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<document id="D53660F11E71EDAA7A8BCD68677BC83E" ID-DOI="10.5281/zenodo.5360042" ID-ISSN="2345-7600" ID-Zenodo-Dep="5360042" ID-ZooBank="2034ADBF-59D3-4CA1-BDB1-5C7D02A1CC4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630229305742" checkinUser="valdenar" docAuthor="Kóbor, Péter" docDate="2018" docId="03E88791FFDF270228CD5785FD9BFA49" docLanguage="en" docName="66rbz580-586.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Geocoris (Geocoris) margaretarum Kóbor 2018, new species" docType="treatment" docUuid="A1634EA4-12D0-47B1-8059-CBBE29AFC058" docUuidSource="ZooBank" docVersion="4" lastPageNumber="583" masterDocId="FFD1FFE9FFDE270128775211FFE8FFCD" masterDocTitle="Geocoris margaretarum: description of a new species from the Oriental region with remarks on allied taxa (Heteroptera: Lygaeoidea: Geocoridae)" masterLastPageNumber="586" masterPageNumber="580" pageNumber="581" updateTime="1699192735050" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="3959A4B6CDA7D9088A1C17F221A2D023">Geocoris margaretarum: description of a new species from the Oriental region with remarks on allied taxa (Heteroptera: Lygaeoidea: Geocoridae)</mods:title>
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<mods:date id="2095906824DF12ADFE1881F54CF926C3">2018</mods:date>
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<emphasis id="B935EA95FFDF270028CD5785FD3DFA66" bold="true" box="[186,725,1427,1452]" pageId="1" pageNumber="581">
<taxonomicName id="4C414D04FFDF270028CD5785FDA8FA66" authority="Kóbor, 2018" authorityName="Kóbor" authorityYear="2018" box="[186,576,1427,1452]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="581" phylum="Arthropoda" rank="species" species="margaretarum" status="sp. nov." subGenus="Geocoris">
<emphasis id="B935EA95FFDF270028CD5785FEF4FA61" bold="true" box="[186,284,1428,1452]" italics="true" pageId="1" pageNumber="581">Geocoris</emphasis>
(
<emphasis id="B935EA95FFDF2700295A5785FE67FA61" bold="true" box="[301,399,1428,1452]" italics="true" pageId="1" pageNumber="581">Geocoris</emphasis>
)
<emphasis id="B935EA95FFDF270029D75785FDA8FA66" bold="true" box="[416,576,1428,1451]" italics="true" pageId="1" pageNumber="581">margaretarum</emphasis>
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,
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<emphasis id="B935EA95FFDF270029FC57A2FDEBFA06" bold="true" box="[395,515,1459,1483]" pageId="1" pageNumber="581">
(
<figureCitation id="137A2A02FFDF270029E557A2FE13FA06" box="[402,507,1459,1483]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="1.[835,870,826,848]" captionStartId-1="2.[113,148,1390,1412]" captionStartId-2="2.[813,850,1108,1130]" captionStartId-3="3.[136,171,1151,1173]" captionTargetBox-0="[835,1474,189,808]" captionTargetBox-1="[113,752,189,1374]" captionTargetBox-2="[813,1452,189,1093]" captionTargetBox-3="[136,1474,189,1135]" captionTargetId-0="figure-730@1.[835,1474,189,808]" captionTargetId-1="figure-298@2.[113,752,189,1375]" captionTargetId-2="figure-330@2.[813,1452,189,1093]" captionTargetId-3="figure-7@3.[136,1474,189,1135]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="3" captionText-0="Fig. 1. Habitus of Geocoris margaretarum new species (holotype, NHMW). A, dorsal view; B, lateral view." captionText-1="Fig. 2. Variability of head pattern in Geocoris margaretarum sp. n. A, holotype from Yunnan, China (NHMW); B, paratype from Arunachal province, India (MMB); C, paratype from Kambaiti Pass, Mianmar (NHRS)." captionText-2="Fig. 3. Genitalia of Geocoris margaretarum sp. n. A, male pygophore; B, male paramere from posterior view; C, male paramere from dorsal view; D, male aedeagus; E, female spemartheca." captionText-3="Fig. 4. Distribution of Geocoris margaretarum new species." figureDoi-0="http://doi.org/10.5281/zenodo.5360044" figureDoi-1="http://doi.org/10.5281/zenodo.5360048" figureDoi-2="http://doi.org/10.5281/zenodo.5360046" figureDoi-3="http://doi.org/10.5281/zenodo.5360050" httpUri-0="https://zenodo.org/record/5360044/files/figure.png" httpUri-1="https://zenodo.org/record/5360048/files/figure.png" httpUri-2="https://zenodo.org/record/5360046/files/figure.png" httpUri-3="https://zenodo.org/record/5360050/files/figure.png" pageId="1" pageNumber="581">Figs. 14</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
