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<document id="49D8A2829F08173A14079CE1E2DC677F" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF92F758036516DB49ABC9A5" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada euroa Köhler 2010, n.sp." docType="treatment" docUuid="284664D6-0AEB-4201-B4CC-DEE14524AA3E" docUuidSource="ZooBank" docVersion="3" lastPageNumber="221" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="220" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="29BAD99DA01FC8FF9300CF2638C494A4">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="4FFE89B59CF660798C87B3423403D869">Köhler, Frank</mods:namePart>
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<mods:title id="0252061AC04064ADF60CD0E645F91BA9">Records of the Australian Museum</mods:title>
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<mods:date id="717C4E13969C37EF9FCD4B51A557BC65">2010</mods:date>
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<mods:number id="A1B3CA039BBA6C997A2CA974646B7703">2010-11-24</mods:number>
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<mods:url id="2083D84F493E7D397DF0304B26A859E0">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<subSubSection id="C34A6558FF92F75F036516DB4F40C8CC" box="[272,597,1694,1720]" pageId="3" pageNumber="220" type="nomenclature">
<paragraph id="8BEF36D3FF92F75F036516DB4F40C8CC" blockId="3.[272,597,1694,1720]" box="[272,597,1694,1720]" pageId="3" pageNumber="220">
<heading id="D0A781BFFF92F75F036516DB4F40C8CC" box="[272,597,1694,1720]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="220" reason="2">
<emphasis id="B924EAC1FF92F75F036516DB4F40C8CC" bold="true" box="[272,597,1694,1720]" pageId="3" pageNumber="220">
<taxonomicName id="4C504D50FF92F75F036516DB4F04C8CC" ID-CoL="D7QW" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[272,529,1694,1720]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="220" phylum="Mollusca" rank="species" species="euroa" status="sp. nov.">
<emphasis id="B924EAC1FF92F75F036516DB4F04C8CC" bold="true" box="[272,529,1694,1720]" italics="true" pageId="3" pageNumber="220">Amplirhagada euroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF92F75F006C16DB4F40C8CC" box="[537,597,1694,1720]" pageId="3" pageNumber="220" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BEF36D3FF92F75F020C169D4CB6C9F6" blockId="3.[121,749,1752,1922]" pageId="3" pageNumber="220">
<materialsCitation id="3B383C8EFF92F75F020C169D4CB5C9F6" collectingDate="2008-08-07" collectionCode="V" county="eastern Kimberley" latitude="-15.10889" location="Adolphus Island" longLatPrecision="21" longitude="128.15222" municipality="Cambridge Gulf" pageId="3" pageNumber="220" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF92F75F020C169D4C1DC884" bold="true" box="[121,264,1752,1776]" pageId="3" pageNumber="220">
<typeStatus id="54EB8871FF92F75F020C169D4DA4C884" box="[121,177,1752,1776]" pageId="3" pageNumber="220">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF92F75F0361169D4C5AC89B" box="[276,335,1752,1775]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="3" pageNumber="220">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF92F75F0311169D4F35C89B" box="[356,544,1752,1775]" country="Australia" name="Western Australia" pageId="3" pageNumber="220">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF92F75F005D169D4FFDC89B" box="[552,744,1752,1775]" pageId="3" pageNumber="220">eastern Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF92F75F020C16B34C2EC979" box="[121,315,1782,1805]" pageId="3" pageNumber="220">Cambridge Gulf</collectingMunicipality>
, eastern section of
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,
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<geoCoordinate id="EE645014FF92F75F028317564C6AC95E" box="[246,383,1811,1834]" degrees="128" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="220" precision="15" seconds="08" value="128.15222">128°09'08&quot;E</geoCoordinate>
; KIS 3113.
<location id="8E8F6008FF92F75F007017564F68C95E" LSID="urn:lsid:plazi:treatment:03F987C5FF92F758036516DB49ABC9A5:8E8F6008FF92F75F007017564F68C95E" box="[517,637,1811,1834]" county="eastern Kimberley" latitude="-15.10889" longLatPrecision="21" longitude="128.15222" municipality="Cambridge Gulf" name="East-facing" pageId="3" pageNumber="220" stateProvince="Western Australia">East-facing</location>
gully with ephemeral stream and patches of vine thicket, fig trees.
