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209 lines
22 KiB
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<document id="FDB03AC118FEC328BAFE95335C8DF9B2" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. & Marusik, Yuri M." docDate="2021" docId="03F98785037DEF631C5E7B05FCEE6AA9" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Berinda Roewer 1928" docType="treatment" docVersion="7" lastPageNumber="70" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="70" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="37F617B6E722ED23C8A3259FD048CE76">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="04F5579D886510B6B7950C6CEBDA68E1">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="DD85B521389AC3C4B3689E80BBB5C9FB">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="540E50CD5713C89DA6A4258CB438C78D" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="43DC93F1F359A82E1F19D10F70C7D4F6">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="4564746555D9C2DCD84C7DD3B4CF3D63">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="D71631AC4D4A2BFF1A0B51926B9F5A43">Sergei.</mods:affiliation>
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<mods:title id="0C3A3AA3E882D116F128BC8F38BE287F">European Journal of Taxonomy</mods:title>
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<treatment id="03F98785037DEF631C5E7B05FCEE6AA9" ID-DOI="http://doi.org/10.5281/zenodo.4906015" ID-GBIF-Taxon="182597824" ID-Zenodo-Dep="4906015" LSID="urn:lsid:plazi:treatment:03F98785037DEF631C5E7B05FCEE6AA9" httpUri="http://treatment.plazi.org/id/03F98785037DEF631C5E7B05FCEE6AA9" lastPageNumber="70" pageId="2" pageNumber="70">
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Genus
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<taxonomicName id="4C504D10037DEF631C897B05FC0869D1" ID-CoL="39PB" authority="Roewer, 1928" authorityName="Roewer" authorityYear="1928" box="[701,969,1142,1169]" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA81037DEF631C897B05FCDE69D0" bold="true" box="[701,799,1142,1169]" italics="true" pageId="2" pageNumber="70">Berinda</emphasis>
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<bibRefCitation id="EFC14B62037DEF631D127B04FC0869D1" author="Roewer C. F." box="[806,969,1142,1169]" pageId="2" pageNumber="70" pagination="92 - 123" refId="ref18119" refString="Roewer C. F. 1928. Araneae. In: Zoologische Streifzuge in Attika, Morea, und besonders auf der Insel Kreta, II. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 27: 92 - 123." type="journal article" year="1928">Roewer, 1928</bibRefCitation>
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<paragraph id="8BEF3693037DEF631E897BCBFDC86993" blockId="2.[189,521,1208,1235]" box="[189,521,1208,1235]" pageId="2" pageNumber="70">
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<treatmentCitation id="0AF11082037DEF631E897BCBFDC46993" author="Roewer C. F." box="[189,517,1208,1235]" page="106" pageId="2" pageNumber="70" year="1928">
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<taxonomicName id="4C504D10037DEF631E897BCBFDC46993" ID-CoL="39PB" authority="Roewer, 1928: 106" authorityName="Roewer" authorityPageNumber="106" authorityYear="1928" box="[189,517,1208,1235]" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA81037DEF631E897BCBFEDD6993" box="[189,284,1208,1234]" italics="true" pageId="2" pageNumber="70">Berinda</emphasis>
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<bibRefCitation id="EFC14B62037DEF631F177BCAFDC46993" author="Roewer C. F." box="[291,517,1208,1235]" pageId="2" pageNumber="70" pagination="92 - 123" refId="ref18119" refString="Roewer C. F. 1928. Araneae. In: Zoologische Streifzuge in Attika, Morea, und besonders auf der Insel Kreta, II. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 27: 92 - 123." type="journal article" year="1928">Roewer, 1928: 106</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8BEF3693037DEF631E897B89FCB76854" blockId="2.[189,886,1274,1301]" box="[189,886,1274,1301]" pageId="2" pageNumber="70">
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<emphasis id="B924EA81037DEF631E897B89FEDD6855" box="[189,284,1274,1300]" italics="true" pageId="2" pageNumber="70">Berinda</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0AF11082037DEF631F0D7B89FDCB6854" author="Murphy J." box="[313,522,1274,1301]" page="37" pageId="2" pageNumber="70" year="2007">
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<bibRefCitation id="EFC14B62037DEF631F0D7B89FDCB6854" author="Murphy J." box="[313,522,1274,1301]" pageId="2" pageNumber="70" refId="ref17893" refString="Murphy J. 2007. Gnaphosid Genera of the World. British Arachnological Society St Neots, Cambridgeshire." type="book" year="2007">Murphy 2007: 37</bibRefCitation>
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</treatmentCitation>
