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<document id="E92C0575249E417DC889A4F60D9150BD" ID="10.11646/zootaxa.4244.1.1SLASH10518" ID-CLB-Dataset="36256" ID-DOI="10.11646/zootaxa.4244.1.1" ID-GBIF-Dataset="05f1a37b-29f3-4bca-a735-178a3373bb8a" ID-ISSN="1175-5326" ID-Zenodo-Dep="400400" ID-ZooBank="781D71C8-4632-4D1B-8D82-F77CA1146029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489738266586" checkinUser="plazi" docAuthor="Hogans, W. E." docDate="2017" docId="03F777535B78FFEFD6A0FDD273A4F3D7" docLanguage="en" docName="zootaxa.4244.1.1SLASH10518.pdf" docOrigin="Zootaxa 4244 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pennella diodontis Oken 1816" docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="FFCE0F2B5B77FFFFD637FFC57039F11B" masterDocTitle="Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean" masterLastPageNumber="38" masterPageNumber="1" pageNumber="16" updateTime="1698438091026" updateUser="ExternalLinkService">
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<mods:title id="3981A1253E20966BE8CA386F0A461F97">Review of Pennella Oken, 1816 (Copepoda: Pennellidae) with a description of Pennella benzi sp. nov., a parasite of Escolar, Lepidocybium flavobrunneum (Pisces) in the northwest Atlantic Ocean</mods:title>
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<treatment id="03F777535B78FFEFD6A0FDD273A4F3D7" ID-DOI="http://doi.org/10.5281/zenodo.6052533" ID-GBIF-Taxon="127847758" ID-Zenodo-Dep="6052533" LSID="urn:lsid:plazi:treatment:03F777535B78FFEFD6A0FDD273A4F3D7" httpUri="http://treatment.plazi.org/id/03F777535B78FFEFD6A0FDD273A4F3D7" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C34495CE5B78FFF0D6A0FDD270D2F348" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BE1C6455B78FFF0D6A0FDD27237F329" blockId="15.[151,526,535,595]" box="[151,526,535,562]" pageId="15" pageNumber="16">
<heading id="D0A971295B78FFF0D6A0FDD27237F329" bold="true" box="[151,526,535,562]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C5EBDC65B78FFF0D6A0FDD27237F329" authority="Oken, 1816" authorityName="Oken" authorityYear="1816" box="[151,526,535,562]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B78FFF0D6A0FDD27237F329" bold="true" box="[151,526,535,562]" pageId="15" pageNumber="16">
<emphasis id="B92A1A575B78FFF0D6A0FDD27141F32A" bold="true" box="[151,376,535,561]" italics="true" pageId="15" pageNumber="16">Pennella diodontis</emphasis>
<bibRefCitation id="EFCFBBB45B78FFF0D7B7FDDD7237F329" author="Oken" box="[384,526,536,562]" pageId="15" pageNumber="16" refString="Oken, L. (1816) Lehrbuch der Naturgeschichte. Dritter Theil, Zoologie. Fol. 2. A. Schmid, Jena, 1299 pp. [pp. 181, 357]" type="book" year="1816">Oken, 1816</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE1C6455B78FFF0D6A0FDFF70D2F348" blockId="15.[151,526,535,595]" box="[151,235,570,595]" pageId="15" pageNumber="16">
(
<figureCitation id="1365DAC05B78FFF0D6A8FDFF70DAF348" box="[159,227,570,595]" captionStart="FIGURE 7" captionStartId="15.[151,250,1955,1977]" captionTargetBox="[264,1324,775,1918]" captionTargetId="figure@15.[263,1324,754,1936]" captionTargetPageId="15" captionText="FIGURE 7. Pennella diodontis, a. entire parasite; b. cephalothorax, dorsal; c. same, ventral; d. first and second antenna, dorsal; e. second and third swimming legs (redrawn from Kirtisinghe, 1935). Scale bars: a = 2.0 mm; b, c = 0.2 mm; d, e = 0.02 mm." httpUri="https://zenodo.org/record/400406/files/figure.png" pageId="15" pageNumber="16">Fig. 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34495CE5B78FFEFD6A0FD47722BF1E3" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BE1C6455B78FFF0D6A0FD477366F3A4" blockId="15.[151,1008,642,703]" pageId="15" pageNumber="16">