<location id="8E8F6008FF92F75F00C717754FF8C933" LSID="urn:lsid:plazi:treatment:03F987C5FF92F758036516DB49ABC9A5:8E8F6008FF92F75F00C717754FF8C933" box="[690,749,1840,1863]" county="eastern Kimberley" latitude="-15.10889" longLatPrecision="21" longitude="128.15222" municipality="Cambridge Gulf" name="Scree" pageId="3" pageNumber="220" stateProvince="Western Australia">Scree</location>
on very steep slopes.
<location id="8E8F6008FF92F75F032C170B4C7AC911" LSID="urn:lsid:plazi:treatment:03F987C5FF92F758036516DB49ABC9A5:8E8F6008FF92F75F032C170B4C7AC911" box="[345,367,1870,1893]" county="eastern Kimberley" latitude="-15.10889" longLatPrecision="21" longitude="128.15222" municipality="Cambridge Gulf" name="In" pageId="3" pageNumber="220" stateProvince="Western Australia">In</location>
loose soil under large boulders (leg.
<collectionCode id="ED41AE16FF92F75F020C172E4D99C9F6" box="[121,140,1899,1922]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="220" type="Museum">V</collectionCode>
. Kessner,
<date id="FFEE1013FF92F75F028D172E4C8DC9F6" box="[248,408,1899,1922]" pageId="3" pageNumber="220" value="2008-08-07">
<collectingDate id="EFAAE9FBFF92F75F028D172E4C8DC9F6" box="[248,408,1899,1922]" pageId="3" pageNumber="220" value="2008-08-07">7 August 2008</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF92F75F020C17E14FF9C9BB" blockId="3.[121,748,1956,1999]" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F020C17E14C11C9C3" bold="true" box="[121,260,1956,1975]" pageId="3" pageNumber="220">Type material</emphasis>
.
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<typeStatus id="54EB8871FF92F75F036117E14C7CC9C3" box="[276,361,1956,1975]" pageId="3" pageNumber="220" type="holotype">Holotype</typeStatus>
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(Pl. 1.1)
</materialsCitation>
.
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<typeStatus id="54EB8871FF92F75F002917E14FA0C9C3" box="[604,693,1956,1975]" pageId="3" pageNumber="220" type="paratype">Paratypes</typeStatus>
AMS C463680 (6 preserved specimens, as
<typeStatus id="54EB8871FF92F75F03AE17F94F3AC9BB" box="[475,559,1980,1999]" pageId="3" pageNumber="220" type="holotype">holotype</typeStatus>
),
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(12
</paragraph>
<paragraph id="8BEF36D3FF92F75F016F10DF492CCF79" blockId="3.[794,1422,154,269]" pageId="3" pageNumber="220">
preserved specimens, as
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<typeStatus id="54EB8871FF92F75F018610DF4954CED9" box="[1011,1089,154,173]" pageId="3" pageNumber="220" type="holotype">holotype</typeStatus>
),
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(3 shells,
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)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF92F75F01D510F74846CEB1" accessionNumber="S37391" box="[928,1363,178,197]" collectionCode="WAM" isTailEnumeration="true" latitude="-15.109722" longLatPrecision="21" longitude="128.16583" pageId="3" pageNumber="220" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED41AE16FF92F75F01D510F74EC0CEB1" box="[928,981,178,197]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="220" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF92F75F01AB10F74937CEB1" box="[990,1058,178,197]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37391" pageId="3" pageNumber="220" type="EnaNcbi">S37391</accessionNumber>
(shell,
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)
</materialsCitation>
, AMS C463723 (6 shells,
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),
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<collectionCode id="ED41AE16FF92F75F06DA108F49F6CEA9" box="[1199,1251,202,221]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="220" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF92F75F069F108F4839CEA9" box="[1258,1324,202,221]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37392" pageId="3" pageNumber="220" type="EnaNcbi">S37392</accessionNumber>
(11 shells,
