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<document id="4F547DED16BA8F5135230E3CEBAF7B7B" ID-DOI="10.11646/phytotaxa.172.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="5142201" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627482379846" checkinUser="felipe" docAuthor="Fateryga, Alexander V., Fateryga, Valentina V. &amp; Efimov, Petr G." docDate="2014" docId="03A99C58EB3C273A0AC6FCF3FA76F90C" docLanguage="en" docName="Phytotaxa.172.1.022-030.pdf" docOrigin="Phytotaxa 172 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.172.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Epipactis (sect. Epipactis) sect. Epipactis" docType="key" docVersion="3" lastPageNumber="28" masterDocId="FF90E420EB3A273C0A4EFFFFFFDDFFA5" masterDocTitle="Epipactis krymmontana (Orchidaceae), a new species endemic to the Crimean Mountains and notes on the related taxa in the Crimea and bordering Russian Caucasus" masterLastPageNumber="30" masterPageNumber="22" pageNumber="28" updateTime="1699025980482" updateUser="plazi" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="28AAE12B1BEBAA8180A440C77BD5E195">Epipactis krymmontana (Orchidaceae), a new species endemic to the Crimean Mountains and notes on the related taxa in the Crimea and bordering Russian Caucasus</mods:title>
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<mods:namePart id="6E64A66D332B8757BB4D1BF22933DBB0">Fateryga, Alexander V.</mods:namePart>
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<mods:namePart id="CDE3BEA3254A84AD4158E911AE8C52F0">Fateryga, Valentina V.</mods:namePart>
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<mods:namePart id="770DD7012C300D52F01D121A0A5FFC02">Efimov, Petr G.</mods:namePart>
<mods:affiliation id="481DC28913D518EE14827D87A1F80E17">Komarov Botanical Institute of the Russian Academy of Sciences, Prof. Popov str. 2, 197376 St. Petersburg, Russia</mods:affiliation>
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<heading id="D0F79A22EB3C273A0AC6FCF3FB5BFC83" bold="true" box="[136,1158,780,806]" fontSize="11" level="1" pageId="6" pageNumber="28" reason="1">
<emphasis id="B974F15CEB3C273A0AC6FCF3FB5BFC83" bold="true" box="[136,1158,780,806]" pageId="6" pageNumber="28">
Key to the species of
<taxonomicName id="4C0056CDEB3C273A0BCCFCF3FD47FC83" authority="sect. Epipactis" box="[386,666,780,806]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="section" section="Epipactis">
<emphasis id="B974F15CEB3C273A0BCCFCF3FE30FC83" bold="true" box="[386,493,780,806]" italics="true" pageId="6" pageNumber="28">Epipactis</emphasis>
sect.
<emphasis id="B974F15CEB3C273A0861FCF3FD47FC83" bold="true" box="[559,666,780,806]" italics="true" pageId="6" pageNumber="28">Epipactis</emphasis>
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in the
<collectingRegion id="49C4E3ACEB3C273A08A2FCF3FC9BFC83" box="[748,838,780,806]" country="Ukraine" name="Crimea" pageId="6" pageNumber="28">Crimea</collectingRegion>
and the Russian Caucasus
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<keyLead id="30F18E7BEB3C273A0AC6FCAEFA76FCC3" box="[136,1451,848,870]" pageId="6" pageNumber="28">1. Rachis of inflorescence densely pubescent ........................................................................................................................................2</keyLead>
</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6FC91FA76FC27" blockId="6.[136,1451,848,1706]" box="[136,1451,877,899]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FC91FA76FC27" box="[136,1451,877,899]" pageId="6" pageNumber="28">- Rachis of inflorescence glabrous or subglabrous (minute pubescence could sometimes be visible in a microscope) ....................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F435EBEB3C273A0AC6FC75FA76FC7F" pageId="6" pageNumber="28">
<paragraph id="8BBF2D4EEB3C273A0AC6FC75FA76FC3A" blockId="6.[136,1451,848,1706]" box="[136,1451,906,928]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FC75FA76FC3A" box="[136,1451,906,928]" pageId="6" pageNumber="28">2. Ovary strongly pubescent same as rachis of inflorescence; leaves usually ± equal in length to the internodes................................3</keyLead>
</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6FC58FA76FC7F" blockId="6.[136,1451,848,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FC58FA76FC7F" pageId="6" pageNumber="28">- Ovaries glabrous or significantly more sparsely pubescent than the rachis of inflorescence; leaves usually much longer than the internodes ...........................................................................................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F435EBEB3C273A0AC6FC1EFA76FBB7" pageId="6" pageNumber="28">
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<keyLead id="30F18E7BEB3C273A0AC6FC1EFA76FC50" box="[136,1451,992,1014]" pageId="6" pageNumber="28">
3. Epichile green, with one big, irregular, strongly crenate boss near the base; sepals shorter than
<quantity id="4CF880ABEB3C273A0E11FC1EFB4BFC53" box="[1119,1174,993,1014]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="28" unit="mm" value="7.0">7 mm</quantity>
....................