<materialsCitation id="3B36CC185B78FFF0D6A0FD477366F3A4" ID-GBIF-Occurrence="1440667660" country="Philippines" location="Philippines" pageId="15" pageNumber="16" specimenCount="1" typeStatus="holotype">
<emphasis id="B92A1A575B78FFF0D6A0FD477120F380" bold="true" box="[151,281,642,667]" pageId="15" pageNumber="16">Synonyms.</emphasis>
<taxonomicName id="4C5EBDC65B78FFF0D717FD4772FAF380" authority="Heegaard, 1943" authorityName="Heegaard" authorityYear="1943" box="[288,707,642,667]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cervicornis">
<emphasis id="B92A1A575B78FFF0D717FD477231F380" box="[288,520,642,667]" italics="true" pageId="15" pageNumber="16">Pennella cervicornis</emphasis>
Heegaard, 1943
</taxonomicName>
,
<taxonomicName id="4C5EBDC65B78FFF0D4F9FD4173C9F380" authority="Dekay, 1844" authorityName="Dekay" authorityYear="1844" box="[718,1008,642,667]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="plumosa">
<emphasis id="B92A1A575B78FFF0D4F9FD417370F380" box="[718,841,642,667]" italics="true" pageId="15" pageNumber="16">P. plumosa</emphasis>
(
<bibRefCitation id="EFCFBBB45B78FFF0D560FD4773DEF380" author="Dekay" box="[855,999,642,667]" pageId="15" pageNumber="16" refString="Dekay, J. E. (1844) Zoology of New York or the New York fauna. Part IF. Crustacea. W. &amp; A. White &amp; J. Visscher, Albany, N. Y., 415 pp. [pp. 60]" type="book" year="1844">Dekay, 1844</bibRefCitation>
)
</taxonomicName>
<emphasis id="B92A1A575B78FFF0D6F0FD6371EBF3A4" bold="true" box="[199,466,678,703]" pageId="15" pageNumber="16">
<typeStatus id="54E578E75B78FFF0D6F0FD63713BF3A4" box="[199,258,678,703]" pageId="15" pageNumber="16">Type</typeStatus>
host and locality.
</emphasis>
<taxonomicName id="4C5EBDC65B78FFF0D7EEFD6372F2F3A4" box="[473,715,678,703]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="sexmaculatus">
<emphasis id="B92A1A575B78FFF0D7EEFD6372F2F3A4" box="[473,715,678,703]" italics="true" pageId="15" pageNumber="16">Diodon sexmaculatus</emphasis>
</taxonomicName>
,
<collectingCountry id="F34986D55B78FFF0D4EEFD637362F3A4" box="[729,859,678,703]" name="Philippines" pageId="15" pageNumber="16">Philippines</collectingCountry>
.
</materialsCitation>
</paragraph>
<caption id="DF2196CD5B78FFF0D6A0F86675B5F6C3" httpUri="https://zenodo.org/record/400406/files/figure.png" pageId="15" pageNumber="16" targetBox="[264,1324,775,1918]" targetPageId="15">
<paragraph id="8BE1C6455B78FFF0D6A0F86675B5F6C3" blockId="15.[151,1436,1955,2008]" pageId="15" pageNumber="16">
<emphasis id="B92A1A575B78FFF0D6A0F866712BF6A2" bold="true" box="[151,274,1955,1977]" pageId="15" pageNumber="16">FIGURE 7.</emphasis>
<taxonomicName id="4C5EBDC65B78FFF0D72FF86671ECF6A2" box="[280,469,1955,1977]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B78FFF0D72FF86671ECF6A2" box="[280,469,1955,1977]" italics="true" pageId="15" pageNumber="16">Pennella diodontis</emphasis>
</taxonomicName>
, a. entire parasite; b. cephalothorax, dorsal; c. same, ventral; d. first and second antenna, dorsal; e. second and third swimming legs (redrawn from Kirtisinghe, 1935). Scale bars: a = 2.0 mm; b, c = 0.2 mm; d, e = 0.02 mm.
</paragraph>
</caption>
<paragraph id="8BE1C6455B67FFEFD6F0FF52722BF1E3" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">
<emphasis id="B92A1A575B67FFEFD6F0FF527158F1AB" bold="true" box="[199,353,151,176]" pageId="16" pageNumber="17">Morphology.</emphasis>
Size:
<quantity id="4CA66BA05B67FFEFD79FFF5D721FF1B4" box="[424,550,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="16" pageNumber="17" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">2428 mm</quantity>
. Papillae: partial coverage, clavate and club-like, consistent in size and shape, not found in organized groups. Holdfasts: two only, short. First antenna with three segments, second with two segments. Plumes: simple, single.