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<geoCoordinate id="EE645014FF92F75F01F010A74EE3CE81" box="[901,1014,226,245]" degrees="128" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="220" precision="15" seconds="08" value="128.15222">128°09'08&quot;E</geoCoordinate>
)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF92F75F067D10A74920CF79" accessionNumber="S37393" collectionCode="WAM" latitude="-15.071944" location="Adolphus Island" longLatPrecision="21" longitude="128.13834" pageId="3" pageNumber="220" specimenCount="1" typeStatus="holotype">
<collectionCode id="ED41AE16FF92F75F067D10A74928CE81" box="[1032,1085,226,245]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="3" pageNumber="220" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF92F75F063010A74992CE81" box="[1093,1159,226,245]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S37393" pageId="3" pageNumber="220" type="EnaNcbi">S37393</accessionNumber>
(3 shells, north of
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,
<geoCoordinate id="EE645014FF92F75F012C10BF4EAFCF79" box="[857,954,250,269]" degrees="15" direction="south" minutes="04" orientation="latitude" pageId="3" pageNumber="220" precision="15" seconds="19" value="-15.071944">15°04'19&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF92F75F01CA10BF493BCF79" box="[959,1070,250,269]" degrees="128" direction="east" minutes="08" orientation="longitude" pageId="3" pageNumber="220" precision="15" seconds="18" value="128.13834">128°08'18&quot;E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF92F758016F117449ABC9A5" lastPageId="4" lastPageNumber="221" pageId="3" pageNumber="220" type="etymology">
<paragraph id="8BEF36D3FF92F75F016F11744E92CFF7" blockId="3.[794,1421,305,387]" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F016F11744E80CF3D" bold="true" box="[794,917,305,329]" pageId="3" pageNumber="220">Etymology</emphasis>
. From
<taxonomicName id="4C504D50FF92F75F019111744934CF3C" authorityName="Köhler" authorityYear="2010" box="[996,1057,305,328]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="220" phylum="Mollusca" rank="species" species="euroa">
<emphasis id="B924EAC1FF92F75F019111744934CF3C" box="[996,1057,305,328]" italics="true" pageId="3" pageNumber="220">euroa</emphasis>
</taxonomicName>
(Latin = eastern), for this species occurring at the easternmost limits of the known range of the genus.
</paragraph>
<paragraph id="8BEF36D3FF92F75F016F11E34946CFC9" blockId="3.[794,1107,422,445]" box="[794,1107,422,445]" pageId="3" pageNumber="220">Sealing strategy. Rock sealer.</paragraph>
<paragraph id="8BEF36D3FF92F75F016F11A44997CA51" blockId="3.[794,1422,481,1061]" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F016F11A44E47CF8D" bold="true" box="[794,850,481,505]" pageId="3" pageNumber="220">Shell</emphasis>
(
<figureCitation id="136B2A56FF92F75F011711A44ECDCF8C" box="[866,984,481,504]" captionStart="Figure 2" captionStartId="4.[166,229,1565,1586]" captionTargetBox="[167,1467,151,1549]" captionTargetId="figure-85@4.[167,1467,151,1549]" captionTargetPageId="4" captionText="Figure 2. SEM photographs of Amplirhagada euroa n.sp., paratype AMS C463680: (A) apical whorl viewed from above (scale 200 µm); (B) penultimate whorls viewed from above (scale 200 µm); (C) central and lateral radular teeth (scale 20 µm); (D) details of central and lateral teeth (scale 10 µm); (E) details of outer lateral and inner marginal teeth (scale 10 µm); (F) jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239090" httpUri="https://zenodo.org/record/5239090/files/figure.png" pageId="3" pageNumber="220">Fig. 2AB</figureCitation>
, Pl. 1.12). Broadly conical with low to medium high spire, thin to translucent. Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus open, narrowly winding, 5090 percent concealed by columellar reflection. Background colour yellowish brown with chestnut brown, thin to moderately broad, diffuse to well defined sub-sutural and mid-whorl bands, clearly visible on last whorls only; ventral colour brownish horn to whitish; outer lip colour tends to be lighter than shell till whitish; inner lip translucent. Protoconch c.