<taxonomicName id="4C0056CDEB3C273A0F53FC1EFA76FC50" ID-CoL="6G7KR" authorityName="Swartz" authorityYear="1800" baseAuthorityName="Ehrhart" baseAuthorityYear="1789" box="[1309,1451,992,1014]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="microphylla">
<emphasis id="B974F15CEB3C273A0F53FC1EFA76FC50" box="[1309,1451,992,1014]" italics="true" pageId="6" pageNumber="28">E. microphylla</emphasis>
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- Epichile greenish-white to bright-pink, with two separate bosses near the base; sepals longer than
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......................................4
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<keyLead id="30F18E7BEB3C273A0AC6FBE5FA76FB8A" box="[136,1451,1050,1072]" pageId="6" pageNumber="28">
4. Viscidium absent; epichile greenish-white to dirty-pink...........................................................................................
<taxonomicName id="4C0056CDEB3C273A0F42FBE4FA76FB8A" ID-CoL="6G7KT" authorityName="Kreutz, Fateryga &amp; Efimov" authorityYear="2014" box="[1292,1451,1050,1072]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="krymmontana">
<emphasis id="B974F15CEB3C273A0F42FBE4FA76FB8A" box="[1292,1451,1050,1072]" italics="true" pageId="6" pageNumber="28">E. krymmontana</emphasis>
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<paragraph id="8BBF2D4EEB3C273A0AC6FBC8FA76FBE8" blockId="6.[136,1451,848,1706]" box="[136,1451,1079,1101]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FBC8FA76FBE8" box="[136,1451,1079,1101]" pageId="6" pageNumber="28">- Viscidium present; epichile pale-pink to bright-pink .........................................................................................................................5</keyLead>
</paragraph>
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<keyLead id="30F18E7BEB3C273A0AC6FBABFA77FBCC" box="[136,1450,1108,1130]" pageId="6" pageNumber="28">
5. Leaves green, not suffused purple, elliptic, rarely lanceolate ........................................................
<taxonomicName id="4C0056CDEB3C273A0E23FBABFA77FBCC" ID-CoL="7YHPD" authority="subsp. condensata" box="[1133,1450,1108,1130]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="subSpecies" species="condensata" subSpecies="condensata">
<emphasis id="B974F15CEB3C273A0E23FBABFB29FBCC" box="[1133,1268,1108,1129]" italics="true" pageId="6" pageNumber="28">E. condensata</emphasis>
subsp.
<emphasis id="B974F15CEB3C273A0F73FBABFA77FBCC" box="[1341,1450,1108,1129]" italics="true" pageId="6" pageNumber="28">condensata</emphasis>
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</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6FB8EFA76FB20" blockId="6.[136,1451,848,1706]" box="[136,1451,1136,1158]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FB8EFA76FB20" box="[136,1451,1136,1158]" pageId="6" pageNumber="28">
- Leaves suffused by purple, ovate, rarely elliptic...........................................................................
<taxonomicName id="4C0056CDEB3C273A0E2CFB8EFA76FB20" ID-CoL="7YHPF" authorityName="Kreutz, Fateryga &amp; Efimov" authorityYear="2014" baseAuthorityName="Akhalk., H. Baumann, R. Lorenz &amp; Mosul." baseAuthorityYear="2014" box="[1122,1451,1136,1158]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="subSpecies" species="condensata" subSpecies="kuenkeleana">
<emphasis id="B974F15CEB3C273A0E2CFB8EFB37FB20" box="[1122,1258,1136,1158]" italics="true" pageId="6" pageNumber="28">E. condensata</emphasis>
subsp.
