
(5 preserved specimens)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFA7F764013217424FE2C9AD" lastPageId="56" lastPageNumber="273" pageId="54" pageNumber="271" type="etymology">
<paragraph id="8BEF36D3FFA7F76A0132174248AEC948" blockId="54.[839,1467,1799,1852]" pageId="54" pageNumber="271">
<emphasis id="B924EAC1FFA7F76A013217424ED4C96B" bold="true" box="[839,961,1799,1823]" pageId="54" pageNumber="271">Etymology</emphasis>
. From
<taxonomicName id="4C504D50FFA7F76A067F17424960C96A" authorityName="Köhler" authorityYear="2010" box="[1034,1141,1799,1822]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="54" pageNumber="271" phylum="Mollusca" rank="species" species="dubitabile">
<emphasis id="B924EAC1FFA7F76A067F17424960C96A" box="[1034,1141,1799,1822]" italics="true" pageId="54" pageNumber="271">dubitabile</emphasis>
</taxonomicName>
(Latin = doubtful), in reference to Doubtful Bay, where this species occurs on Steep Island.
</paragraph>
<paragraph id="8BEF36D3FFA7F76B0132171A4CDEC867" blockId="54.[839,1467,1887,1998]" lastBlockId="55.[121,749,1238,1555]" lastPageId="55" lastPageNumber="272" pageId="54" pageNumber="271">
<emphasis id="B924EAC1FFA7F76A0132171A4E94C903" bold="true" box="[839,897,1887,1911]" pageId="54" pageNumber="271">Shell</emphasis>
(
<figureCitation id="136B2A56FFA7F76A01E0171A490AC902" box="[917,1055,1887,1910]" captionStart="Figure 85" captionStartId="55.[121,184,1089,1110]" captionTargetBox="[121,1421,151,1077]" captionTargetId="figure-9@55.[121,1421,151,1078]" captionTargetPageId="55" captionText="Figure 85. SEM photographs of shell, jaw and radula of Amplirhagada dubitabile n.sp., paratype FMNH 219047:(A) apical whorl viewed from above (scale 200 µm); (B) sculpture on penultimate whorl viewed rectangular from above (scale 200 µm); (C) close-up showing growth lines on last whorl (scale 200 µm); (D) jaw (scale 100 µm); (E) central and inner lateral teeth (scale 20 µm); (F) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239277" httpUri="https://zenodo.org/record/5239277/files/figure.png" pageId="54" pageNumber="271">Fig. 85A–C</figureCitation>
, Pl. 2.15). Semi-globose to broadly conical, with medium high spire; solid. Periphery slightly angulate; upper and basal sectors of whorls rounded. Umbilicus open, forming narrowly winding opening, 30–70 percent concealed by columellar reflection. Background colour brownish beige, may display irregular pattern of hornish growth lines; sub-sutural band absent; mid-whorl band absent or diffuse, yellowish brown, thin, visible on last whorl only; ventral colour horn; outer lip colour differs from shell, horn; inner lip whitish. Protoconch c.
<quantity id="4CA89B36FFA6F76B00EC152C4FF9CBF4" box="[665,748,1385,1408]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="55" pageNumber="272" unit="mm" value="2.8">2.8 mm</quantity>
in diameter, comprising 2 whorls, smooth. Teleoconch with pronounced axial growth lines. Angle of aperture 45°; outer lip rounded, moderately thick, slightly expanded, not or slightly reflected, basal and palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<caption id="DF2F665BFFA6F76B020C14044CDCCAD2" ID-DOI="http://doi.org/10.5281/zenodo.5239277" ID-Zenodo-Dep="5239277" httpUri="https://zenodo.org/record/5239277/files/figure.png" pageId="55" pageNumber="272" startId="55.[121,184,1089,1110]" targetBox="[121,1421,151,1077]" targetPageId="55">
<paragraph id="8BEF36D3FFA6F76B020C14044CDCCAD2" blockId="55.[121,1421,1089,1190]" pageId="55" pageNumber="272">
Figure 85. SEM photographs of shell, jaw and radula of
<taxonomicName id="4C504D50FFA6F76B00FC14044E6CCA22" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[649,889,1089,1110]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="272" phylum="Mollusca" rank="species" species="dubitabile" status="sp. nov.">
<emphasis id="B924EAC1FFA6F76B00FC14044E6CCA22" box="[649,889,1089,1110]" italics="true" pageId="55" pageNumber="272">Amplirhagada dubitabile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA6F76B010B14044EB8CA22" box="[894,941,1089,1110]" pageId="55" pageNumber="272" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219047: (