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. —
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<treatmentCitation id="0AF11082037DEF631C0D7B89FCB36854" ID-DOI="http://doi.org/10.5281/zenodo.6204142" ID-Zenodo-Dep="6204142" author="Panayiotou E. & Kaltsas D. & Seyyar O. & Chatzaki M." box="[569,882,1274,1301]" httpUri="http://treatment.plazi.org/id/EF4909198464FFA25D96188AFB5DF851" page="45" pageId="2" pageNumber="70" year="2010">
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<bibRefCitation id="EFC14B62037DEF631C0D7B89FCB36854" author="Panayiotou E. & Kaltsas D. & Seyyar O. & Chatzaki M." box="[569,882,1274,1301]" pageId="2" pageNumber="70" pagination="44 - 54" refId="ref18061" refString="Panayiotou E., Kaltsas D., Seyyar O. & Chatzaki M. 2010. Revision of the genus Berinda (Araneae, Gnaphosidae) in the east Mediterranean with the description of two new species. Zootaxa 2362: 44 - 54. https: // doi. org / 10.11646 / zootaxa. 2362.1.2" type="journal article" year="2010">
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||
Panayiotou
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||
<emphasis id="B924EA81037DEF631CF17B88FCC16855" box="[709,768,1274,1300]" italics="true" pageId="2" pageNumber="70">et al.</emphasis>
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2010: 45
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</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6518037DEF631E897A4EFDFB68C7" pageId="2" pageNumber="70" type="type_taxon">
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<paragraph id="8BEF3693037DEF631E897A4EFE996817" blockId="2.[189,344,1340,1367]" box="[189,344,1340,1367]" pageId="2" pageNumber="70">
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<heading id="D0A781FF037DEF631E897A4EFE996817" bold="true" box="[189,344,1340,1367]" fontSize="11" level="3" pageId="2" pageNumber="70" reason="3">
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<emphasis id="B924EA81037DEF631E897A4EFE996817" bold="true" box="[189,344,1340,1367]" pageId="2" pageNumber="70">
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<typeStatus id="54EB8831037DEF631E897A4EFF3B6816" box="[189,250,1341,1367]" pageId="2" pageNumber="70">Type</typeStatus>
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species
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</emphasis>
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||
</heading>
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</paragraph>
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||
<paragraph id="8BEF3693037DEF631E897A1FFDFB68C7" blockId="2.[189,570,1388,1414]" box="[189,570,1388,1414]" pageId="2" pageNumber="70">
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<taxonomicName id="4C504D10037DEF631E897A1FFDF768C7" ID-CoL="68K5C" authority="Roewer, 1928" authorityName="Roewer" authorityYear="1928" box="[189,566,1388,1414]" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="species" species="amabilis">
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||
<emphasis id="B924EA81037DEF631E897A1FFE4968C7" box="[189,392,1388,1414]" italics="true" pageId="2" pageNumber="70">Berinda amabilis</emphasis>
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<bibRefCitation id="EFC14B62037DEF631FBB7A1FFDF768C7" author="Roewer C. F." box="[399,566,1388,1414]" pageId="2" pageNumber="70" pagination="92 - 123" refId="ref18119" refString="Roewer C. F. 1928. Araneae. In: Zoologische Streifzuge in Attika, Morea, und besonders auf der Insel Kreta, II. Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 27: 92 - 123." type="journal article" year="1928">Roewer, 1928</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A6518037DEF631E897ADDFCEE6AA9" pageId="2" pageNumber="70" type="discussion">
|
||
<paragraph id="8BEF3693037DEF631E897ADDFE856889" blockId="2.[189,324,1454,1480]" box="[189,324,1454,1480]" pageId="2" pageNumber="70">
|
||
<heading id="D0A781FF037DEF631E897ADDFE856889" bold="true" box="[189,324,1454,1480]" fontSize="11" level="3" pageId="2" pageNumber="70" reason="3">
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||
<emphasis id="B924EA81037DEF631E897ADDFE856889" bold="true" box="[189,324,1454,1480]" pageId="2" pageNumber="70">Comments</emphasis>
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||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF3693037DEF631E897AAEFD736BEA" blockId="2.[189,1400,1501,1707]" pageId="2" pageNumber="70">
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||
This is a small genus with seven named species that are restricted to the Eastern Mediterranean from
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||
<collectingCountry id="F3477603037DEF631E8A7972FED26B5A" box="[190,275,1537,1563]" name="Greece" pageId="2" pageNumber="70">Greece</collectingCountry>
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||
to