<quantity id="4CA89B36FF92F75F01CA13434914CD69" box="[959,1025,774,797]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="220" unit="mm" value="3.0">3 mm</quantity>
in diameter, comprising about 1.7 whorls, with faint, indistinct axial riblets. Teleoconch with fine axial lirae, curved if viewed from above, pointed in cross-section, irregularly spaced, spaces as wide as thickness of lirae. Lirae evenly distributed across whorl diameter, with reduced height underneath suture; present on all whorls. Angle of aperture 45°. Outer lip simple rounded, sharp, slightly to well expanded, slightly reflected; basal node absent or very weak; palatal node absent. Parietal wall of inner lip absent or inconspicuous.
</paragraph>
<paragraph id="8BEF36D3FF92F75F016F140C49BDCAEF" blockId="3.[794,1421,1097,1179]" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F016F140C4EE9CA15" bold="true" box="[794,1020,1097,1121]" pageId="3" pageNumber="220">Pallial morphology</emphasis>
. Pallial cavity deep, extending one whorl; mottled or spotted black mantle pigmentation. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FF92F75F016F14FB4EABC977" blockId="3.[794,1422,1214,1795]" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F016F14FB4917CAA2" bold="true" box="[794,1026,1214,1238]" pageId="3" pageNumber="220">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF92F75F066614FB496BCAA1" box="[1043,1150,1214,1237]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="5.[121,183,676,697]" captionStartId-1="5.[794,857,1297,1318]" captionTargetBox-0="[124,736,153,647]" captionTargetBox-1="[817,1389,140,1272]" captionTargetId-0="figure-9@5.[121,748,140,655]" captionTargetId-1="figure-67@5.[794,1421,126,1291]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Figure 3. Genitalia of Amplirhagada euroa n.sp., paratype AMS C463680 (7 August, scale 10 mm). Labelling of structures: ag, albumen gland; hd, hermaphroditic duct; p, penial complex (penis and penis sheath); ov, oviduct (free anterior part); rm, penial retractor muscle; sd, spermathecal duct; sh, spermathecal head; sp, spermoviduct (uterus, prostate); t, talon; va,vagina; vd, vas deferens." captionText-1="Figure 4. Interior of penial chamber of Amplirhagada euroa n.sp., paratype AMS C463680 (7 August, scale 5 mm). Labelling of structures: iw, inner penial wall; lp, longitudinal (main) pilaster; rm, penial retractor muscle; sh, penial sheath; vd, vas deferens; vg, penial verge." figureDoi-0="http://doi.org/10.5281/zenodo.5239092" figureDoi-1="http://doi.org/10.5281/zenodo.5239094" httpUri-0="https://zenodo.org/record/5239092/files/figure.png" httpUri-1="https://zenodo.org/record/5239094/files/figure.png" pageId="3" pageNumber="220">Figs. 34</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens coils before entering penis. Penial retractor muscle shorter than penis complex. Penial verge short, less than
<emphasis id="B924EAC1FF92F75F07231553484BCB50" bold="true" box="[1366,1374,1302,1316]" pageId="3" pageNumber="220">
<superScript id="7C259B9BFF92F75F07231553484BCB50" attach="right" box="[1366,1374,1302,1316]" fontSize="6" pageId="3" pageNumber="220">1</superScript>
</emphasis>
<emphasis id="B924EAC1FF92F75F071115584879CB5F" bold="true" box="[1380,1388,1309,1323]" pageId="3" pageNumber="220">
<subScript id="17D43496FF92F75F071115584879CB5F" attach="left" box="[1380,1388,1309,1323]" fontSize="6" pageId="3" pageNumber="220">5</subScript>
</emphasis>
of length of penial chamber, slender with rounded tip. Penial wall pustules normal to elongated, arranged in rows over entire length of inner penial wall. Main stimulatory pilaster large, cone-shaped, comprising entire length of inner penial wall; sculptured by ridges with smooth, flattened pustules. Vas deferens entering penial sheath in upper third. Vagina of medium length, posteriorly inflated; inner vaginal wall supports undulating longitudinal pilasters. Spermatheca short, reaching base of spermoviduct; duct wide, inner wall with smooth longitudinal pilasters; head globular to elongately inflated, connected with oviduct by connective tissue, wall of head delicate, smooth. Free oviduct rather straight comprising about half of length of anterior part of oviduct. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end of albumen gland.