</paragraph>
</subSubSection>
<subSubSection id="C34495CE5B67FFEFD6F0FEC173A4F3D7" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BE1C6455B67FFEFD6F0FEC173A4F3D7" blockId="16.[151,1436,151,717]" pageId="16" pageNumber="17">
<emphasis id="B92A1A575B67FFEFD6F0FEC17102F006" bold="true" box="[199,315,260,285]" pageId="16" pageNumber="17">Remarks.</emphasis>
A valid species. Original description from porcupine fish,
<taxonomicName id="4C5EBDC65B67FFEFD5F9FEC174F9F006" box="[974,1216,260,285]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="sexmaculatus">
<emphasis id="B92A1A575B67FFEFD5F9FEC174F9F006" box="[974,1216,260,285]" italics="true" pageId="16" pageNumber="17">Diodon sexmaculatus</emphasis>
</taxonomicName>
(=
<emphasis id="B92A1A575B67FFEFD2D0FEC075A5F006" box="[1255,1436,260,285]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C5EBDC65B67FFEFD2D0FEC075ADF006" box="[1255,1428,260,285]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="holacanthus">D. holacanthus</taxonomicName>
)
</emphasis>
(
<bibRefCitation id="EFCFBBB45B67FFEFD6A9FEE27118F05B" author="Oken" box="[158,289,295,320]" pageId="16" pageNumber="17" refString="Oken, L. (1816) Lehrbuch der Naturgeschichte. Dritter Theil, Zoologie. Fol. 2. A. Schmid, Jena, 1299 pp. [pp. 181, 357]" type="book" year="1816">Oken 1816</bibRefCitation>
); redescription from the same host species,
<collectingCountry id="F34986D55B67FFEFD526FEE2735BF05B" box="[785,866,295,320]" name="Sri Lanka" pageId="16" pageNumber="17">Ceylon</collectingCountry>
(
<bibRefCitation id="EFCFBBB45B67FFEFD546FEE274BCF05B" author="Kirtisinghe" box="[881,1157,295,320]" pageId="16" pageNumber="17" refString="Kirtisinghe, P. (1932) Pennella zeylanica n. sp. a parasitic copepod of Histiophorous gladius Day. Parasitology, 24, 137 - 139. https: // doi. org / 10.1017 / S 0031182000020473" type="journal article" year="1932">
Kirtisinghe 1932;
<figureCitation id="1365DAC05B67FFEFD276FEE274BCF05B" box="[1089,1157,295,320]" captionStart="FIGURE 7" captionStartId="15.[151,250,1955,1977]" captionTargetBox="[264,1324,775,1918]" captionTargetId="figure@15.[263,1324,754,1936]" captionTargetPageId="15" captionText="FIGURE 7. Pennella diodontis, a. entire parasite; b. cephalothorax, dorsal; c. same, ventral; d. first and second antenna, dorsal; e. second and third swimming legs (redrawn from Kirtisinghe, 1935). Scale bars: a = 2.0 mm; b, c = 0.2 mm; d, e = 0.02 mm." httpUri="https://zenodo.org/record/400406/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
</bibRefCitation>
). Considered incorrectly as infesting two host genera:
<taxonomicName id="4C5EBDC65B67FFEFD7D1FE897279F07E" box="[486,576,332,357]" class="Actinopterygii" family="Lophiidae" genus="Lophius" kingdom="Animalia" order="Lophiiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B92A1A575B67FFEFD7D1FE897279F07E" box="[486,576,332,357]" italics="true" pageId="16" pageNumber="17">Lophius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5EBDC65B67FFEFD44DFE897345F07F" authority="Wilson 1917" authorityName="Wilson" authorityYear="1917" box="[634,892,332,357]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B92A1A575B67FFEFD44DFE8972F7F07E" box="[634,718,332,357]" italics="true" pageId="16" pageNumber="17">Diodon</emphasis>
(
<bibRefCitation id="EFCFBBB45B67FFEFD4E9FE89734DF07F" author="Wilson" box="[734,884,332,357]" pageId="16" pageNumber="17" refString="Wilson, C. B. (1917) North American parasitic copepods belonging to the family Lernaeidae with a revision of the entire family. Proceedings of the United States National Museum, 53, 1 - 150." type="journal article" year="1917">Wilson 1917</bibRefCitation>
)
</taxonomicName>
. The history of
<taxonomicName id="4C5EBDC65B67FFEFD204FE887481F07E" box="[1075,1208,332,357]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD204FE887481F07E" box="[1075,1208,332,357]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
was often confused with that
