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<document id="E8B94CD6E23EB01FF9F2ECF6053C2176" ID-DOI="10.11646/zootaxa.5419.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="10791732" ID-ZooBank="AE1721AA-277C-4699-9A53-9A8A8D982C04" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709794774810" checkinUser="plazi" docAuthor="Pal, Gouri Sankar &amp; Hazra, Niladri" docDate="2024" docId="03A90C56FF87FFE3FF4EF90716445C26" docLanguage="en" docName="zootaxa.5419.3.2.pdf" docOrigin="Zootaxa 5419 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Forcipomyia (Lepidohelea) sulcoa Pal &amp; Hazra 2024, sp. n." docType="treatment" docUuid="B9C7E98F-F2DE-4AA4-9B2F-CC674204CFAB" docUuidSource="ZooBank" docVersion="2" lastPageNumber="356" masterDocId="FF90742EFF80FFEBFFD9FFC0151B5D3B" masterDocTitle="Three new species of Forcipomyia (Lepidohelea) (Diptera: Ceratopogonidae) from West Bengal, India" masterLastPageNumber="360" masterPageNumber="348" pageNumber="355" updateTime="1709795180797" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F4C9E9B7BD568E6BE0788CDD8D8085AE">Three new species of Forcipomyia (Lepidohelea) (Diptera: Ceratopogonidae) from West Bengal, India</mods:title>
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<mods:name id="6A1E0772DA49C7673DA96D4878D41348" type="personal">
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<mods:namePart id="8E76B5A9AD1D889E67CE27410F725095">Pal, Gouri Sankar</mods:namePart>
<mods:nameIdentifier id="8936765DA30B0502793D22687D9B3FF1" type="ORCID">0000-0001-6124-0479</mods:nameIdentifier>
<mods:affiliation id="15DAF92695486919FA69E19AEC1872A1">Biosystematics &amp; Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, India, 713104</mods:affiliation>
<mods:nameIdentifier id="C4E79791736AE69222731E39D277CEC2" type="email">gsankarpal9@gmail.com</mods:nameIdentifier>
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<mods:name id="A757E03E99961E37DBB705E84EFBF08C" type="personal">
<mods:role id="6F72EA2A833873B499B280F653727B47">
<mods:roleTerm id="2C5C94F8FD0A8D97E7DB1021AACEB99E">Author</mods:roleTerm>
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<mods:namePart id="DC304F23B6D349535424DCA816F56F74">Hazra, Niladri</mods:namePart>
<mods:nameIdentifier id="A4B15CA920DF27BC8D04A0DE0DFEBC20" type="ORCID">0000-0001-8149-9445</mods:nameIdentifier>
<mods:affiliation id="B315DE75EF5E453B7800B103BBB6C4AA">Biosystematics &amp; Entomology Research Unit, Department of Zoology, The University of Burdwan, Burdwan, India, 713104</mods:affiliation>
<mods:nameIdentifier id="7DF3AE085B3D99FA7C01A808BA8E0DF0" type="email">nhazra@zoo.buruniv.ac.in</mods:nameIdentifier>
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<mods:title id="11027A0CC128963BC6C55A4EDC6C550F">Zootaxa</mods:title>
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<mods:part id="8A5708FCBD7B59812C12CD62AFC775E9">
<mods:date id="BCB6D9B893100D826160C3F9A85EBA8F">2024</mods:date>
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<mods:number id="2051B4E0CBB1CBCF7290E1ED7E051AA5">2024-03-07</mods:number>
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<mods:number id="488DE7EAA656E552CEE2F7BF57C81460">5419</mods:number>