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||
<collectingCountry id="F3477603037DEF631F0F7972FE676B5A" box="[315,422,1537,1563]" name="Turkey" pageId="2" pageNumber="70">Anatolia</collectingCountry>
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||
. The new species collected in
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<collectingCountry id="F3477603037DEF631D2E7971FC8B6B5D" box="[794,842,1538,1564]" name="Iran" pageId="2" pageNumber="70">Iran</collectingCountry>
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represent the southeasternmost records of the genus. Compared to other gnaphosids, homology of the male palpal structures in this genus is unclear, as it is questionable whether the sclerotized hook-like structure (“posterior hook” sensu
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<bibRefCitation id="EFC14B62037DEF631A81793AFF386BC6" author="Panayiotou E. & Kaltsas D. & Seyyar O. & Chatzaki M." pageId="2" pageNumber="70" pagination="44 - 54" refId="ref18061" refString="Panayiotou E., Kaltsas D., Seyyar O. & Chatzaki M. 2010. Revision of the genus Berinda (Araneae, Gnaphosidae) in the east Mediterranean with the description of two new species. Zootaxa 2362: 44 - 54. https: // doi. org / 10.11646 / zootaxa. 2362.1.2" type="journal article" year="2010">
|
||
Panayiotou
|
||
<emphasis id="B924EA81037DEF631B087939FAB46B22" box="[1340,1397,1609,1635]" italics="true" pageId="2" pageNumber="70">et al.</emphasis>
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||
2010
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||
</bibRefCitation>
|
||
) that guides the tip of the embolus is a greatly modified median apophysis or not. Here, we treat the hook-like structure as a median apophysis.
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</paragraph>
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||
<paragraph id="8BEF3693037DEF631E8979A0FCEE6AA9" blockId="2.[188,1400,1747,2025]" pageId="2" pageNumber="70">
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||
While comparing our specimens to the species reported from Europe, we realized that
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||
<taxonomicName id="4C504D10037DEF631AE879A0FE9C6A50" authority="Lissner, 2016" authorityName="Lissner" authorityYear="2016" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="species" species="idae">
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<emphasis id="B924EA81037DEF631AE879A0FAB96BAC" box="[1244,1400,1747,1773]" italics="true" pageId="2" pageNumber="70">Berinda idae</emphasis>
|
||
<bibRefCitation id="EFC14B62037DEF631E8B7984FE9C6A50" author="Lissner J. & Chatzaki M." box="[191,349,1783,1809]" pageId="2" pageNumber="70" pagination="9 - 11" refId="ref17628" refString="Lissner J. & Chatzaki M. 2016. A new species of Berinda (Araneae, Gnaphosidae) from the eastern Aegean Islands, Greece. Arachnologische Mitteilungen 51: 9 - 11. https: // doi. org / 10.5431 / aramit 5102" type="journal article" year="2016">Lissner, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
(described from
|
||
<collectingCountry id="F3477603037DEF631C107984FDB66A50" box="[548,631,1783,1809]" name="Greece" pageId="2" pageNumber="70">Greece</collectingCountry>
|
||
and later reported from
|
||
<collectingCountry id="F3477603037DEF631DA77984FC2B6A50" box="[915,1002,1783,1809]" name="Cyprus" pageId="2" pageNumber="70">Cyprus</collectingCountry>
|
||
) is undoubtedly a junior synonym of
|
||
<taxonomicName id="4C504D10037DEF631EE8786FFCC46A74" authority="(O. Pickard-Cambridge, 1872)" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[220,773,1819,1845]" class="Arachnida" family="Gnaphosidae" genus="Heser" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="species" species="infumatus">
|
||
<emphasis id="B924EA81037DEF631EE8786FFE5D6A74" box="[220,412,1819,1845]" italics="true" pageId="2" pageNumber="70">Heser infumatus</emphasis>
|
||
(O. Pickard-Cambridge, 1872)
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="F3477603037DEF631D607868FC6B6A74" box="[852,938,1819,1845]" name="Greece" pageId="2" pageNumber="70">Greece</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3477603037DEF631D807868FBE06A74" box="[948,1057,1819,1845]" name="Tanzania" pageId="2" pageNumber="70">Tanzania</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3477603037DEF631A1F7868FBB06A74" box="[1067,1137,1819,1845]" name="Egypt" pageId="2" pageNumber="70">Egypt</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3477603037DEF631A9C7868FB2C6A74" box="[1192,1261,1819,1845]" name="Israel" pageId="2" pageNumber="70">Israel</collectingCountry>
|
||
; introduced to
|
||
<collectingCountry id="F3477603037DEF631EE9784CFEE46A18" box="[221,293,1855,1881]" name="Japan" pageId="2" pageNumber="70">Japan</collectingCountry>
|
||
). The copulatory organs are identical (cf.