</paragraph>
<paragraph id="8BEF36D3FF92F758016F17634E04C955" blockId="3.[794,1422,1830,2000]" lastBlockId="4.[166,794,1743,1825]" lastPageId="4" lastPageNumber="221" pageId="3" pageNumber="220">
<emphasis id="B924EAC1FF92F75F016F17634901C94A" bold="true" box="[794,1044,1830,1854]" pageId="3" pageNumber="220">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FF92F75F0652176349B7C949" box="[1063,1186,1830,1853]" captionStart="Figure 2" captionStartId="4.[166,229,1565,1586]" captionTargetBox="[167,1467,151,1549]" captionTargetId="figure-85@4.[167,1467,151,1549]" captionTargetPageId="4" captionText="Figure 2. SEM photographs of Amplirhagada euroa n.sp., paratype AMS C463680: (A) apical whorl viewed from above (scale 200 µm); (B) penultimate whorls viewed from above (scale 200 µm); (C) central and lateral radular teeth (scale 20 µm); (D) details of central and lateral teeth (scale 10 µm); (E) details of outer lateral and inner marginal teeth (scale 10 µm); (F) jaw (scale 100 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239090" httpUri="https://zenodo.org/record/5239090/files/figure.png" pageId="3" pageNumber="220">Fig. 2CE</figureCitation>
). Rectangular. Tooth formula C+1213+34+2021. Average number of rows of teeth 146±7 with 36.2±0.7 rows of teeth per mm (n = 2). Central teeth with sharply pointed triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with bluntly pointed triangular mesocones, length equal to base of tooth; small ectocones; endocones absent. Marginal teeth multicuspic, mesocone and endocone similar in length, ectocone smaller than endocone, occasionally subdivided.
</paragraph>
<caption id="DF2F665BFF95F75802D31658481FC813" ID-DOI="http://doi.org/10.5281/zenodo.5239090" ID-Zenodo-Dep="5239090" httpUri="https://zenodo.org/record/5239090/files/figure.png" pageId="4" pageNumber="221" startId="4.[166,229,1565,1586]" targetBox="[167,1467,151,1549]" targetPageId="4">
<paragraph id="8BEF36D3FF95F75802D31658481FC813" blockId="4.[166,1467,1565,1640]" pageId="4" pageNumber="221">
Figure 2. SEM photographs of
<taxonomicName id="4C504D50FF95F75803BA16584F83C846" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[463,662,1565,1586]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="euroa" status="sp. nov.">
<emphasis id="B924EAC1FF95F75803BA16584F83C846" box="[463,662,1565,1586]" italics="true" pageId="4" pageNumber="221">Amplirhagada euroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF95F75800EE16584FDEC846" box="[667,715,1565,1586]" pageId="4" pageNumber="221" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463680: (
<emphasis id="B924EAC1FF95F75801A416584EF5C846" box="[977,992,1565,1586]" italics="true" pageId="4" pageNumber="221">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF95F75802DB167D4DA8C839" box="[174,189,1592,1613]" italics="true" pageId="4" pageNumber="221">B</emphasis>
) penultimate whorls viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF95F758009E167D4FEEC839" box="[747,763,1592,1613]" italics="true" pageId="4" pageNumber="221">C</emphasis>
) central and lateral radular teeth (scale 20 µm); (
<emphasis id="B924EAC1FF95F75806A7167D49F6C839" box="[1234,1251,1592,1613]" italics="true" pageId="4" pageNumber="221">D</emphasis>
) details of central and lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FF95F75803C116164CD6C81C" box="[436,451,1619,1640]" italics="true" pageId="4" pageNumber="221">E</emphasis>
) details of outer lateral and inner marginal teeth (scale 10 µm); (
<emphasis id="B924EAC1FF95F75806451616492AC81C" box="[1072,1087,1619,1640]" italics="true" pageId="4" pageNumber="221">F</emphasis>
) jaw (scale 100 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FF95F75802D3170149ABC9A5" blockId="4.[166,794,1860,2001]" lastBlockId="4.[839,1467,1743,2001]" pageId="4" pageNumber="221">
<emphasis id="B924EAC1FF95F75802D317014C89C928" bold="true" box="[166,412,1860,1884]" pageId="4" pageNumber="221">Comparative remarks</emphasis>
.