<emphasis id="B92A1A575B67FFEFD6C8FEB5712CF093" box="[255,277,368,392]" italics="true" pageId="16" pageNumber="17">of</emphasis>
<taxonomicName id="4C5EBDC65B67FFEFD72BFEB47215F093" authority="Wilson 1917" authorityName="Wilson" authorityYear="1917" box="[284,556,367,392]" class="Anthozoa" family="Pennatulidae" genus="Pennatula" kingdom="Animalia" order="Pennatulacea" pageId="16" pageNumber="17" phylum="Cnidaria" rank="species" species="sagitta">
<emphasis id="B92A1A575B67FFEFD72BFEB471BBF093" box="[284,386,369,392]" italics="true" pageId="16" pageNumber="17">P. sagitta</emphasis>
(
<bibRefCitation id="EFCFBBB45B67FFEFD7A6FEAA721CF093" author="Wilson" box="[401,549,367,392]" pageId="16" pageNumber="17" refString="Wilson, C. B. (1917) North American parasitic copepods belonging to the family Lernaeidae with a revision of the entire family. Proceedings of the United States National Museum, 53, 1 - 150." type="journal article" year="1917">Wilson 1917</bibRefCitation>
)
</taxonomicName>
, however,
<taxonomicName id="4C5EBDC65B67FFEFD493FEB4731EF093" box="[676,807,367,392]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD493FEB4731EF093" box="[676,807,367,392]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
reported from
<taxonomicName id="4C5EBDC65B67FFEFD5F9FEAA7411F093" box="[974,1064,367,392]" class="Actinopterygii" family="Lophiidae" genus="Lophius" kingdom="Animalia" order="Lophiiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B92A1A575B67FFEFD5F9FEAA7411F093" box="[974,1064,367,392]" italics="true" pageId="16" pageNumber="17">Lophius</emphasis>
</taxonomicName>
was an error of
<bibRefCitation id="EFCFBBB45B67FFEFD2EBFEAA75A5F093" author="Blainville" box="[1244,1436,367,392]" pageId="16" pageNumber="17" refString="Blainville, M. H. D. (1822) Memoire sur les Lernaees (Lernaea, Linn.). Journal Physique, 95, 437 - 447." type="journal article" year="1822">Blainville (1822)</bibRefCitation>
based on a poor description and incorrect host identification;
<taxonomicName id="4C5EBDC65B67FFEFD568FE5073DCF0B6" box="[863,997,404,429]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD568FE5073DCF0B6" box="[863,997,404,429]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
is in fact a parasite of porcupine fish (
<taxonomicName id="4C5EBDC65B67FFEFD6A9FE727116F0CB" box="[158,303,439,464]" class="Actinopterygii" family="Diodontidae" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="family">Diodontidae</taxonomicName>
:
<taxonomicName id="4C5EBDC65B67FFEFD773FE7271A1F0CB" box="[324,408,439,464]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B92A1A575B67FFEFD773FE7271A1F0CB" box="[324,408,439,464]" italics="true" pageId="16" pageNumber="17">Diodon</emphasis>
</taxonomicName>
) and
<taxonomicName id="4C5EBDC65B67FFEFD7DBFE7C7266F0CB" box="[492,607,441,464]" class="Anthozoa" family="Pennatulidae" genus="Pennatula" kingdom="Animalia" order="Pennatulacea" pageId="16" pageNumber="17" phylum="Cnidaria" rank="species" species="sagitta">
<emphasis id="B92A1A575B67FFEFD7DBFE7C7266F0CB" box="[492,607,441,464]" italics="true" pageId="16" pageNumber="17">P. sagitta</emphasis>
</taxonomicName>
is found so far only on frogfish (Antennaridae:
<emphasis id="B92A1A575B67FFEFD2E0FE7C7514F0CB" box="[1239,1325,441,464]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C5EBDC65B67FFEFD2E0FE7C7510F0CB" box="[1239,1321,441,464]" class="Actinopterygii" family="Antennariidae" genus="Histrio" kingdom="Animalia" order="Lophiiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">Histrio</taxonomicName>
,
</emphasis>
possibly
<emphasis id="B92A1A575B67FFEFD6A0FE197110F0E8" box="[151,297,476,499]" italics="true" pageId="16" pageNumber="17">
<taxonomicName id="4C5EBDC65B67FFEFD6A0FE197118F0E8" box="[151,289,476,499]" class="Actinopterygii" family="Antennariidae" genus="Antennarius" kingdom="Animalia" order="Lophiiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">Antennarius</taxonomicName>
)
</emphasis>
).
<bibRefCitation id="EFCFBBB45B67FFEFD777FE18729CF0EF" author="Lasarus" box="[320,677,476,501]" pageId="16" pageNumber="17" refString="Lasarus, S. &amp; Sreenivasan, P. V. (1977) On a copepod parasite Pennella diodontis Oken with epizoic cirripede Chonchoderma virgatum Spengler on a new host Zanculus canascens (Linnaeus). Indian Journal of Fisheries, 24 (2), 204 - 206." type="journal article" year="1977">Lasarus and Sreenivasan (1977)</bibRefCitation>
reported
<taxonomicName id="4C5EBDC65B67FFEFD538FE1873ABF0EE" box="[783,914,476,501]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD538FE1873ABF0EE" box="[783,914,476,501]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
from the moorish idol (
<emphasis id="B92A1A575B67FFEFD2ABFE197545F0EF" box="[1180,1404,476,501]" italics="true" pageId="16" pageNumber="17">Zanculus canascens</emphasis>
(=
<emphasis id="B92A1A575B67FFEFD6A0FDC47137F303" box="[151,270,513,536]" italics="true" pageId="16" pageNumber="17">Z. cornuta</emphasis>
) in the Indian Ocean. The single habitus figure and brief description are not sufficient to confidently place their specimens (eight) in
<taxonomicName id="4C5EBDC65B67FFEFD43CFDE072ABF326" box="[523,658,548,573]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD43CFDE072ABF326" box="[523,658,548,573]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
. However, the size of the parasite (
<quantity id="4CA66BA05B67FFEFD207FDE17443F326" box="[1072,1146,548,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="17" unit="mm" value="26.0">26mm</quantity>
) and the cephalothoracic papillae (few small at center, and larger branched papillae around the cephalothorax periphery) are reminiscent of the species. As a parasite of
<taxonomicName id="4C5EBDC65B67FFEFD7D3FDA97201F39E" box="[484,568,620,645]" class="Actinopterygii" family="Diodontidae" genus="Diodon" kingdom="Animalia" order="Tetraodontiformes" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B92A1A575B67FFEFD7D3FDA97201F39E" box="[484,568,620,645]" italics="true" pageId="16" pageNumber="17">Diodon</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5EBDC65B67FFEFD47EFDA872F6F39E" box="[585,719,620,645]" class="Maxillopoda" family="Pennellidae" genus="Pennella" kingdom="Animalia" order="Siphonostomatoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="diodontis">
<emphasis id="B92A1A575B67FFEFD47EFDA872F6F39E" box="[585,719,620,645]" italics="true" pageId="16" pageNumber="17">P. diodontis</emphasis>
</taxonomicName>
is a small species with uniformity in most structures between specimens (
<bibRefCitation id="EFCFBBB45B67FFEFD72AFD5571FFF3B3" author="Hogans" box="[285,454,656,680]" pageId="16" pageNumber="17" refString="Hogans, W. E. (1988 a) Redescription of Pennella sagitta (Copepoda: Pennellidae) from Histrio histrio (Pisces) in the northwest Atlantic Ocean with a provisional review of the genus Pennella Oken, 1816. Journal of Zoology, London, 216 (2), 379 - 390." type="journal article" year="1988" yearSuffix="a">Hogans 1988a</bibRefCitation>
), however, its cephalothoracic papillae may vary slightly in arrangement and number depending on the individual parasite (
<bibRefCitation id="EFCFBBB45B67FFEFD40AFD7172DAF3D7" author="Hogans" box="[573,739,692,716]" pageId="16" pageNumber="17" refString="Hogans, W. E. (1988 a) Redescription of Pennella sagitta (Copepoda: Pennellidae) from Histrio histrio (Pisces) in the northwest Atlantic Ocean with a provisional review of the genus Pennella Oken, 1816. Journal of Zoology, London, 216 (2), 379 - 390." type="journal article" year="1988" yearSuffix="a">Hogans 1988a</bibRefCitation>
;
<bibRefCitation id="EFCFBBB45B67FFEFD4D8FD7173A9F3D7" author="Heegard" box="[751,912,692,717]" pageId="16" pageNumber="17" refString="Heegard, P. (1943) Parasitic copepods mainly from tropical and Antarctic seas. Arkives Zoology, 34 (A), 1 - 37." type="journal article" year="1943">Heegard 1943</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>