|
||
<bibRefCitation id="EFC14B62037DEF631D23784CFB876A18" author="Lissner J. & Chatzaki M." box="[791,1094,1855,1881]" pageId="2" pageNumber="70" pagination="9 - 11" refId="ref17628" refString="Lissner J. & Chatzaki M. 2016. A new species of Berinda (Araneae, Gnaphosidae) from the eastern Aegean Islands, Greece. Arachnologische Mitteilungen 51: 9 - 11. https: // doi. org / 10.5431 / aramit 5102" type="journal article" year="2016">Lissner & Chatzaki 2016</bibRefCitation>
|
||
: figs 2–3 and
|
||
<bibRefCitation id="EFC14B62037DEF631ADA784CFAB26A18" author="Levy G." box="[1262,1395,1855,1881]" pageId="2" pageNumber="70" pagination="93 - 158" refId="ref17565" refString="Levy G. 1998. The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology 44: 93 - 158." type="journal article" year="1998">Levy 1998</bibRefCitation>
|
||
: figs 112–115) and both have been recorded from Greek islands. The long filiform embolus, robust conductor and distinctive tibial apophysis conform to
|
||
<taxonomicName id="4C504D10037DEF631D6778F5FC726AE1" authorityName="Roewer" authorityYear="1928" box="[851,947,1926,1952]" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924EA81037DEF631D6778F5FC726AE1" box="[851,947,1926,1952]" italics="true" pageId="2" pageNumber="70">Berinda</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C504D10037DEF631A7D78F4FAFC6AE0" authority="Tuneva, 2004" authorityName="Tuneva" authorityYear="2004" box="[1097,1341,1927,1953]" class="Arachnida" family="Gnaphosidae" genus="Heser" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924EA81037DEF631A7D78F4FB516AE1" box="[1097,1168,1927,1952]" italics="true" pageId="2" pageNumber="70">Heser</emphasis>
|
||
Tuneva, 2004
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFC14B62037DEF631E8B78D9FE0B6A84" author="Panayiotou E. & Kaltsas D. & Seyyar O. & Chatzaki M." box="[191,458,1962,1989]" pageId="2" pageNumber="70" pagination="44 - 54" refId="ref18061" refString="Panayiotou E., Kaltsas D., Seyyar O. & Chatzaki M. 2010. Revision of the genus Berinda (Araneae, Gnaphosidae) in the east Mediterranean with the description of two new species. Zootaxa 2362: 44 - 54. https: // doi. org / 10.11646 / zootaxa. 2362.1.2" type="journal article" year="2010">
|
||
Panayiotou
|
||
<emphasis id="B924EA81037DEF631F7C78D8FE426A85" box="[328,387,1962,1988]" italics="true" pageId="2" pageNumber="70">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). Thus, a generic transfer is proposed, resulting in the new combination
|
||
<taxonomicName id="4C504D10037DEF631B2C78D9FEF06AA9" authorityName="Zamani & Chatzaki & Esyunin & Marusik" authorityYear="2021" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" class="Arachnida" family="Gnaphosidae" genus="Berinda" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="70" phylum="Arthropoda" rank="species" species="infumatus" status="comb. nov.">
|
||
<emphasis id="B924EA81037DEF631B2C78D9FEF06AA9" italics="true" pageId="2" pageNumber="70">Berinda infumatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21757FA037DEF631F0C78BDFE7B6AA8" box="[312,442,1998,2025]" pageId="2" pageNumber="70" rank="species">comb. nov.</taxonomicNameLabel>
|
||
(O. Pickard-Cambridge, 1872).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |