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<document id="414F803BB2CD4535169FA7F0D4ED37DD" ID-DOI="10.11646/zootaxa.2458.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316658" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629675028716" checkinUser="felipe" docAuthor="Anderson, Robert S." docDate="2010" docId="03E88799FFABFFF72CB4FDEF487B1BFE" docLanguage="en" docName="zt02458p127.pdf" docOrigin="Zootaxa 2458 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2458.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Theognete denticulata Anderson 2010, new species" docType="treatment" docUuid="473AFD17-2AC2-4C97-B8AB-CDA916F0154D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="24" masterDocId="FFD1FFE1FFBCFFE02D59FFD84C3D1E42" masterDocTitle="A taxonomic monograph of the Middle American leaf-litter inhabiting weevil genus Theognete Champion (Coleoptera: Curculionidae; Molytinae; Lymantini) 2458" masterLastPageNumber="127" masterPageNumber="1" pageNumber="24" updateTime="1699094135467" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="19EA9AB6ED147DFCA783D2C580EA9C42">A taxonomic monograph of the Middle American leaf-litter inhabiting weevil genus Theognete Champion (Coleoptera: Curculionidae; Molytinae; Lymantini) 2458</mods:title>
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<subSubSection id="C35B6504FFABFFF72CB4FDEF4FFA1C36" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BFE368FFFABFFF72CB4FDEF4FFA1C36" blockId="23.[493,1093,567,628]" pageId="23" pageNumber="24">
<emphasis id="B935EA9DFFABFFF72CB4FDEF48781C13" bold="true" box="[493,1093,567,593]" pageId="23" pageNumber="24">
11.
<taxonomicName id="4C414D0CFFABFFF72F41FDEF4F951C13" authority="Anderson" authorityName="Anderson" authorityYear="2010" box="[536,936,567,593]" class="Insecta" family="Curculionidae" genus="Theognete" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B935EA9DFFABFFF72F41FDEF4F181C13" bold="true" box="[536,805,567,593]" italics="true" pageId="23" pageNumber="24">Theognete denticulata</emphasis>
Anderson
</taxonomicName>
,
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</emphasis>
(
<figureCitation id="137A2A0AFFABFFF72F2AFD824EC91C36" box="[627,756,602,628]" captionStart="FIGURES 5963" captionStartId="95.[151,269,1435,1459]" captionTargetBox="[290,1296,211,1403]" captionTargetId="figure-40@95.[285,1301,194,1411]" captionTargetPageId="95" captionText="FIGURES 5963. Habitus, dorsal view, Theognete denticulata species group. 59. T. caviventris. 60. T. thibodeaui. 61. T. fulgida. 62. T. denticulata. 63. T. stefurinoi." figureDoi="http://doi.org/10.5281/zenodo.5362249" httpUri="https://zenodo.org/record/5362249/files/figure.png" pageId="23" pageNumber="24">Figures 62</figureCitation>
,
<figureCitation id="137A2A0AFFABFFF72E59FD824F1D1C36" box="[768,800,602,628]" captionStart="FIGURES 6467" captionStartId="96.[151,269,1950,1974]" captionTargetBox="[461,1120,204,1915]" captionTargetId="figure-43@96.[443,1144,194,1926]" captionTargetPageId="96" captionText="FIGURES 6467. Aedeagus, dorsal and lateral view, Theognete denticulata species group. 64. T. caviventris. 65. T. thibodeaui. 66. T. fulgida. 67. T. denticulata." figureDoi="http://doi.org/10.5281/zenodo.5362251" httpUri="https://zenodo.org/record/5362251/files/figure.png" pageId="23" pageNumber="24">67</figureCitation>
,
<figureCitation id="137A2A0AFFABFFF72E72FD824F761C36" box="[811,843,602,628]" captionStart="FIGURES 6872" captionStartId="97.[151,269,701,725]" captionTargetBox="[323,1267,213,661]" captionTargetId="figure-84@97.[315,1273,194,677]" captionTargetPageId="97" captionText="FIGURES 6872. Abdominal ventrite 5, male and female, ventral view, Theognete denticulata species group. 68. T. caviventris, male. 69. T. thibodeaui, male. 70. T. fulgida, male. 71. T. denticulata, male. 72. T. stefurinoi, female." figureDoi="http://doi.org/10.5281/zenodo.5362255" httpUri="https://zenodo.org/record/5362255/files/figure.png" pageId="23" pageNumber="24">71</figureCitation>