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<emphasis id="B935EA95FFDF270028FF57E2FE86F9C6" bold="true" box="[136,366,1523,1547]" pageId="1" pageNumber="581">Material examined.</emphasis>
<materialsCitation id="3B293CDAFFDF2700290F57E5FD3FF981" collectionCode="NHMW" collectorName="E. Jendek &amp; O. Sausa." country="China" location="Baoshan" pageId="1" pageNumber="581" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
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: male (
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),
<collectingCountry id="F3567617FFDF27002ADD57E5FCEAF9C1" box="[682,770,1524,1548]" name="China" pageId="1" pageNumber="581">CHINA</collectingCountry>
,
<collectingRegion id="4985F865FFDF270028FF5405FF0EF9E6" box="[136,230,1556,1579]" country="China" name="Yunnan" pageId="1" pageNumber="581">Yunnan</collectingRegion>
1993 /
<quantity id="4CB99B62FFDF270029235405FE5BF9E1" box="[340,435,1556,1580]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="581" unit="km" value="100.0">100 km</quantity>
W
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/ 14.-21. 6. / Gaoligongshan Nat. Res. / leg.
<collectorName id="26B45351FFDF270029A85425FDA1F981" box="[479,585,1588,1612]" pageId="1" pageNumber="581">E. Jendek</collectorName>
&amp;
<collectorName id="26B45351FFDF27002A185425FD3FF981" box="[623,727,1588,1612]" pageId="1" pageNumber="581">O. Sausa.</collectorName>
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</paragraph>
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<typeStatus id="54FA8825FFDF270028FF5465FEE8F941" box="[136,256,1652,1676]" pageId="1" pageNumber="581" type="paratype">Paratypes</typeStatus>
:
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,
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(
<collectionCode id="ED50AE42FFDF27002A7B5465FD8EF941" box="[524,614,1652,1676]" pageId="1" pageNumber="581">MZMB</collectionCode>
) NE
<collectingCountry id="F3567617FFDF27002ADA5465FCE8F941" box="[685,768,1652,1676]" name="India" pageId="1" pageNumber="581">INDIA</collectingCountry>
, ARUNACHAL PR. / ETALIN vicinity,
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</quantity>
/
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, /
<collectorName id="26B45351FFDF2700295454A5FE5AF901" box="[291,434,1716,1740]" pageId="1" pageNumber="581">L. Dembický</collectorName>
leg.
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<date id="FFFF1047FFDF2700299D54A5FD91F901" box="[490,633,1716,1740]" pageId="1" pageNumber="581" value="2012-05-12" valueMax="2012-05-25" valueMin="2012-05-12">12-25.v.2012</date>
/ AP 3/2012 MZM Expedition;
<materialsCitation id="3B293CDAFFDF2700292254C5FDFCF8E1" collectionCode="HNHM" collectorName="L. Dembicky" country="India" elevation="700" latitude="28.615557" location="INDIA" longLatPrecision="20" longitude="95.88917" pageId="1" pageNumber="581" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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(
<collectionCode id="ED50AE42FFDF27002A6A54C5FD9CF921" box="[541,628,1748,1772]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="581">HNHM</collectionCode>
), NE
<collectingCountry id="F3567617FFDF27002AC354C5FCEAF921" box="[692,770,1748,1772]" name="India" pageId="1" pageNumber="581">INDIA</collectingCountry>
, ARUNACHAL PR. / ETALIN vicinity,
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/
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, /
<collectorName id="26B45351FFDF2700294B5505FE33F8E1" box="[316,475,1812,1836]" pageId="1" pageNumber="581">L. Dembický</collectorName>
leg.