<emphasis id="B924EAC1FFA6F76B06C7140449D4CA22" box="[1202,1217,1089,1110]" italics="true" pageId="55" pageNumber="272">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFA6F76B03E014194CB1CA05" box="[405,420,1116,1137]" italics="true" pageId="55" pageNumber="272">B</emphasis>
) sculpture on penultimate whorl viewed rectangular from above (scale 200 µm); (
<emphasis id="B924EAC1FFA6F76B06B0141949C0CA05" box="[1221,1237,1116,1137]" italics="true" pageId="55" pageNumber="272">C</emphasis>
) close-up showing growth lines on last whorl (scale 200 µm); (
<emphasis id="B924EAC1FFA6F76B006E14334F39CAFF" box="[539,556,1142,1163]" italics="true" pageId="55" pageNumber="272">D</emphasis>
) jaw (scale 100 µm); (
<emphasis id="B924EAC1FFA6F76B017014334E01CAFF" box="[773,788,1142,1163]" italics="true" pageId="55" pageNumber="272">E</emphasis>
) central and inner lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FFA6F76B06A0143349F1CAFF" box="[1237,1252,1142,1163]" italics="true" pageId="55" pageNumber="272">F</emphasis>
) outer lateral and inner marginal teeth (scale 20 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFA6F76B016F14934977CB81" blockId="55.[794,1422,1238,1525]" pageId="55" pageNumber="272">Vagina rather short, posteriorly inflated; inner vaginal wall with weakly developed longitudinal pilasters. Spermatheca slightly extending over base of spermoviduct. Spermathecal duct wide, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising about half of anterior part of oviduct, more or less straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end of albumen gland.</paragraph>
<paragraph id="8BEF36D3FFA6F76B020C16734F12C8FC" blockId="55.[121,749,1590,1672]" pageId="55" pageNumber="272">
<emphasis id="B924EAC1FFA6F76B020C16734C41C83A" bold="true" box="[121,340,1590,1614]" pageId="55" pageNumber="272">Pallial morphology</emphasis>
. Pallial cavity short, extending half a whorl; mantle pigmentation mottled, greyish black. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFA6F764020C16EE4F8EC95D" blockId="55.[121,749,1707,1995]" lastBlockId="56.[166,794,1575,1833]" lastPageId="56" lastPageNumber="273" pageId="55" pageNumber="272">
<emphasis id="B924EAC1FFA6F76B020C16EE4C77C8B7" bold="true" box="[121,354,1707,1731]" pageId="55" pageNumber="272">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFA6F76B030116E94CEEC8B7" box="[372,507,1708,1731]" captionStart-0="Figure 86" captionStart-1="Figure 87" captionStartId-0="55.[794,857,1921,1942]" captionStartId-1="56.[166,229,1414,1435]" captionTargetBox-0="[799,1411,1556,1911]" captionTargetBox-1="[186,764,157,1404]" captionTargetId-0="figure-94@55.[794,1421,1542,1923]" captionTargetId-1="figure-9@56.[166,793,148,1410]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionText-0="Figure 86. Genitalia of Amplirhagada dubitabile n.sp., paratype FMNH 219047 (8 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 87. Interior of penial chamber of Amplirhagada dubitabile n.sp., paratype FMNH 219047 (8 July, scale 3 mm). Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239279" figureDoi-1="http://doi.org/10.5281/zenodo.5239281" httpUri-0="https://zenodo.org/record/5239279/files/figure.png" httpUri-1="https://zenodo.org/record/5239281/files/figure.png" pageId="55" pageNumber="272">Figs. 86–87</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens coils once before entering penis. Penial retractor muscle shorter than penis. Penial verge short (&lt;
<emphasis id="B924EAC1FFA6F76B00F917414F81C966" bold="true" box="[652,660,1796,1810]" pageId="55" pageNumber="272">
<superScript id="7C259B9BFFA6F76B00F917414F81C966" attach="none" box="[652,660,1796,1810]" fontSize="6" pageId="55" pageNumber="272">1</superScript>
</emphasis>
⁄
<emphasis id="B924EAC1FFA6F76B00EC174E4FB4C96D" bold="true" box="[665,673,1803,1817]" pageId="55" pageNumber="272">
<subScript id="17D43496FFA6F76B00EC174E4FB4C96D" attach="left" box="[665,673,1803,1817]" fontSize="6" pageId="55" pageNumber="272">8</subScript>
</emphasis>
penial chamber), broad, pointed. Penial wall pustules absent; wall covered by delicate, narrow, longitudinal pilasters. Main stimulatory pilaster well-differentiated; smooth undulating, well developed at apical portion, decreasing in size in median portion, largely reduced at basal portion of penial chamber. Vas deferens entering penial sheath in upper third.