<emphasis id="B974F15CEB3C273A0F7DFB8FFA76FB20" box="[1331,1451,1136,1157]" italics="true" pageId="6" pageNumber="28">kuenkeleana</emphasis>
</taxonomicName>
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</keyStep>
<keyStep id="30F435EBEB3C273A0AC6FB72FA76FB5C" pageId="6" pageNumber="28">
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<keyLead id="30F18E7BEB3C273A0AC6FB72FA76FB1A" pageId="6" pageNumber="28">
6. Viscidium absent, pollinia touching the stigma even in young flowers; epichile pinkish-green, ± flat or with two indistinct small bosses near the base; petals whitish-green; pedicel completely green ...............................................................................
<taxonomicName id="4C0056CDEB3C273A0F0EFB54FA76FB1A" ID-CoL="6G7L3" authorityName="Godfery" authorityYear="1921" box="[1344,1451,1194,1216]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="muelleri">
<emphasis id="B974F15CEB3C273A0F0EFB54FA76FB1A" box="[1344,1451,1194,1216]" italics="true" pageId="6" pageNumber="28">E. muelleri</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6FB38FA76FB5C" blockId="6.[136,1451,848,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FB38FA76FB5C" pageId="6" pageNumber="28">- Viscidium present at least in young flowers, pollinia not touching the stigma; epichile usually with two large bosses near the base, if bosses absent, then the epichile is green; petals of various coloration; pedicel purplish at least at the base ........................7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F435EBEB3C273A0AC6FAFEFA76FAC9" pageId="6" pageNumber="28">
<paragraph id="8BBF2D4EEB3C273A0AC6FAFEFA76FA96" blockId="6.[136,1451,848,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FAFEFA76FA96" pageId="6" pageNumber="28">
7. Epichile completely green, ± flat or with two indistinct small bosses near the base; sepals
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long; viscidium present only in young flowers ..................................................................................................................................................................
<taxonomicName id="4C0056CDEB3C273A0F06FAE1FA76FA96" ID-CoL="6FVKJ" authorityName="Taubenheim" authorityYear="1975" box="[1352,1451,1310,1331]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="pontica">
<emphasis id="B974F15CEB3C273A0F06FAE1FA76FA96" box="[1352,1451,1310,1331]" italics="true" pageId="6" pageNumber="28">E. pontica</emphasis>
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- Epichile whitish-pink to bright-pink or crimson, with two large bosses near the base; sepals
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long; viscidium usually present not only in young flowers ......................................................................................................................................................8
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8. Leaves concentrated mainly at the base of the stem; inflorescence occupies about 2/3 of the stem; epichile usually with yellow longitudinal stripe................................................................................................................................................................
<taxonomicName id="4C0056CDEB3C273A0F03FA6EFA76FA03" ID-CoL="7VZ2N" authorityName="Kreutz" authorityYear="1997" box="[1357,1451,1425,1446]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="turcica">
<emphasis id="B974F15CEB3C273A0F03FA6EFA76FA03" box="[1357,1451,1425,1446]" italics="true" pageId="6" pageNumber="28">E. turcica</emphasis>
</taxonomicName>
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<paragraph id="8BBF2D4EEB3C273A0AC6FA51FA76FA7A" blockId="6.[136,1451,848,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6FA51FA76FA7A" pageId="6" pageNumber="28">- Leaves scattered regularly along the stem; inflorescence occupies about 1/3 of the stem; epichile without yellow longitudinal stripe ...................................................................................................................................................................................................9</keyLead>
</paragraph>
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<keyLead id="30F18E7BEB3C273A0AC6FA18FA76FA59" box="[136,1451,1511,1533]" pageId="6" pageNumber="28">
9. Epichile pale pink to pink, paler than the petals; leaves ± flat .......................................................
<taxonomicName id="4C0056CDEB3C273A0E21FA17FA76FA59" ID-CoL="7YHQB" authorityName="Klein" authorityYear="1997" baseAuthorityName="Richter" baseAuthorityYear="1887" box="[1135,1451,1511,1533]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="subSpecies" species="helleborine" subSpecies="orbicularis">
<emphasis id="B974F15CEB3C273A0E21FA17FB2AFA59" box="[1135,1271,1511,1533]" italics="true" pageId="6" pageNumber="28">E. helleborine</emphasis>
subsp.