<taxonomicName id="4C504D50FF95F75803DD17004F94C928" authorityName="Köhler" authorityYear="2010" box="[424,641,1861,1884]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="euroa">
<emphasis id="B924EAC1FF95F75803DD17004F94C928" box="[424,641,1861,1884]" italics="true" pageId="4" pageNumber="221">Amplirhagada euroa</emphasis>
</taxonomicName>
is geographically well separated from most other congeneric species.
<taxonomicName id="4C504D50FF95F75800A117274F31C9E2" authority="Solem, 1988" authorityName="Solem" authorityYear="1988" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="cambridgensis">
<emphasis id="B924EAC1FF95F75800A117274C89C9E3" italics="true" pageId="4" pageNumber="221">Amplirhagada cambridgensis</emphasis>
<bibRefCitation id="EFC14B22FF95F75803D4173A4F31C9E2" author="Solem, A" box="[417,548,1919,1942]" pageId="4" pageNumber="221" pagination="27 - 58" refId="ref39380" refString="Solem, A., 1988. New camaenid land snails from the northeast Kimberley, Western Australia. Journal of the Malacological Society of Australia 9: 27 - 58." type="journal article" year="1988">Solem, 1988</bibRefCitation>
</taxonomicName>
(from the western bank of Cambridge Sound) and
<taxonomicName id="4C504D50FF95F75803C517D84FCFC9C0" authority="Solem, 1981 a" authorityName="Solem" authorityYear="1981" box="[432,730,1949,1972]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="questronana">
<emphasis id="B924EAC1FF95F75803C517D84F5EC9C0" box="[432,587,1949,1972]" italics="true" pageId="4" pageNumber="221">A. questronana</emphasis>
<bibRefCitation id="EFC14B22FF95F758003B17D84FCFC9C0" author="Solem, A" box="[590,730,1949,1972]" pageId="4" pageNumber="221" pagination="147 - 320" refId="ref39183" refString="Solem, A., 1981 a. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Supplement 11: 147 - 320." type="journal article" year="1981">Solem, 1981a</bibRefCitation>
</taxonomicName>
(from El Questro Station near Wyndham, c.
<quantity id="4CA89B36FF95F758005A17FF4F6AC9A5" box="[559,639,1978,2001]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="221" unit="km" value="100.0">100 km</quantity>
S of Adolphus Island) occur in closer proximity. The shell of
<taxonomicName id="4C504D50FF95F758074516954E6EC970" authorityName="Solem" authorityYear="1988" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="cambridgensis">
<emphasis id="B924EAC1FF95F758074516954E6EC970" italics="true" pageId="4" pageNumber="221">A. cambridgensis</emphasis>
</taxonomicName>
is very similar but this species differs in the morphology of the inner penial wall with pustules being arranged to form corrugated longitudinal pilasters.
<taxonomicName id="4C504D50FF95F75806E8176D4861C94B" authorityName="Köhler" authorityYear="2010" box="[1181,1396,1832,1855]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="euroa">
<emphasis id="B924EAC1FF95F75806E8176D4861C94B" box="[1181,1396,1832,1855]" italics="true" pageId="4" pageNumber="221">Amplirhagada euroa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C504D50FF95F75801F717004937C928" authorityName="Solem" authorityYear="1981" box="[898,1058,1861,1884]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="questronana">
<emphasis id="B924EAC1FF95F75801F717004937C928" box="[898,1058,1861,1884]" italics="true" pageId="4" pageNumber="221">A. questronana</emphasis>
</taxonomicName>
by umbilicus forming a chink instead of being open and by its long main stimulatory pilaster that supports flattened pustules (
<taxonomicName id="4C504D50FF95F758061417C549E9C9E3" authorityName="Solem" authorityYear="1981" box="[1121,1276,1920,1943]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="221" phylum="Mollusca" rank="species" species="questronana">
<emphasis id="B924EAC1FF95F758061417C549E9C9E3" box="[1121,1276,1920,1943]" italics="true" pageId="4" pageNumber="221">A. questronana</emphasis>
</taxonomicName>
has a short pilaster with corrugations). Otherwise, both species have rather similar shells with regard to shape and size.
</paragraph>
</subSubSection>
</treatment>
</document>