<emphasis id="B924EAC1FFA9F76402D316624C88C84B" bold="true" box="[166,413,1575,1599]" pageId="56" pageNumber="273">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFA9F76403DA166D4F27C84B" box="[431,562,1576,1599]" captionStart="Figure 85" captionStartId="55.[121,184,1089,1110]" captionTargetBox="[121,1421,151,1077]" captionTargetId="figure-9@55.[121,1421,151,1078]" captionTargetPageId="55" captionText="Figure 85. SEM photographs of shell, jaw and radula of Amplirhagada dubitabile n.sp., paratype FMNH 219047:(A) apical whorl viewed from above (scale 200 µm); (B) sculpture on penultimate whorl viewed rectangular from above (scale 200 µm); (C) close-up showing growth lines on last whorl (scale 200 µm); (D) jaw (scale 100 µm); (E) central and inner lateral teeth (scale 20 µm); (F) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239277" httpUri="https://zenodo.org/record/5239277/files/figure.png" pageId="56" pageNumber="273">Fig. 85E–F</figureCitation>
). Rectangular. Tooth formula C+16+3+22. With 139 rows of teeth, 32.3 rows per mm (n = 1). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular to ovate mesocones, length equal to base of tooth; ectocones small; endocones vestigial. Marginal teeth with elongate to triangular mesocones; ectocones shorter and narrower than mesocones; endocones smaller than ectocones.
</paragraph>
<caption id="DF2F665BFFA6F76B016F17C44EE7C9B8" ID-DOI="http://doi.org/10.5281/zenodo.5239279" ID-Zenodo-Dep="5239279" httpUri="https://zenodo.org/record/5239279/files/figure.png" pageId="55" pageNumber="272" startId="55.[794,857,1921,1942]" targetBox="[799,1411,1556,1911]" targetPageId="55">
<paragraph id="8BEF36D3FFA6F76B016F17C44EE7C9B8" blockId="55.[794,1422,1921,1996]" pageId="55" pageNumber="272">
Figure 86. Genitalia of
<taxonomicName id="4C504D50FFA6F76B067617C749E2C9E3" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1027,1271,1922,1943]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="55" pageNumber="272" phylum="Mollusca" rank="species" species="dubitabile" status="sp. nov.">
<emphasis id="B924EAC1FFA6F76B067617C749E2C9E3" box="[1027,1271,1922,1943]" italics="true" pageId="55" pageNumber="272">Amplirhagada dubitabile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA6F76B068A17C4483AC9E2" box="[1279,1327,1921,1942]" pageId="55" pageNumber="272" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219047 (8 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFA9F76402D315C34CCACBA5" ID-DOI="http://doi.org/10.5281/zenodo.5239281" ID-Zenodo-Dep="5239281" httpUri="https://zenodo.org/record/5239281/files/figure.png" pageId="56" pageNumber="273" startId="56.[166,229,1414,1435]" targetBox="[186,764,157,1404]" targetPageId="56">
<paragraph id="8BEF36D3FFA9F76402D315C34CCACBA5" blockId="56.[166,794,1414,1489]" pageId="56" pageNumber="273">
Figure 87. Interior of penial chamber of
<taxonomicName id="4C504D50FFA9F764005D15C24E0CCBE8" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[552,793,1415,1436]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="273" phylum="Mollusca" rank="species" species="dubitabile" status="sp. nov.">
<emphasis id="B924EAC1FFA9F764005D15C24E0CCBE8" box="[552,793,1415,1436]" italics="true" pageId="56" pageNumber="273">Amplirhagada dubitabile</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA9F76402D315E44DC3CBC2" box="[166,214,1441,1462]" pageId="56" pageNumber="273" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219047 (8 July, scale 3 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FFA9F76402D317084FE2C9AD" blockId="56.[166,794,1869,2009]" pageId="56" pageNumber="273">
<emphasis id="B924EAC1FFA9F76402D317084CB1C911" bold="true" box="[166,420,1869,1893]" pageId="56" pageNumber="273">Comparative remarks</emphasis>
. Material of this species has been labelled as “
<taxonomicName id="4C504D50FFA9F764035E172E4CFFC9F5" box="[299,490,1898,1922]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="273" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FFA9F764035E172E4CD0C9F6" box="[299,453,1899,1922]" italics="true" pageId="56" pageNumber="273">Amplirhagada</emphasis>
sp.
</taxonomicName>
63” by Solem. Shell similar to
<taxonomicName id="4C504D50FFA9F76402B217CD4C97C9EB" authorityName="Köhler" authorityYear="2010" box="[199,386,1928,1951]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="273" phylum="Mollusca" rank="species" species="kimberleyana">
<emphasis id="B924EAC1FFA9F76402B217CD4C97C9EB" box="[199,386,1928,1951]" italics="true" pageId="56" pageNumber="273">A. kimberleyana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FFA9F76403B417CD4F23C9EB" authorityName="Köhler" authorityYear="2010" box="[449,566,1928,1951]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="56" pageNumber="273" phylum="Mollusca" rank="species" species="gemina">
<emphasis id="B924EAC1FFA9F76403B417CD4F23C9EB" box="[449,566,1928,1951]" italics="true" pageId="56" pageNumber="273">A. gemina</emphasis>
</taxonomicName>
in shape, size, and colouration. Inner penial wall structure (lamellae, no pustules with long, undulating main pilaster) very characteristic.
</paragraph>
</subSubSection>
</treatment>
</document>
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