,
<figureCitation id="137A2A0AFFABFFF72E01FD824F821C36" box="[856,959,602,628]" captionStart="FIGURES 117132" captionStartId="103.[151,269,1424,1448]" captionTargetBox="[204,1373,207,1381]" captionTargetId="figure-51@103.[196,1390,194,1400]" captionTargetPageId="103" captionText="FIGURES 117132. Female genitalia (top to bottom; spermatheca, tergite VIII, sternite VIII, tergite VII), Theognete species, various species groups, representatives. 117120. T. kovariki. 121124. T. laevis. 125128. T. denticulata. 129 132. T. semistriata." figureDoi="http://doi.org/10.5281/zenodo.5362275" httpUri="https://zenodo.org/record/5362275/files/figure.png" pageId="23" pageNumber="24">125128</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35B6504FFABFFF72DCEFD724F981AE6" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BFE368FFFABFFF72DCEFD72480F1D9E" blockId="23.[151,1436,682,1468]" pageId="23" pageNumber="24">
<emphasis id="B935EA9DFFABFFF72DCEFD724D321C86" bold="true" box="[151,271,682,708]" pageId="23" pageNumber="24">Diagnosis</emphasis>
. Size. Length, male,
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; female,
<quantity id="4CB99B6AFFABFFF72E6BFD724FE51C86" box="[818,984,682,708]" metricMagnitude="-3" metricUnit="m" metricValue="4.585" metricValueMax="4.84" metricValueMin="4.33" pageId="23" pageNumber="24" unit="mm" value="4.585" valueMax="4.84" valueMin="4.33">4.334.84mm</quantity>
. Width, male,
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; female,
<quantity id="4CB99B6AFFABFFF72DCEFD0A4D031CAE" box="[151,318,722,748]" metricMagnitude="-3" metricUnit="m" metricValue="2.205" metricValueMax="2.36" metricValueMin="2.05" pageId="23" pageNumber="24" unit="mm" value="2.205" valueMax="2.36" valueMin="2.05">2.052.36mm</quantity>
. Pronotum with disk weakly cordate, scales appressed to very slightly arched and scattered; surface with isodiametric microsculpture. Elytra with disk flattened in middle, striae 1, 2 and 3 each deeply linearly impressed for a short length in this flat area; scales sparse, appressed to very slightly arched and scattered. Abdomen in male with ventrite 1 deeply longitudinally impressed, sides of impression near apical margin inwardly tuberculate. Ventrite 5 with pair of large subbasal punctures, pair of smaller median, more laterally placed punctures and apical margin deeply irregularly excavate. Legs in male with middle and hind tibiae with 35 distinct sharp teeth along inner margin. Aedeagus as in
<figureCitation id="137A2A0AFFABFFF72E81FC1A48131D9E" box="[984,1070,962,988]" captionStart="FIGURES 6467" captionStartId="96.[151,269,1950,1974]" captionTargetBox="[461,1120,204,1915]" captionTargetId="figure-43@96.[443,1144,194,1926]" captionTargetPageId="96" captionText="FIGURES 6467. Aedeagus, dorsal and lateral view, Theognete denticulata species group. 64. T. caviventris. 65. T. thibodeaui. 66. T. fulgida. 67. T. denticulata." figureDoi="http://doi.org/10.5281/zenodo.5362251" httpUri="https://zenodo.org/record/5362251/files/figure.png" pageId="23" pageNumber="24">Fig. 67</figureCitation>
.
</paragraph>
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<emphasis id="B935EA9DFFABFFF72D9FFC324E361A46" bold="true" box="[198,523,1002,1028]" pageId="23" pageNumber="24">Geographical distribution</emphasis>
.
<collectingCountry id="F356761FFFABFFF72F43FC324E4E1A46" box="[538,627,1002,1028]" name="Mexico" pageId="23" pageNumber="24">México</collectingCountry>
(
<collectingRegion id="4985F86DFFABFFF72FDAFC324EE41A46" box="[643,729,1002,1028]" country="Mexico" name="Puebla" pageId="23" pageNumber="24">Puebla</collectingRegion>
).
</paragraph>
<paragraph id="8BFE368FFFABFFF72D9FFBCA4D041A16" blockId="23.[151,1436,682,1468]" pageId="23" pageNumber="24">
<emphasis id="B935EA9DFFABFFF72D9FFBCA4DB51A6E" bold="true" box="[198,392,1042,1068]" pageId="23" pageNumber="24">Natural history</emphasis>
. Collected from berlese extraction of cloud forest leaf litter at elevations ranging from
<quantity id="4CB99B6AFFABFFF72DCEFBE24D081A16" box="[151,309,1082,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.6" metricValueMin="1.35" pageId="23" pageNumber="24" unit="m" value="1475.0" valueMax="1600.0" valueMin="1350.0">13501600m</quantity>
.
</paragraph>
<paragraph id="8BFE368FFFABFFF72D9FFBBA4F981AE6" blockId="23.[151,1436,682,1468]" pageId="23" pageNumber="24">
<emphasis id="B935EA9DFFABFFF72D9FFBBA4E211A3E" bold="true" box="[198,540,1122,1148]" pageId="23" pageNumber="24">Derivation of specific name</emphasis>
. Named from the Latin
<emphasis id="B935EA9DFFABFFF72E63FBBB4FF41A3E" box="[826,969,1123,1148]" italics="true" pageId="23" pageNumber="24">denticulatus</emphasis>
, meaning with small teeth, in reference to the distinct teeth on the inner margin of the hind tibiae of males.
</paragraph>
</subSubSection>
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<emphasis id="B935EA9DFFABFFF72D9FFB6A4D8E1A8E" bold="true" box="[198,435,1202,1228]" pageId="23" pageNumber="24">Material examined</emphasis>
.
<materialsCitation id="3B293CD2FFABFFF72C98FB6A4E851A8E" box="[449,696,1202,1228]" collectionCode="CMNC" pageId="23" pageNumber="24" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="holotype">
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,
<specimenCount id="9D47FD06FFABFFF72F50FB6A4E791A8E" box="[521,580,1202,1228]" pageId="23" pageNumber="24" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED50AE4AFFABFFF72F0AFB6A4E8D1A8E" box="[595,688,1202,1228]" pageId="23" pageNumber="24">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B293CD2FFABFFF72F9AFB6A4FFC1A8E" box="[707,961,1202,1228]" collectionCode="CMNC" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54FA882DFFABFFF72F9AFB6A4F0D1A8E" box="[707,816,1202,1228]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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(
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)
</materialsCitation>
:
<materialsCitation id="3B293CD2FFABFFF72E95FB6A4EA11AB6" collectorName="N. Teziutlan &amp; S. &amp; J. Peck" country="Mexico" location="Puebla" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Puebla" typeStatus="holotype">
<collectingCountry id="F356761FFFABFFF72E95FB6A48791A8E" box="[972,1092,1202,1228]" name="Mexico" pageId="23" pageNumber="24">MÉXICO</collectingCountry>
:
<collectingRegion id="4985F86DFFABFFF72917FB6A489F1A8E" box="[1102,1186,1202,1228]" country="Mexico" name="Puebla" pageId="23" pageNumber="24">Puebla</collectingRegion>
.
<quantity id="4CB99B6AFFABFFF729F5FB6A48C31A8E" box="[1196,1278,1202,1228]" metricMagnitude="3" metricUnit="m" metricValue="7.0811136" pageId="23" pageNumber="24" unit="mi" value="4.4">4.4 mi.</quantity>
<collectorName id="26B45359FFABFFF7285CFB6A49AA1A8E" box="[1285,1431,1202,1228]" pageId="23" pageNumber="24">N. Teziutlán</collectorName>
, 4500', 19.vi.83,
<collectorName id="26B45359FFABFFF72C0AFB024DD91AB6" box="[339,484,1242,1268]" pageId="23" pageNumber="24">S. &amp; J. Peck</collectorName>
, cloud for. litter
</materialsCitation>
/
<materialsCitation id="3B293CD2FFABFFF72FEBFB024F641B06" collectingDate="1987-07-22" collectionCode="CMNC" collectorName="R. S. Anderson. Aedeagus &amp; R. Anderson" country="Mexico" elevation="1401" location="Zacapoaxtla" municipality="Apulco" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="holotype">
<typeStatus id="54FA882DFFABFFF72FEBFB024F701AB6" box="[690,845,1242,1268]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
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,
<taxonomicName id="4C414D0CFFABFFF72E2EFB0348411AB6" authority="Anderson, 2010" authorityName="Anderson" authorityYear="2010" box="[887,1148,1243,1268]" class="Insecta" family="Curculionidae" genus="Theognete" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="denticulata" status="sp. nov.">
<emphasis id="B935EA9DFFABFFF72E2EFB0348411AB6" box="[887,1148,1243,1268]" italics="true" pageId="23" pageNumber="24">Theognete denticulata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657E6FFABFFF729DBFB0248E31AB6" box="[1154,1246,1242,1268]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectorName id="26B45359FFABFFF729B3FB024D371B5E" pageId="23" pageNumber="24">R.S. Anderson. Aedeagus</collectorName>
extracted.