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<mods:identifier id="EF6933870805232F6EE882419EB484AE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A90C56FF87FFE3FF4EF90716445C26" ID-DOI="http://doi.org/10.5281/zenodo.10791746" ID-Zenodo-Dep="10791746" LSID="urn:lsid:zoobank.org:act:B9C7E98F-F2DE-4AA4-9B2F-CC674204CFAB" httpUri="http://treatment.plazi.org/id/03A90C56FF87FFE3FF4EF90716445C26" lastPageId="8" lastPageNumber="356" pageId="7" pageNumber="355">
<subSubSection id="C31AEECBFF87FFECFF4EF90716C35A3E" pageId="7" pageNumber="355" type="nomenclature">
<paragraph id="8BBFBD40FF87FFECFF4EF907179B5BDA" blockId="7.[151,984,1734,1833]" box="[151,640,1734,1761]" pageId="7" pageNumber="355">
<heading id="D0F70A2CFF87FFECFF4EF907179B5BDA" bold="true" box="[151,640,1734,1761]" fontSize="11" level="1" pageId="7" pageNumber="355" reason="1">
<emphasis id="B9746152FF87FFECFF4EF907179B5BDA" bold="true" box="[151,640,1734,1761]" pageId="7" pageNumber="355">
<taxonomicName id="4C00C6C3FF87FFECFF4EF907172D5BDA" authority="Pal &amp; Hazra, 2024" authorityName="Pal &amp; Hazra" authorityYear="2024" box="[151,566,1734,1761]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="355" phylum="Arthropoda" rank="species" species="sulcoa" status="sp. nov." subGenus="Lepidohelea">
<emphasis id="B9746152FF87FFECFF4EF90714295BDA" bold="true" box="[151,306,1735,1761]" italics="true" pageId="7" pageNumber="355">Forcipomyia</emphasis>
(
<emphasis id="B9746152FF87FFECFE9AF90614CC5BDA" bold="true" box="[323,471,1734,1761]" italics="true" pageId="7" pageNumber="355">Lepidohelea</emphasis>
)
<emphasis id="B9746152FF87FFECFE31F906172D5BDA" bold="true" box="[488,566,1734,1761]" italics="true" pageId="7" pageNumber="355">sulcoa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A247DC29FF87FFECFDE7F907179B5BDA" box="[574,640,1735,1761]" pageId="7" pageNumber="355" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBFBD40FF87FFECFF4EF92B16C35A3E" blockId="7.[151,984,1734,1833]" box="[151,984,1771,1797]" pageId="7" pageNumber="355">
<uri id="FF91B142FF87FFECFF4EF92B16C35A3E" box="[151,984,1771,1797]" pageId="7" pageNumber="355">
urn:lsid:zoobank.org:act:
<uuid id="FFA68795FF87FFECFE76F92B16C35A3E" box="[431,984,1771,1797]" pageId="7" pageNumber="355">B9C7E98F-F2DE-4AA4-9B2F-CC674204CFAB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C31AEECBFF87FFECFF4EF8CF14365A12" box="[151,301,1807,1833]" pageId="7" pageNumber="355" type="description">
<paragraph id="8BBFBD40FF87FFECFF4EF8CF14365A12" blockId="7.[151,984,1734,1833]" box="[151,301,1807,1833]" pageId="7" pageNumber="355">
(
<figureCitation id="133BA1C5FF87FFECFF78F8CF143E5A12" box="[161,293,1807,1833]" captionStart="FIGURE 4" captionStartId="9.[152,255,1841,1865]" captionTargetBox="[205,1383,191,1808]" captionTargetId="figure-28@9.[196,1392,181,1817]" captionTargetPageId="9" captionText="FIGURE 4. Forcipomyia (Lepidohelea) sulcoa sp. n. AI Male (♂): A, Antenna; B, Maxillary palpus; C, Wing; D, Banding patterns of legs (femur, tibia and tarsomeres); E, Hind tibial comb; F, Parameres; G, Aedeagus; H, Photograph of male genitalia (dorsal view); I, Illustration of male genitalia (dorsal view); JO Female (♀): J, Antenna K, Maxillary palpus; L, Wing; M, Hind tibial comb; N, Abdomen; O, Spermatheca. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10791744" httpUri="https://zenodo.org/record/10791744/files/figure.png" pageId="7" pageNumber="355">Figs. 4AO</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31AEECBFF87FFECFF4EF897141E5AAE" pageId="7" pageNumber="355" type="etymology">