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<date id="FFFF1047FFDF27002A565505FD29F8E1" box="[545,705,1812,1836]" pageId="1" pageNumber="581" value="2012-05-12" valueMax="2012-05-25" valueMin="2012-05-12">12-25.v.2012</date>
/ AP 3/2012 MZM Expedition;
<materialsCitation id="3B293CDAFFDF270029C25525FD04F841" collectionCode="NHMW" collectorName="Gaoligongshan Nat. Res. &amp; E. Jendek &amp; O. Sausa" country="China" location="100 km W Baoshan" pageId="1" pageNumber="581" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
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,
<specimenCount id="9D47FD0EFFDF27002A695525FD63F881" box="[542,651,1844,1868]" pageId="1" pageNumber="581" type="female">4 females</specimenCount>
(
<collectionCode id="ED50AE42FFDF27002AEA5525FD13F881" box="[669,763,1844,1868]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="1" pageNumber="581" type="Museum">NHMW</collectionCode>
),
<collectingCountry id="F3567617FFDF270028FF5545FF08F8A1" box="[136,224,1876,1900]" name="China" pageId="1" pageNumber="581">CHINA</collectingCountry>
,
<collectingRegion id="4985F865FFDF270028995545FEABF8A6" box="[238,323,1876,1899]" country="China" name="Yunnan" pageId="1" pageNumber="581">Yunnan</collectingRegion>
1993 /
<location id="8E9E605CFFDF270029D75545FD97F8A1" LSID="urn:lsid:plazi:treatment:03E88791FFDF270228CD5785FD9BFA49:8E9E605CFFDF270029D75545FD97F8A1" box="[416,639,1876,1900]" country="China" name="100 km W Baoshan" pageId="1" pageNumber="581" stateProvince="Yunnan">
<quantity id="4CB99B62FFDF270029D75545FE1CF8A1" box="[416,500,1876,1900]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="581" unit="km" value="100.0">100 km</quantity>
W Baoshan
</location>
/ 14.-21. 6. /
<collectorName id="26B45351FFDF270028EE5565FE42F846" box="[153,426,1908,1932]" pageId="1" pageNumber="581">Gaoligongshan Nat. Res.</collectorName>
/ leg.
<collectorName id="26B45351FFDF270029835565FDB7F841" box="[500,607,1908,1932]" pageId="1" pageNumber="581">E. Jendek</collectorName>
&amp;
<collectorName id="26B45351FFDF27002AF15565FD04F841" box="[646,748,1908,1932]" pageId="1" pageNumber="581">O. Sausa</collectorName>
</materialsCitation>
;
<materialsCitation id="3B293CDAFFDF27002A8E5565FF5EF821" collectingDate="1936-11-10" collectionCode="BMNH" collectorName="Alt. &amp; M. Steele. &amp; Blackett MJ" country="China" county="Mishmi Hills" location="Taphlogam." municipality="Dalai Valley" pageId="1" pageNumber="581" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D47FD0EFFDF27002A8E5565FF54F861" pageId="1" pageNumber="581" type="male">1 male</specimenCount>
(
<collectionCode id="ED50AE42FFDF270028B85585FECDF861" box="[207,293,1940,1964]" pageId="1" pageNumber="581">BMNH</collectionCode>
), ASSAM:
<collectingCounty id="629F4E0BFFDF270029DD5585FDD7F861" box="[426,575,1940,1964]" pageId="1" pageNumber="581">Mishmi Hills</collectingCounty>
, /
<collectingMunicipality id="6B9AACFDFFDF27002A175585FD18F861" box="[608,752,1940,1964]" pageId="1" pageNumber="581">Dalai Valley</collectingMunicipality>
, /
<location id="8E9E605CFFDF270028FF55A5FEE1F801" LSID="urn:lsid:plazi:treatment:03E88791FFDF270228CD5785FD9BFA49:8E9E605CFFDF270028FF55A5FEE1F801" box="[136,265,1972,1996]" country="China" county="Mishmi Hills" municipality="Dalai Valley" name="Taphlogam." pageId="1" pageNumber="581" stateProvince="Yunnan">Taphlogam.</location>
/
<date id="FFFF1047FFDF2700296C55A5FE71F801" box="[283,409,1972,1996]" pageId="1" pageNumber="581" value="1936-11-10">