<typeStatus id="54FA882DFFABFFF72CFFFADA4E201B5E" box="[422,541,1282,1308]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F356761FFFABFFF72F6CFADA4E901B5E" box="[565,685,1282,1308]" name="Mexico" pageId="23" pageNumber="24">MÉXICO</collectingCountry>
:
<collectingRegion id="4985F86DFFABFFF72F9FFADA4F271B5E" box="[710,794,1282,1308]" country="Mexico" name="Puebla" pageId="23" pageNumber="24">Puebla</collectingRegion>
:
<collectingMunicipality id="6B9AACF5FFABFFF72E6BFADA4FB51B5E" box="[818,904,1282,1308]" pageId="23" pageNumber="24">Apulco</collectingMunicipality>
(
<quantity id="4CB99B6AFFABFFF72EFFFADA48381B5E" box="[934,1029,1282,1308]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="23" pageNumber="24" unit="km" value="2.7">2.7 km</quantity>
S. near
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),
<quantity id="4CB99B6AFFABFFF7281AFADA49AA1B5E" box="[1347,1431,1282,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="23" pageNumber="24" unit="m" value="1401.0">
<elevation id="006CD1BCFFABFFF7281AFADA49AA1B5E" box="[1347,1431,1282,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.401" pageId="23" pageNumber="24" unit="m" value="1401.0">1401m</elevation>
</quantity>
,
<date id="FFFF104FFFABFFF72DCEFAF24D141B06" box="[151,297,1322,1348]" pageId="23" pageNumber="24" value="1987-07-22">
<collectingDate id="EFBBE9A7FFABFFF72DCEFAF24D141B06" box="[151,297,1322,1348]" pageId="23" pageNumber="24" value="1987-07-22">22.VII.1987</collectingDate>
</date>
,
<collectorName id="26B45359FFABFFF72C6CFAF24DF01B06" box="[309,461,1322,1348]" pageId="23" pageNumber="24">R. Anderson</collectorName>
, cloud forest, 87-36 (
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<collectionCode id="ED50AE4AFFABFFF72FACFAF24F6C1B06" box="[757,849,1322,1348]" pageId="23" pageNumber="24">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B293CD2FFABFFF72E3DFAF248A51B06" box="[868,1176,1322,1348]" collectionCode="W" country="Mexico" location="La Cumbre" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Puebla" typeStatus="paratype">
<location id="8E9E6054FFABFFF72E3DFAF24FD41B06" LSID="urn:lsid:plazi:treatment:03E88799FFABFFF72CB4FDEF487B1BFE:8E9E6054FFABFFF72E3DFAF24FD41B06" box="[868,1001,1322,1348]" country="Mexico" name="La Cumbre" pageId="23" pageNumber="24" stateProvince="Puebla">La Cumbre</location>
(
<quantity id="4CB99B6AFFABFFF72EA0FAF248711B06" box="[1017,1100,1322,1348]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="23" pageNumber="24" unit="km" value="7.6">7.6 km</quantity>
S.
<collectionCode id="ED50AE4AFFABFFF72930FAF248B61B06" box="[1129,1163,1322,1348]" pageId="23" pageNumber="24">W</collectionCode>
.)
</materialsCitation>
,
<materialsCitation id="3B293CD2FFABFFF729FCFAF24F321B2E" collectingDate="1987-07-23" collectionCode="CMNC" collectorName="Anderson" country="Mexico" elevation="1585" pageId="23" pageNumber="24" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
<quantity id="4CB99B6AFFABFFF729FCFAF248C41B06" box="[1189,1273,1322,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="23" pageNumber="24" unit="m" value="1585.0">
<elevation id="006CD1BCFFABFFF729FCFAF248C41B06" box="[1189,1273,1322,1348]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="23" pageNumber="24" unit="m" value="1585.0">1585m</elevation>
</quantity>
,
<date id="FFFF104FFFABFFF7285FFAF249AA1B06" box="[1286,1431,1322,1348]" pageId="23" pageNumber="24" value="1987-07-23">
<collectingDate id="EFBBE9A7FFABFFF7285FFAF249AA1B06" box="[1286,1431,1322,1348]" pageId="23" pageNumber="24" value="1987-07-23">23.VII.1987</collectingDate>
</date>
, R.