<paragraph id="8BBFBD40FF87FFECFF4EF897141E5AAE" blockId="7.[151,1436,1879,2013]" pageId="7" pageNumber="355">
<emphasis id="B9746152FF87FFECFF4EF89714055A4A" bold="true" box="[151,286,1879,1905]" pageId="7" pageNumber="355">Etymology.</emphasis>
The specific name
<taxonomicName id="4C00C6C3FF87FFECFDDFF89717475A4A" authorityName="Pal &amp; Hazra" authorityYear="2024" box="[518,604,1879,1905]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="355" phylum="Arthropoda" rank="species" species="sulcoa">
<emphasis id="B9746152FF87FFECFDCAF89717475A4A" box="[531,604,1879,1905]" italics="true" pageId="7" pageNumber="355">sulcoa</emphasis>
</taxonomicName>
” derives from a Latin word, indicating “a small groove” present in the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C31AEECBFF87FFE3FF1EF85E11FA5DEE" lastPageId="8" lastPageNumber="356" pageId="7" pageNumber="355" type="diagnosis">
<paragraph id="8BBFBD40FF87FFE3FF1EF85E11FA5DEE" blockId="7.[151,1436,1879,2013]" lastBlockId="8.[151,1436,151,285]" lastPageId="8" lastPageNumber="356" pageId="7" pageNumber="355">
<emphasis id="B9746152FF87FFECFF1EF85E14595A83" bold="true" box="[199,322,1950,1976]" pageId="7" pageNumber="355">Diagnosis.</emphasis>
Only species of
<taxonomicName id="4C00C6C3FF87FFECFDDFF85F16275A82" authorityName="Kieffer" authorityYear="1917" box="[518,828,1951,1977]" class="Insecta" family="Ceratopogonidae" genus="Forcipomyia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="355" phylum="Arthropoda" rank="subGenus" subGenus="Lepidohelea">
<emphasis id="B9746152FF87FFECFDDFF85F178D5A83" box="[518,662,1951,1976]" italics="true" pageId="7" pageNumber="355">Forcipomyia</emphasis>
(
<emphasis id="B9746152FF87FFECFD71F85F16295A82" box="[680,818,1951,1977]" italics="true" pageId="7" pageNumber="355">Lepidohelea</emphasis>
)
</taxonomicName>
from the
<collectingRegion id="49C473A2FF87FFECFC68F85F11165A82" box="[945,1037,1951,1977]" country="Democratic Republic of the Congo" name="Orientale" pageId="7" pageNumber="355">Oriental</collectingRegion>
realm possesses flattened, roughly square shaped aedeagus with a notch at middle of apical margin (
<figureCitation id="133BA1C5FF87FFECFC70F80311135AE6" box="[937,1032,1987,2013]" captionStart="FIGURE 4" captionStartId="9.[152,255,1841,1865]" captionTargetBox="[205,1383,191,1808]" captionTargetId="figure-28@9.[196,1392,181,1817]" captionTargetPageId="9" captionText="FIGURE 4. Forcipomyia (Lepidohelea) sulcoa sp. n. AI Male (♂): A, Antenna; B, Maxillary palpus; C, Wing; D, Banding patterns of legs (femur, tibia and tarsomeres); E, Hind tibial comb; F, Parameres; G, Aedeagus; H, Photograph of male genitalia (dorsal view); I, Illustration of male genitalia (dorsal view); JO Female (♀): J, Antenna K, Maxillary palpus; L, Wing; M, Hind tibial comb; N, Abdomen; O, Spermatheca. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10791744" httpUri="https://zenodo.org/record/10791744/files/figure.png" pageId="7" pageNumber="355">Fig. 4G</figureCitation>