<emphasis id="B974F15CEB3C273A0F0EFA18FA76FA59" box="[1344,1451,1511,1532]" italics="true" pageId="6" pageNumber="28">orbicularis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6F9FBFA76F9BC" blockId="6.[136,1451,848,1706]" box="[136,1451,1540,1562]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6F9FBFA76F9BC" box="[136,1451,1540,1562]" pageId="6" pageNumber="28">- Epichile pale pink to bright-pink and crimson, not paler and usually brighter than the petals; leaves ± arching............................10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F435EBEB3C273A0AC6F9DEFA77F9F7" pageId="6" pageNumber="28">
<paragraph id="8BBF2D4EEB3C273A0AC6F9DEFA77F990" blockId="6.[136,1451,848,1706]" box="[136,1450,1568,1590]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6F9DEFA77F990" box="[136,1450,1568,1590]" pageId="6" pageNumber="28">
10. Epichile brighter than petals; medial leaves elliptic to elliptically lanceolate ...................................
<taxonomicName id="4C0056CDEB3C273A0ECFF9DEFA77F990" ID-CoL="7YHQ2" authorityName="Kreutz, Ovari &amp; Shifman" authorityYear="2011" box="[1153,1450,1568,1590]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="subSpecies" species="helleborine" subSpecies="levantina">
<emphasis id="B974F15CEB3C273A0ECFF9DEFAD5F990" box="[1153,1288,1568,1590]" italics="true" pageId="6" pageNumber="28">E. helleborine</emphasis>
subsp.
<emphasis id="B974F15CEB3C273A0F1FF9DFFA77F990" box="[1361,1450,1568,1589]" italics="true" pageId="6" pageNumber="28">levantina</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6F9C1FA77F9F7" blockId="6.[136,1451,848,1706]" box="[136,1450,1597,1619]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6F9C1FA77F9F7" box="[136,1450,1597,1619]" pageId="6" pageNumber="28">
- Epichile ± same in colour as petals; medial leaves ovate to elliptic ...............................................
<taxonomicName id="4C0056CDEB3C273A0E23F9C1FA77F9F7" ID-CoL="7YHPV" authority="subsp. helleborine" box="[1133,1450,1597,1619]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="subSpecies" species="helleborine" subSpecies="helleborine">
<emphasis id="B974F15CEB3C273A0E23F9C1FB29F9F7" box="[1133,1268,1597,1619]" italics="true" pageId="6" pageNumber="28">E. helleborine</emphasis>
subsp.
<emphasis id="B974F15CEB3C273A0F73F9C2FA77F9F7" box="[1341,1450,1597,1618]" italics="true" pageId="6" pageNumber="28">helleborine</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30F435EBEB3C273A0AC6F9A5FA76F90C" pageId="6" pageNumber="28">
<paragraph id="8BBF2D4EEB3C273A0AC6F9A5FA77F929" blockId="6.[136,1451,848,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6F9A5FA77F929" pageId="6" pageNumber="28">
11. Leaves erect; stem completely green or purplish at the base; pedicel sometimes completely green; flowers ± campanulate ............ ..............................................................................................................................................................................................
<taxonomicName id="4C0056CDEB3C273A0F05F987FA77F929" ID-CoL="6FVLV" authorityName="Fateryga &amp; Kreutz" authorityYear="2012" box="[1355,1450,1655,1677]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="taurica">
<emphasis id="B974F15CEB3C273A0F05F987FA77F929" box="[1355,1450,1655,1677]" italics="true" pageId="6" pageNumber="28">E. taurica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BBF2D4EEB3C273A0AC6F96BFA76F90C" blockId="6.[136,1451,848,1706]" box="[136,1451,1684,1706]" pageId="6" pageNumber="28">
<keyLead id="30F18E7BEB3C273A0AC6F96BFA76F90C" box="[136,1451,1684,1706]" pageId="6" pageNumber="28">
- Leaves arching; stem always purplish at the base; pedicel always purple at least at the base; tepals ± spreading.............
<taxonomicName id="4C0056CDEB3C273A0F07F96BFA76F90C" ID-CoL="6FVKK" authorityName="Nannfeldt" authorityYear="1946" baseAuthorityName="Soo" baseAuthorityYear="1927" box="[1353,1451,1684,1705]" class="Liliopsida" family="Orchidaceae" genus="Epipactis" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="28" phylum="Tracheophyta" rank="species" species="persica">
<emphasis id="B974F15CEB3C273A0F07F96BFA76F90C" box="[1353,1451,1684,1705]" italics="true" pageId="6" pageNumber="28">E. persica</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>