<collectorName id="26B45359FFABFFF72DE0FA8A4D131B2E" box="[185,302,1362,1388]" pageId="23" pageNumber="24">Anderson</collectorName>
, oak-pine forest, 87-39 (
<specimenCount id="9D47FD06FFABFFF72F17FA8A4E4C1B2E" box="[590,625,1362,1388]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
,
<specimenCount id="9D47FD06FFABFFF72F26FA8A4E991B2E" box="[639,676,1362,1388]" pageId="23" pageNumber="24" type="female">1♀</specimenCount>
<collectionCode id="ED50AE4AFFABFFF72FF2FA8A4F3A1B2E" box="[683,775,1362,1388]" pageId="23" pageNumber="24">CMNC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B293CD2FFABFFF72E40FA8A4E3C1BD6" collectingDate="1983-06-19" collectionCode="CMNC" collectorName="S. &amp; J. Peck" country="Mexico" location="Teziutlan" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Puebla" typeStatus="paratype">
<location id="8E9E6054FFABFFF72E40FA8A4FBB1B2E" LSID="urn:lsid:plazi:treatment:03E88799FFABFFF72CB4FDEF487B1BFE:8E9E6054FFABFFF72E40FA8A4FBB1B2E" box="[793,902,1362,1388]" country="Mexico" name="Teziutlan" pageId="23" pageNumber="24" stateProvince="Puebla">Teziutlán</location>
(
<quantity id="4CB99B6AFFABFFF72ECCFA8A4FD41B2E" box="[917,1001,1362,1388]" metricMagnitude="3" metricUnit="m" metricValue="7.0811136" pageId="23" pageNumber="24" unit="mi" value="4.4">4.4 mi.</quantity>
N.), 4500',
<date id="FFFF104FFFABFFF7292BFA8A48C41B2E" box="[1138,1273,1362,1388]" pageId="23" pageNumber="24" value="1983-06-19">
<collectingDate id="EFBBE9A7FFABFFF7292BFA8A48C41B2E" box="[1138,1273,1362,1388]" pageId="23" pageNumber="24" value="1983-06-19">19.VI.1983</collectingDate>
</date>
,
<collectorName id="26B45359FFABFFF7285FFA8A49A51B2E" box="[1286,1432,1362,1388]" pageId="23" pageNumber="24">S. &amp; J. Peck</collectorName>
, cloud forest litter (
<specimenCount id="9D47FD06FFABFFF72C2AFAA24DAB1BD6" box="[371,406,1402,1428]" pageId="23" pageNumber="24" type="male">1♂</specimenCount>
<collectionCode id="ED50AE4AFFABFFF72CC7FAA24DC71BD6" box="[414,506,1402,1428]" pageId="23" pageNumber="24">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE368FFFABFFF72D9FFA7A487B1BFE" blockId="23.[151,1436,682,1468]" box="[198,1094,1442,1468]" pageId="23" pageNumber="24">
<emphasis id="B935EA9DFFABFFF72D9FFA7A4E2B1BFE" bold="true" box="[198,534,1442,1468]" pageId="23" pageNumber="24">Chorological relationships.</emphasis>
Sympatric with
<taxonomicName id="4C414D0CFFABFFF72F80FA7B4FB11BFE" authorityName="Anderson" authorityYear="2010" box="[729,908,1443,1468]" class="Insecta" family="Curculionidae" genus="Theognete" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="araneiformis">
<emphasis id="B935EA9DFFABFFF72F80FA7B4FB11BFE" box="[729,908,1443,1468]" italics="true" pageId="23" pageNumber="24">T. araneiformis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C414D0CFFABFFF72E9FFA7B487D1BFE" authorityName="Anderson" authorityYear="2010" box="[966,1088,1443,1468]" class="Insecta" family="Curculionidae" genus="Theognete" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="newtoni">
<emphasis id="B935EA9DFFABFFF72E9FFA7B487D1BFE" box="[966,1088,1443,1468]" italics="true" pageId="23" pageNumber="24">T. newtoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>