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<document id="8CCE3C3991BC72BA0A913F6D8C185AA7" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF86CD6CFF4CE98AB384EB5E" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heterocaucaseuma mauriesi Golovatch &amp; Makarov 2011, comb. nov." docType="treatment" docVersion="8" lastPageNumber="83" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="82" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="F52EE744C3254330B846A98D12EF30CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="45968E778DE2E2B6FF704C40B1F14B10">
<mods:title id="ACA8BB20525435661E976189BBE5865D">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="5F5A717275DAFDDE6E50D22B0CCCBAFB" type="personal">
<mods:role id="E79DAE7FF0568F2F992517CB96B1EAE2">
<mods:roleTerm id="68763BA5BB9C6078F99735D8B30F42DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2EDF2DADA9B31C03ECAD35E489ABDBA">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="4393B6B4E4370AEE38BFDB212F9F25EA" type="personal">
<mods:role id="C4853037BC24DEF135138AC8A156876B">
<mods:roleTerm id="7C28DBA36A33067EA5AA84DFE8BDE0DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A5070CE2CEE216F3E2D24CF6FDF9080E">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="07885B4C4A23D33EBAAD90342F3E4361">text</mods:typeOfResource>
<mods:relatedItem id="D27253DA47B015E127F3C0197CC00EE3" type="host">
<mods:titleInfo id="9B6968070A529C70FA6BD045D732E0FA">
<mods:title id="64D8CEF0FDF98E1E73945B8202CA87E6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="02FF757F4E957031DA700F6B7AE4E697">
<mods:date id="FE0D299D7FFAFC89D161016654A56C87">2016</mods:date>
<mods:detail id="1B87E3C74949F773C98C7AD4C945A1A5" type="volume">
<mods:number id="0D5E718604AFB3884735DDA0C2D5846D">4211</mods:number>
</mods:detail>
<mods:detail id="616BCA969A696B53CAFB7D9D80BB45C7" type="issue">
<mods:number id="B04F09C1EC60F9E61BFFAE5D2B2A234F">1</mods:number>
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<mods:start id="795B25BAC0D3DA7DE0EBDBC5500650E6">1</mods:start>
<mods:end id="4CDD83E9E9A4636DB3F0C21D514396EB">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A5B420E1E62281C555CA0393C68607E8">journal article</mods:classification>
<mods:identifier id="D81FE84D3DCEA93452E6FFD9C70C4D70" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="8B0F47FD6D36462481CCDDA5FC8C7924" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="79F330E8E5B646A6CAAE5F5C9BB09176" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="466BE93AB8E3B81E996B9E7717AA7FAB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="071459B4ED0D3606086272A0A4824722" type="Zenodo-Dep">272750</mods:identifier>
<mods:identifier id="F4E5BFE9E6AE57598AA83CA1ADFDB30A" type="ZooBank">6B86C6BA-6AFE-4AAD-870D-04794C138D47</mods:identifier>
</mods:mods>
<treatment id="03B7878CFF86CD6CFF4CE98AB384EB5E" ID-DOI="http://doi.org/10.5281/zenodo.6074411" ID-GBIF-Taxon="127419154" ID-Zenodo-Dep="6074411" LSID="urn:lsid:plazi:treatment:03B7878CFF86CD6CFF4CE98AB384EB5E" httpUri="http://treatment.plazi.org/id/03B7878CFF86CD6CFF4CE98AB384EB5E" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<subSubSection id="C3046511FF86CD6FFF4CE98AB195EB72" pageId="81" pageNumber="82" type="nomenclature">
<paragraph id="8BA1369AFF86CD6FFF4CE98AB289EB53" blockId="81.[151,1018,511,571]" box="[151,1018,511,538]" pageId="81" pageNumber="82">
<heading id="D0E981F6FF86CD6FFF4CE98AB289EB53" bold="true" box="[151,1018,511,538]" fontSize="11" level="1" pageId="81" pageNumber="82" reason="1">
<emphasis id="B96AEA88FF86CD6FFF4CE98AB289EB53" bold="true" box="[151,1018,511,538]" pageId="81" pageNumber="82">
<taxonomicName id="4C1E4D19FF86CD6FFF4CE98AB21FEB53" ID-CoL="849BR" authority="Golovatch &amp; Makarov, 2011" authorityName="Golovatch &amp; Makarov" authorityYear="2011" box="[151,876,511,538]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mauriesi" status="comb. nov.">
<emphasis id="B96AEA88FF86CD6FFF4CE98AB082EB50" bold="true" box="[151,497,511,537]" italics="true" pageId="81" pageNumber="82">Heterocaucaseuma mauriesi</emphasis>
(
<bibRefCitation id="EF8F4B6BFF86CD6FFDDBEA75B216EB53" author="Golovatch" box="[512,869,512,538]" pageId="81" pageNumber="82" refString="Golovatch, S. I. &amp; Makarov, S. E. (2011) Two new, easternmost species of the millipede genus Anamastigona Silvestri, 1898 from Israel and the Caucasus (Diplopoda: Chordeumatida: Anthroleucosomatidae). Arthropoda Selecta, 20 (1), 11 - 20." type="journal article" year="2011">Golovatch &amp; Makarov, 2011</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25957F3FF86CD6FFCAFEA75B289EB53" box="[884,1018,512,538]" pageId="81" pageNumber="82">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF86CD6FFF4CEA57B195EB72" blockId="81.[151,1018,511,571]" box="[151,230,546,571]" pageId="81" pageNumber="82">
<figureCitation id="13252A1FFF86CD6FFF4CEA57B195EB72" box="[151,230,546,571]" captionStart="FIGURE 66" captionStartId="81.[151,250,1763,1785]" captionTargetBox="[188,1398,901,1741]" captionTargetId="figure@81.[188,1398,901,1741]" captionTargetPageId="81" captionText="FIGURE 66. Heterocaucaseuma mauriesi (Golovatch &amp; Makarov, 2011), comb. nov., paratype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272816/files/figure.png" pageId="81" pageNumber="82">Fig. 66</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF86CD6FFF4CEA1EB273EBD6" pageId="81" pageNumber="82" type="reference_group">
<paragraph id="8BA1369AFF86CD6FFF4CEA1EB273EBD6" blockId="81.[151,892,618,671]" pageId="81" pageNumber="82">
<treatmentCitationGroup id="AB0E11B4FF86CD6FFF4CEA1EB273EBD6" pageId="81" pageNumber="82">
<taxonomicName id="4C1E4D19FF86CD6FFF4CEA1EB3A6EBC9" ID-CoL="7R6JG" authority="Golovatch &amp; Makarov, 2011: 16" authorityName="Golovatch &amp; Makarov" authorityPageNumber="16" authorityYear="2011" box="[151,725,618,640]" class="Diplopoda" family="Anthroleucosomatidae" genus="Anamastigona" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mauriesi">
<emphasis id="B96AEA88FF86CD6FFF4CEA1EB0FBEBC9" box="[151,392,619,640]" italics="true" pageId="81" pageNumber="82">Anamastigona mauriesi</emphasis>
<treatmentCitation id="0ABF108BFF86CD6FFE55EA1FB3A6EBC9" author="Golovatch" box="[398,725,618,640]" page="16" pageId="81" pageNumber="82" year="2011">
<bibRefCitation id="EF8F4B6BFF86CD6FFE55EA1FB3C0EBC9" author="Golovatch" box="[398,691,618,640]" pageId="81" pageNumber="82" refString="Golovatch, S. I. &amp; Makarov, S. E. (2011) Two new, easternmost species of the millipede genus Anamastigona Silvestri, 1898 from Israel and the Caucasus (Diplopoda: Chordeumatida: Anthroleucosomatidae). Arthropoda Selecta, 20 (1), 11 - 20." type="journal article" year="2011">Golovatch &amp; Makarov, 2011</bibRefCitation>
: 16
</treatmentCitation>
</taxonomicName>
–20, figs 31–45.
<emphasis id="B96AEA88FF86CD6FFF4CEAFFB0EDEBD6" box="[151,414,650,671]" italics="true" pageId="81" pageNumber="82">
<taxonomicName id="4C1E4D19FF86CD6FFF4CEAFFB05AEBD6" ID-CoL="7NPLP" box="[151,297,650,671]" class="Diplopoda" family="Anthroleucosomatidae" genus="Anamastigona" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus">Anamastigona</taxonomicName>
mauriesi—
</emphasis>
<treatmentCitation id="0ABF108BFF86CD6FFE45EAFCB38FEBD6" author="Vagalinski" box="[414,764,649,671]" page="12" pageId="81" pageNumber="82" year="2016">
<bibRefCitation id="EF8F4B6BFF86CD6FFE45EAFCB3A5EBD6" author="Vagalinski" box="[414,726,649,671]" pageId="81" pageNumber="82" refString="Vagalinski, B. &amp; Golovatch, S. I. (2016) Two new species of Anamastigona from Cyprus and an updated key to species of the genus (Diplopoda: Chordeumatida: Anthroleucosomatidae). European Journal of Taxonomy, 227, 1 - 19." type="journal article" year="2016">Vagalinski &amp; Golovatch, 2016</bibRefCitation>
: 12
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF86CD6FFF4CEAB9B37AEA06" pageId="81" pageNumber="82" type="diagnosis">
<paragraph id="8BA1369AFF86CD6FFF4CEAB9B37AEA06" blockId="81.[151,1436,716,848]" pageId="81" pageNumber="82">
<emphasis id="B96AEA88FF86CD6FFF4CEAB9B062EBAC" bold="true" box="[151,273,716,741]" pageId="81" pageNumber="82">Diagnosis.</emphasis>
Differs from the other
<taxonomicName id="4C1E4D19FF86CD6FFDC2EAB8B383EBAD" box="[537,752,717,740]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96AEA88FF86CD6FFDC2EAB8B383EBAD" box="[537,752,717,740]" italics="true" pageId="81" pageNumber="82">Heterocaucaseuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957F3FF86CD6FFD23EAB9B211EBAC" box="[760,866,716,741]" pageId="81" pageNumber="82" rank="genus">
<emphasis id="B96AEA88FF86CD6FFD23EAB9B211EBAC" bold="true" box="[760,866,716,741]" pageId="81" pageNumber="82">gen. nov.</emphasis>
</taxonomicNameLabel>
species by the presence of lateral horns which are higher than the medial part, but significantly shorter than in
<taxonomicName id="4C1E4D19FF86CD6FFC90EA84B29AEA41" box="[843,1001,751,776]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="longicorne">
<emphasis id="B96AEA88FF86CD6FFC90EA84B29AEA41" box="[843,1001,751,776]" italics="true" pageId="81" pageNumber="82">H. longicorne</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF86CD6FFC2AEA9AB5D6EA41" bold="true" box="[1009,1189,751,776]" pageId="81" pageNumber="82">gen. et sp. nov.</emphasis>
, by the presence of a squarish medial part, in anterior view, of the anterior gonopods, and by the presence of a somewhat subtriangular coxal process on male leg-pair 7.
</paragraph>
</subSubSection>
<caption id="DF616612FF86CD6FFF4CEE96B1B5EE5E" httpUri="https://zenodo.org/record/272816/files/figure.png" pageId="81" pageNumber="82" targetBox="[188,1398,901,1741]" targetPageId="81">
<paragraph id="8BA1369AFF86CD6FFF4CEE96B1B5EE5E" blockId="81.[151,1435,1763,1815]" pageId="81" pageNumber="82">
<emphasis id="B96AEA88FF86CD6FFF4CEE96B052EFB1" bold="true" box="[151,289,1763,1785]" pageId="81" pageNumber="82">FIGURE 66.</emphasis>
<taxonomicName id="4C1E4D19FF86CD6FFEF2EE91B2FCEFB0" authority="Golovatch &amp; Makarov, 2011" authorityName="Golovatch &amp; Makarov" authorityYear="2011" box="[297,911,1763,1785]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="mauriesi" status="comb. nov.">
<emphasis id="B96AEA88FF86CD6FFEF2EE91B33EEFB0" box="[297,589,1764,1785]" italics="true" pageId="81" pageNumber="82">Heterocaucaseuma mauriesi</emphasis>
(Golovatch &amp; Makarov, 2011)
</taxonomicName>
,
<taxonomicNameLabel id="A25957F3FF86CD6FFC40EE96B563EFB0" box="[923,1040,1763,1785]" pageId="81" pageNumber="82">comb. nov.</taxonomicNameLabel>
, paratype male habitus. Scale line = 1 mm.
</paragraph>
</caption>
<subSubSection id="C3046511FF86CD6FFF1CEF4BB518EED6" pageId="81" pageNumber="82" type="materials_examined">
<paragraph id="8BA1369AFF86CD6FFF1CEF4BB280EE1E" blockId="81.[151,1436,1854,2023]" box="[199,1011,1854,1879]" pageId="81" pageNumber="82">
<materialsCitation id="3B763CC7FF86CD6FFF1CEF4BB280EE1E" ID-GBIF-Occurrence="1424567024" box="[199,1011,1854,1879]" location="Cave Tarkiladze" municipality="Duripsh" pageId="81" pageNumber="82" specimenCount="1" stateProvince="ABKHAZIA" typeStatus="holotype">
<emphasis id="B96AEA88FF86CD6FFF1CEF4BB016EE1E" bold="true" box="[199,357,1854,1879]" pageId="81" pageNumber="82">
<typeStatus id="54A58838FF86CD6FFF1CEF4BB071EE1E" box="[199,258,1854,1879]" pageId="81" pageNumber="82">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49DAF878FF86CD6FFEB7EF4AB370EE1E" box="[364,515,1855,1879]" country="Georgia" name="Abkhazia" pageId="81" pageNumber="82">ABKHAZIA</collectingRegion>
:
<collectingMunicipality id="6BC5ACE0FF86CD6FFDD6EF4BB319EE1E" box="[525,618,1854,1879]" pageId="81" pageNumber="82">Duripsh</collectingMunicipality>
,
<location id="8EC16041FF86CD6FFDAEEF4AB25FEE1E" LSID="urn:lsid:plazi:treatment:03B7878CFF86CD6CFF4CE98AB384EB5E:8EC16041FF86CD6FFDAEEF4AB25FEE1E" box="[629,812,1854,1879]" municipality="Duripsh" name="Cave Tarkiladze" pageId="81" pageNumber="82" stateProvince="ABKHAZIA">Cave Tarkiladze</location>
(= Cave Tarkila).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF86CD6FFF1CEF16B518EED6" blockId="81.[151,1436,1854,2023]" pageId="81" pageNumber="82">
<materialsCitation id="3B763CC7FF86CD6FFF1CEF16B3B9EE35" ID-GBIF-Occurrence="1424567056" box="[199,714,1891,1916]" location="Material" pageId="81" pageNumber="82" specimenCount="4" specimenCount-juvenile="3" specimenCount-male="1">
<emphasis id="B96AEA88FF86CD6FFF1CEF16B0FCEE35" bold="true" box="[199,399,1891,1916]" pageId="81" pageNumber="82">
<location id="8EC16041FF86CD6FFF1CEF16B05CEE35" LSID="urn:lsid:plazi:treatment:03B7878CFF86CD6CFF4CE98AB384EB5E:8EC16041FF86CD6FFF1CEF16B05CEE35" box="[199,303,1891,1916]" name="Material" pageId="81" pageNumber="82">Material</location>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF86CD6FFE3FEF16B347EE35" box="[484,564,1891,1916]" pageId="81" pageNumber="82" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF86CD6FFD9AEF16B3B1EE35" box="[577,706,1891,1916]" pageId="81" pageNumber="82" type="juvenile">3 juveniles</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B763CC7FF86CD6FFD0DEF16B5F8EE35" ID-GBIF-Occurrence="1424567073" box="[726,1163,1891,1916]" location="Topotypes" pageId="81" pageNumber="82" specimenCount="4" specimenCount-juvenile="3" specimenCount-male="1">
<location id="8EC16041FF86CD6FFD0DEF16B223EE35" LSID="urn:lsid:plazi:treatment:03B7878CFF86CD6CFF4CE98AB384EB5E:8EC16041FF86CD6FFD0DEF16B223EE35" box="[726,848,1891,1916]" name="Topotypes" pageId="81" pageNumber="82">
<emphasis id="B96AEA88FF86CD6FFD0DEF16B223EE35" bold="true" box="[726,848,1891,1916]" pageId="81" pageNumber="82">Topotypes</emphasis>
</location>
(total:
<specimenCount id="9D18FD13FF86CD6FFC7FEF16B287EE35" box="[932,1012,1891,1916]" pageId="81" pageNumber="82" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF86CD6FFBD9EF16B5F1EE35" box="[1026,1154,1891,1916]" pageId="81" pageNumber="82" type="juvenile">3 juveniles</specimenCount>
)
</materialsCitation>
.
<materialsCitation id="3B763CC7FF86CD6FFB4CEF16B518EED6" ID-GBIF-Occurrence="1424567067" collectingDate="1943-11-19" collectionCode="ZMUM" collectorName="Y. A. Birstein" location="Cave Tarkiladze" municipality="All" pageId="81" pageNumber="82" specimenCount="4" specimenCount-juvenile="3" specimenCount-male="1" stateProvince="ABKHAZIA">
<collectingMunicipality id="6BC5ACE0FF86CD6FFB4CEF16B5C8EE35" box="[1175,1211,1891,1916]" pageId="81" pageNumber="82">All</collectingMunicipality>
from
<collectingRegion id="49DAF878FF86CD6FFAD8EF16B4EBEE32" box="[1283,1432,1891,1915]" country="Georgia" name="Abkhazia" pageId="81" pageNumber="82">ABKHAZIA</collectingRegion>
,
<location id="8EC16041FF86CD6FFF4CEFF2B022EED6" LSID="urn:lsid:plazi:treatment:03B7878CFF86CD6CFF4CE98AB384EB5E:8EC16041FF86CD6FFF4CEFF2B022EED6" box="[151,337,1926,1951]" municipality="All" name="Cave Tarkiladze" pageId="81" pageNumber="82" stateProvince="ABKHAZIA">Cave Tarkiladze</location>
:
<specimenCount id="9D18FD13FF86CD6FFE87EFF2B0D9EED6" box="[348,426,1926,1951]" pageId="81" pageNumber="82" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF86CD6FFE6EEFF2B341EED6" box="[437,562,1926,1951]" pageId="81" pageNumber="82" type="juvenile">3 juveniles</specimenCount>
,
<collectingDate id="EFE4E9B2FF86CD6FFDE5EFF2B3ABEED6" box="[574,728,1927,1951]" pageId="81" pageNumber="82" value="1943-11-19">
<date id="FFA0105AFF86CD6FFDE5EFF2B3ABEED6" box="[574,728,1927,1951]" pageId="81" pageNumber="82" value="1943-11-19">19 Nov. 1943</date>
</collectingDate>
,
<collectorName id="26EB534CFF86CD6FFD38EFF2B205EED6" box="[739,886,1926,1951]" pageId="81" pageNumber="82">Y.A. Birstein</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF86CD6FFC6FEFF2B57EEED6" box="[948,1037,1927,1951]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="81" pageNumber="82">ZMUM</collectionCode>
ρ3309).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF86CD6CFF1CEFDEB384EB5E" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="distribution">
<paragraph id="8BA1369AFF86CD6FFF1CEFDEB5F9EE8D" blockId="81.[151,1436,1854,2023]" box="[199,1162,1963,1988]" pageId="81" pageNumber="82">
<emphasis id="B96AEA88FF86CD6FFF1CEFDEB013EE8D" bold="true" box="[199,352,1963,1988]" pageId="81" pageNumber="82">Distribution.</emphasis>
<collectingRegion id="49DAF878FF86CD6FFEBCEFDEB0A1EE8D" box="[359,466,1963,1988]" country="Georgia" name="Abkhazia" pageId="81" pageNumber="82">Abkhazia</collectingRegion>
(known only from Cave Tarkiladze) (
<figureCitation id="13252A1FFF86CD6FFCA0EFDEB2A8EE8D" box="[891,987,1963,1988]" captionStart="FIGURE 167" captionStartId="196.[151,250,2007,2029]" captionTargetBox="[188,1398,1136,1986]" captionTargetId="figure@196.[188,1398,1136,1986]" captionTargetPageId="196" captionText="FIGURE 167. Distribution of the Caucasian Anthroleucosomatidae: the Caucaseuma complex." httpUri="https://zenodo.org/record/272916/files/figure.png" pageId="81" pageNumber="82">Fig. 167</figureCitation>
, orange circle).
</paragraph>
<paragraph id="8BA1369AFF86CD6CFF1CEFBBB294E899" blockId="81.[151,1436,1854,2023]" lastBlockId="82.[151,1436,151,536]" lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<emphasis id="B96AEA88FF86CD6FFF1CEFBBB075EEAE" bold="true" box="[199,262,1998,2023]" pageId="81" pageNumber="82">Note.</emphasis>
This,
<emphasis id="B96AEA88FF86CD6FFE97EFA5B015EEAE" box="[332,358,2000,2023]" italics="true" pageId="81" pageNumber="82">ca</emphasis>
<quantity id="4CE69B7FFF86CD6FFEB5EFBAB0CEEEAE" box="[366,445,1999,2023]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="81" pageNumber="82" unit="mm" value="30.0">30 mm</quantity>
long species is characterized as by the presence of some troglomorphic features, such as reduced body pigmentation and somewhat elongated walking legs and antennae.
<bibRefCitation id="EF8F4B6BFF85CD6CFBFEE8E2B04CE99D" author="Golovatch" pageId="82" pageNumber="83" refString="Golovatch, S. I. &amp; Makarov, S. E. (2011) Two new, easternmost species of the millipede genus Anamastigona Silvestri, 1898 from Israel and the Caucasus (Diplopoda: Chordeumatida: Anthroleucosomatidae). Arthropoda Selecta, 20 (1), 11 - 20." type="journal article" year="2011">Golovatch &amp; Makarov (2011: 18, figs 33 and 34)</bibRefCitation>
mistakenly show the same drawings for leg-pairs 4 and 5. In this species, leg-pair 5 is characterized by the presence of a well-developed, oral, basal protrusion on the prefemur. As in the previous two species of this genus, the gonopods of this species are also characterized by the presence of an anterior, hairy, sternal sac, and by the presence of a medial syncoxal part and two lateral parts. The syncoxal medial part is squarish, with an anterior projection and with coxal vesicles posteriorly. The lateral parts are significantly shorter than those in
<taxonomicName id="4C1E4D19FF85CD6CFADAE938B4EFE82C" box="[1281,1436,332,357]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="longicorne" status="sp. nov.">
<emphasis id="B96AEA88FF85CD6CFADAE938B4EFE82C" box="[1281,1436,332,357]" italics="true" pageId="82" pageNumber="83">H. longicorne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957F3FF85CD6CFF4CE91AB183E8C1" box="[151,240,367,392]" pageId="82" pageNumber="83" rank="species">
<emphasis id="B96AEA88FF85CD6CFF4CE91AB183E8C1" bold="true" box="[151,240,367,392]" pageId="82" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
, but higher than in
<taxonomicName id="4C1E4D19FF85CD6CFE11E904B3DCE8C1" box="[458,687,368,392]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="feminaepectorum" status="sp. nov.">
<emphasis id="B96AEA88FF85CD6CFE11E904B3DCE8C1" box="[458,687,368,392]" italics="true" pageId="82" pageNumber="83">H. feminaepectorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957F3FF85CD6CFD6EE91AB27CE8C1" box="[693,783,367,392]" pageId="82" pageNumber="83" rank="species">
<emphasis id="B96AEA88FF85CD6CFD6EE91AB27CE8C1" bold="true" box="[693,783,367,392]" pageId="82" pageNumber="83">sp. nov.</emphasis>
</taxonomicNameLabel>
The posterior gonopods consist of the same parts as in the other species of
<taxonomicName id="4C1E4D19FF85CD6CFE8AE9E0B35BE8E5" box="[337,552,405,428]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="202" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96AEA88FF85CD6CFE8AE9E0B35BE8E5" box="[337,552,405,428]" italics="true" pageId="82" pageNumber="83">Heterocaucaseuma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D19FF85CD6CFDBAE9E0B381E8E5" box="[609,754,405,428]" class="Diplopoda" family="Anthroleucosomatidae" genus="Caucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEA88FF85CD6CFDBAE9E0B381E8E5" box="[609,754,405,428]" italics="true" pageId="82" pageNumber="83">Caucaseuma</emphasis>
</taxonomicName>
: coxites divided, each with a pair of flagelliform processes; posterolaterally with telopodital remnants; coxal vesicles present anteriorly.
</paragraph>
<paragraph id="8BA1369AFF85CD6CFF1CE9A9B384EB5E" blockId="82.[151,1436,151,536]" pageId="82" pageNumber="83">
<bibRefCitation id="EF8F4B6BFF85CD6CFF1CE9A9B35EE8BC" author="Vagalinski" box="[199,557,476,501]" pageId="82" pageNumber="83" refString="Vagalinski, B. &amp; Golovatch, S. I. (2016) Two new species of Anamastigona from Cyprus and an updated key to species of the genus (Diplopoda: Chordeumatida: Anthroleucosomatidae). European Journal of Taxonomy, 227, 1 - 19." type="journal article" year="2016">Vagalinski &amp; Golovatch (2016)</bibRefCitation>
, in a recent review of
<taxonomicName id="4C1E4D19FF85CD6CFCF6E9A9B2BCE8BA" box="[813,975,476,499]" class="Diplopoda" family="Anthroleucosomatidae" genus="Anamastigona" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEA88FF85CD6CFCF6E9A9B2BCE8BA" box="[813,975,476,499]" italics="true" pageId="82" pageNumber="83">Anamastigona</emphasis>
</taxonomicName>
, took a more conservative approach and still treated
<taxonomicName id="4C1E4D19FF85CD6CFEC0EA74B0ECEB51" box="[283,415,513,536]" class="Diplopoda" family="Anthroleucosomatidae" genus="Heterocaucaseuma" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="mauriesi">
<emphasis id="B96AEA88FF85CD6CFEC0EA74B0ECEB51" box="[283,415,513,536]" italics="true" pageId="82" pageNumber="83">H. mauriesi</emphasis>
</taxonomicName>
as a species of
<taxonomicName id="4C1E4D19FF85CD6CFD95EA75B383EB5E" box="[590,752,512,535]" class="Diplopoda" family="Anthroleucosomatidae" genus="Anamastigona" kingdom="Animalia" order="Chordeumatida" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEA88FF85CD6CFD95EA75B383EB5E" box="[590,752,512,535]" italics="true" pageId="82" pageNumber="83">Anamastigona</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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