); submedian process of parameres moderately long and a little outwardly directed (
<figureCitation id="133BA1C5FF88FFE3FD77FF5717E55D8A" box="[686,766,151,177]" captionStart="FIGURE 4" captionStartId="9.[152,255,1841,1865]" captionTargetBox="[205,1383,191,1808]" captionTargetId="figure-28@9.[196,1392,181,1817]" captionTargetPageId="9" captionText="FIGURE 4. Forcipomyia (Lepidohelea) sulcoa sp. n. AI Male (♂): A, Antenna; B, Maxillary palpus; C, Wing; D, Banding patterns of legs (femur, tibia and tarsomeres); E, Hind tibial comb; F, Parameres; G, Aedeagus; H, Photograph of male genitalia (dorsal view); I, Illustration of male genitalia (dorsal view); JO Female (♀): J, Antenna K, Maxillary palpus; L, Wing; M, Hind tibial comb; N, Abdomen; O, Spermatheca. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10791744" httpUri="https://zenodo.org/record/10791744/files/figure.png" pageId="8" pageNumber="356">Fig. 4F</figureCitation>
); female with a large irregular sensory pit on third maxillary palp segment (
<figureCitation id="133BA1C5FF88FFE3FEE5FF7B148D5DEE" box="[316,406,187,213]" captionStart="FIGURE 4" captionStartId="9.[152,255,1841,1865]" captionTargetBox="[205,1383,191,1808]" captionTargetId="figure-28@9.[196,1392,181,1817]" captionTargetPageId="9" captionText="FIGURE 4. Forcipomyia (Lepidohelea) sulcoa sp. n. AI Male (♂): A, Antenna; B, Maxillary palpus; C, Wing; D, Banding patterns of legs (femur, tibia and tarsomeres); E, Hind tibial comb; F, Parameres; G, Aedeagus; H, Photograph of male genitalia (dorsal view); I, Illustration of male genitalia (dorsal view); JO Female (♀): J, Antenna K, Maxillary palpus; L, Wing; M, Hind tibial comb; N, Abdomen; O, Spermatheca. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10791744" httpUri="https://zenodo.org/record/10791744/files/figure.png" pageId="8" pageNumber="356">Fig. 4K</figureCitation>
); single spermatheca with a neck and without hyaline punctation (
<figureCitation id="133BA1C5FF88FFE3FBA2FF7B11CF5DEE" box="[1147,1236,187,213]" captionStart="FIGURE 4" captionStartId="9.[152,255,1841,1865]" captionTargetBox="[205,1383,191,1808]" captionTargetId="figure-28@9.[196,1392,181,1817]" captionTargetPageId="9" captionText="FIGURE 4. Forcipomyia (Lepidohelea) sulcoa sp. n. AI Male (♂): A, Antenna; B, Maxillary palpus; C, Wing; D, Banding patterns of legs (femur, tibia and tarsomeres); E, Hind tibial comb; F, Parameres; G, Aedeagus; H, Photograph of male genitalia (dorsal view); I, Illustration of male genitalia (dorsal view); JO Female (♀): J, Antenna K, Maxillary palpus; L, Wing; M, Hind tibial comb; N, Abdomen; O, Spermatheca. Scale bars 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.10791744" httpUri="https://zenodo.org/record/10791744/files/figure.png" pageId="8" pageNumber="356">Fig. 4O</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31AEECBFF88FFE3FF1EFF1F16445C26" pageId="8" pageNumber="356" type="materials_examined">
<paragraph id="8BBFBD40FF88FFE3FF1EFF1F16445C26" blockId="8.[151,1436,151,285]" pageId="8" pageNumber="356">
<emphasis id="B9746152FF88FFE3FF1EFF1F14635DC3" bold="true" box="[199,376,222,249]" pageId="8" pageNumber="356">Type Material.</emphasis>