<collectingDate id="EFBBE9AFFFDF2700296C55A5FE71F801" box="[283,409,1972,1996]" pageId="1" pageNumber="581" value="1936-11-10">10. xi. 1936</collectingDate>
</date>
//
<collectorName id="26B45351FFDF270029C455A5FE35F801" box="[435,477,1972,1996]" pageId="1" pageNumber="581">Alt.</collectorName>
ft. /
<collectorName id="26B45351FFDF27002A7B55A5FD91F801" box="[524,633,1972,1996]" pageId="1" pageNumber="581">M. Steele.</collectorName>
/
<collectorName id="26B45351FFDF27002AFC55A5FD2AF806" box="[651,706,1972,1995]" name="Blackett MJ" pageId="1" pageNumber="581">B.M.</collectorName>
1937- 324
</materialsCitation>
.;
<materialsCitation id="3B293CDAFFDF270028BF55C5FCEBF7C1" collectingDate="1937-05-02" collectingDateMax="1937-05-03" collectingDateMin="1937-05-02" collectionCode="NHRS" country="Myanmar" elevation="2134" location="Kambaiti" pageId="1" pageNumber="581" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D47FD0EFFDF270028BF55C5FEF5F821" box="[200,285,2004,2028]" pageId="1" pageNumber="581" type="male">3 males</specimenCount>
(
<collectionCode id="ED50AE42FFDF2700295855C5FE93F821" box="[303,379,2004,2028]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="1" pageNumber="581">NHRS</collectionCode>
), N. E.
<collectingCountry id="F3567617FFDF270029A155C5FDD0F821" box="[470,568,2004,2028]" name="Myanmar" pageId="1" pageNumber="581">BURMA</collectingCountry>
/
<location id="8E9E605CFFDF27002A2455C5FD5EF821" LSID="urn:lsid:plazi:treatment:03E88791FFDF270228CD5785FD9BFA49:8E9E605CFFDF27002A2455C5FD5EF821" box="[595,694,2004,2028]" country="Myanmar" name="Kambaiti" pageId="1" pageNumber="581">Kambaiti</location>
<quantity id="4CB99B62FFDF27002AB755C5FCEEF821" box="[704,774,2004,2028]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="1" pageNumber="581" unit="ft" value="7000.0">
<elevation id="006CD1B4FFDF27002AB755C5FCEEF821" box="[704,774,2004,2028]" metricMagnitude="3" metricUnit="m" metricValue="2.1336" pageId="1" pageNumber="581" unit="ft" value="7000.0">7000ft</elevation>
</quantity>
/
<date id="FFFF1047FFDF270028EF55E5FEE6F7C1" box="[152,270,2036,2060]" pageId="1" pageNumber="581" value="1937-05-02" valueMax="1937-05-03" valueMin="1937-05-02">
<collectingDate id="EFBBE9AFFFDF270028EF55E5FEE6F7C1" box="[152,270,2036,2060]" pageId="1" pageNumber="581" value="1937-05-02" valueMax="1937-05-03" valueMin="1937-05-02">2-3/5 1937</collectingDate>
</date>
/ R. MALAISE // NHRS-GULI / 000006386
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE3687FFDF27002B3451B5FAFBFA51" blockId="1.[834,1476,932,1436]" pageId="1" pageNumber="581">
<emphasis id="B935EA95FFDF27002B3451B5FC3CFC71" bold="true" box="[835,980,932,956]" pageId="1" pageNumber="581">Colouration.</emphasis>
Head variably dark ochraceous or orange with extended, irregular black marking of various extent basally (
<figureCitation id="137A2A02FFDF27002B3A51F5FC7FFC31" box="[845,919,996,1020]" captionStart="Fig" captionStartId="2.[113,148,1390,1412]" captionTargetBox="[113,752,189,1374]" captionTargetId="figure-298@2.[113,752,189,1375]" captionTargetPageId="2" captionText="Fig. 2. Variability of head pattern in Geocoris margaretarum sp. n. A, holotype from Yunnan, China (NHMW); B, paratype from Arunachal province, India (MMB); C, paratype from Kambaiti Pass, Mianmar (NHRS)." figureDoi="http://doi.org/10.5281/zenodo.5360048" httpUri="https://zenodo.org/record/5360048/files/figure.png" pageId="1" pageNumber="581">Fig. 2</figureCitation>