<materialsCitation id="3B68B71DFF88FFE3FEA7FF1F14295C26" collectingDate="2023-05-08" collectorName="Col. G. S. Pal." country="India" latitude="27.058592" location="Paren" longLatPrecision="1" longitude="88.864914" municipality="Kalimpong" pageId="8" pageNumber="356" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="holotype">
<typeStatus id="54BB03E2FF88FFE3FEA7FF1F14FD5DC2" box="[382,486,223,249]" pageId="8" pageNumber="356" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0676C9FF88FFE3FE34FF1F171F5DC2" box="[493,516,223,249]" count="1" pageId="8" pageNumber="356" type="male"></specimenCount>
,
<collectingCountry id="F317FDD0FF88FFE3FDD6FF1F17505DC2" box="[527,587,223,249]" name="India" pageId="8" pageNumber="356">India</collectingCountry>
,
<collectingRegion id="49C473A2FF88FFE3FD8FFF1F17FD5DC2" box="[598,742,223,249]" country="India" name="West Bengal" pageId="8" pageNumber="356">West Bengal</collectingRegion>
,
<collectingMunicipality id="6BDB273AFF88FFE3FD28FF1F16695DC2" box="[753,882,223,249]" pageId="8" pageNumber="356">Kalimpong</collectingMunicipality>
,
<location id="8EDFEB9BFF88FFE3FCA5FF1F16A75DC2" LSID="urn:lsid:plazi:treatment:03A90C56FF87FFE3FF4EF90716445C26:8EDFEB9BFF88FFE3FCA5FF1F16A75DC2" box="[892,956,223,249]" country="India" latitude="27.058592" longLatPrecision="1" longitude="88.864914" municipality="Kalimpong" name="Paren" pageId="8" pageNumber="356" stateProvince="West Bengal">Paren</location>
(
<geoCoordinate id="EE34DB87FF88FFE3FC13FF1E11705DC2" box="[970,1131,222,249]" degrees="27.058592" direction="north" orientation="latitude" pageId="8" pageNumber="356" precision="1" value="27.058592">27.058592 °N</geoCoordinate>
,
<geoCoordinate id="EE34DB87FF88FFE3FBAFFF1E10085DC2" box="[1142,1299,222,249]" degrees="88.864915" direction="east" orientation="longitude" pageId="8" pageNumber="356" precision="1" value="88.864914">88.864915 °E</geoCoordinate>
),
<date id="FFBE9B80FF88FFE3FAFEFF1F10835DC2" box="[1319,1432,223,249]" pageId="8" pageNumber="356" value="2023-05-08">
<collectingDate id="EFFA6268FF88FFE3FAFEFF1F10835DC2" box="[1319,1432,223,249]" pageId="8" pageNumber="356" value="2023-05-08">08.v.2023</collectingDate>
</date>
,
<collectorName id="26F5D896FF88FFE3FF4EFEC314295C26" box="[151,306,259,285]" pageId="8" pageNumber="356">Col. G.S. Pal.</collectorName>
</materialsCitation>
<materialsCitation id="3B68B71DFF88FFE3FEE1FEC316405C26" box="[312,859,259,285]" collectingDate="2023-05-08" collectorName="Col. G. S. Pal." country="India" latitude="27.058592" location="Paren" longLatPrecision="1" longitude="88.864914" municipality="Kalimpong" pageId="8" pageNumber="356" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="West Bengal" typeStatus="paratype">
<typeStatus id="54BB03E2FF88FFE3FEE1FEC314BE5C26" box="[312,421,259,285]" pageId="8" pageNumber="356" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D0676C9FF88FFE3FE75FEC314FF5C26" box="[428,484,259,285]" count="3" pageId="8" pageNumber="356" type="male">3♂♂</specimenCount>
and
<specimenCount id="9D0676C9FF88FFE3FDC2FEC3174F5C26" box="[539,596,259,285]" count="2" pageId="8" pageNumber="356" type="female">2♀♀</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>