). Clypeus sometimes with dark brown marking basally. Antennal segments I and IV ochraceous, segment I sometimes infused with fuscous ventrally; segments II and III dark brownish with apex ochraceous. Labial segments uniformly ochraceous. Thorax: Pronotum and scutellum dark brownish, pronotal callosities and humeral angles slightly paler in hue. Clavus and corium of hemelytron pale ochraceous with fuscous punctures. Membrane hyaline. Thoracic pleura and sterna dark brownish; prothorax with ochraceous collar ventrolaterally, irregular ochraceous spots above coxae (
<figureCitation id="137A2A02FFDF27002BAE5735FBC5FAF1" box="[985,1069,1316,1340]" captionStart="Fig" captionStartId="1.[835,870,826,848]" captionTargetBox="[835,1474,189,808]" captionTargetId="figure-730@1.[835,1474,189,808]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Geocoris margaretarum new species (holotype, NHMW). A, dorsal view; B, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5360044" httpUri="https://zenodo.org/record/5360044/files/figure.png" pageId="1" pageNumber="581">Fig. 1B</figureCitation>
). Orifice of metathoracic scent gland ochraceous. Legs ochraceous with an irregular fuscous band on apices of femora dorsally. Abdomen: Segments uniformly dark brownish with silvery setae ventrally.
</paragraph>
<paragraph id="8BFE3687FFDF27032B3457D5FEFDF804" blockId="1.[835,1476,1476,2108]" lastBlockId="2.[113,754,1553,1994]" lastPageId="2" lastPageNumber="582" pageId="1" pageNumber="581">
<emphasis id="B935EA95FFDF27002B3457D5FC50FA11" bold="true" box="[835,952,1476,1500]" pageId="1" pageNumber="581">Structure.</emphasis>
Head surface impunctate, shiny. Ocular sulcus reduced, only visible near ocelli. Ocelli situated almost equidistantly from the compound eyes and each other; ratio of ocellar distance to eye-ocellus distance: 1: 1.22. Clypeus rounded, exceeding mandibular plates. Antennal segment I shortest, II and III subequal, segment IV longest; ratio of antennal segments: 1: 1.86: 1.71: 2.14 (1: 1.83: 1.8: 2.331: 1.86: 1.71: 2.14). Labial segment I not reaching prothorax; segment II shorter than III, segments III longest IV subequal to segment II; ratio of labial segments: 1: 0.69: 1.07: 0.77 (1: 0.72: 1.09: 0.811: 0.69: 1.07: 0.77). Thorax: Pronotum widened, slightly trapezoidal; surface shiny with dense, strong punctuation. Pronotum ratio: 1:1.75. Pronotal calli, humeral angles and posterior margin impunctate; calli slightly elevated. Lateral margin of pronotum with a blunt but well-defined carina, disc with slight impression medially above carina. Scutellum nearly equilateral with sharp apex; ratio of length and basal width: 1: 1.07; surface with dense, strong punctuation except slightly elevated triradiate carina. Clavus of hemelytron narrowing gradually towards apex of scutellum, claval commissure absent; clavus with one complete row of punctures at corial margin. R-M region with sparse, deep, irregular punctuation. Apical margin of corium S-shape with deep concavity. Membrane without visible veins. Pronotal pleuron with dense, deep but fine punctation except supracoxal lobes and orifice of metathoracic scent gland. Opening of metathoracic scent gland situated ventrolaterally close to anterior margin of metathorax; peritreme ear-shaped, somewhat protruding, evaporative area small. Abdomen: Abdominal dorsum with fine, dense punctuation and pubescence medially. Genitalia: Female spermatheca as in
<figureCitation id="137A2A02FFDC2703294C5560FE66F847" box="[315,398,1905,1930]" captionStart="Fig" captionStartId="2.[813,850,1108,1130]" captionTargetBox="[813,1452,189,1093]" captionTargetId="figure-330@2.[813,1452,189,1093]" captionTargetPageId="2" captionText="Fig. 3. Genitalia of Geocoris margaretarum sp. n. A, male pygophore; B, male paramere from posterior view; C, male paramere from dorsal view; D, male aedeagus; E, female spemartheca." figureDoi="http://doi.org/10.5281/zenodo.5360046" httpUri="https://zenodo.org/record/5360046/files/figure.png" pageId="2" pageNumber="582">Fig. 3E</figureCitation>
. Male pygophore as in
<figureCitation id="137A2A02FFDC27032AF85560FD0FF847" box="[655,743,1905,1930]" captionStart="Fig" captionStartId="2.[813,850,1108,1130]" captionTargetBox="[813,1452,189,1093]" captionTargetId="figure-330@2.[813,1452,189,1093]" captionTargetPageId="2" captionText="Fig. 3. Genitalia of Geocoris margaretarum sp. n. A, male pygophore; B, male paramere from posterior view; C, male paramere from dorsal view; D, male aedeagus; E, female spemartheca." figureDoi="http://doi.org/10.5281/zenodo.5360046" httpUri="https://zenodo.org/record/5360046/files/figure.png" pageId="2" pageNumber="582">Fig. 3A</figureCitation>
.; paramere as in
<figureCitation id="137A2A02FFDC2703296A5580FE7BF867" box="[285,403,1937,1962]" captionStart="Fig" captionStartId="2.[813,850,1108,1130]" captionTargetBox="[813,1452,189,1093]" captionTargetId="figure-330@2.[813,1452,189,1093]" captionTargetPageId="2" captionText="Fig. 3. Genitalia of Geocoris margaretarum sp. n. A, male pygophore; B, male paramere from posterior view; C, male paramere from dorsal view; D, male aedeagus; E, female spemartheca." figureDoi="http://doi.org/10.5281/zenodo.5360046" httpUri="https://zenodo.org/record/5360046/files/figure.png" pageId="2" pageNumber="582">Fig. 3B, C</figureCitation>
; gonoporal process of aedeagus with ~17 coils.
</paragraph>
</subSubSection>
<caption id="DF3E660FFFDC27032B5A5645FA6AFB69" ID-DOI="http://doi.org/10.5281/zenodo.5360046" ID-Zenodo-Dep="5360046" httpUri="https://zenodo.org/record/5360046/files/figure.png" pageId="2" pageNumber="582" startId="2.[813,850,1108,1130]" targetBox="[813,1452,189,1093]" targetPageId="2">
<paragraph id="8BFE3687FFDC27032B5A5645FA6AFB69" blockId="2.[811,1451,1108,1188]" pageId="2" pageNumber="582">
Fig. 3. Genitalia of
<taxonomicName id="4C414D04FFDC27032C7F5645FAEFFBA7" authority="Kóbor, 2018" authorityName="Kóbor" authorityYear="2018" box="[1032,1287,1108,1130]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="margaretarum" status="sp. nov.">
<emphasis id="B935EA95FFDC27032C7F5645FAEFFBA7" box="[1032,1287,1108,1130]" italics="true" pageId="2" pageNumber="582">Geocoris margaretarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657EEFFDC27032D665645FAA5FBA4" box="[1297,1357,1108,1129]" pageId="2" pageNumber="582" rank="species">sp. n.</taxonomicNameLabel>
A, male pygophore; B, male paramere from posterior view; C, male paramere from dorsal view; D, male aedeagus; E, female spemartheca.
</paragraph>
</caption>
<caption id="DF3E660FFFDC27032806577FFE88FA17" ID-DOI="http://doi.org/10.5281/zenodo.5360048" ID-Zenodo-Dep="5360048" httpUri="https://zenodo.org/record/5360048/files/figure.png" pageId="2" pageNumber="582" startId="2.[113,148,1390,1412]" targetBox="[113,752,189,1374]" targetPageId="2">
<paragraph id="8BFE3687FFDC27032806577FFE88FA17" blockId="2.[113,752,1390,1498]" pageId="2" pageNumber="582">
Fig. 2. Variability of head pattern in
<taxonomicName id="4C414D04FFDC27032997577FFD24FA49" authority="Kóbor, 2018" authorityName="Kóbor" authorityYear="2018" box="[480,716,1390,1412]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="margaretarum" status="sp. nov.">
<emphasis id="B935EA95FFDC27032997577FFD24FA49" box="[480,716,1390,1412]" italics="true" pageId="2" pageNumber="582">Geocoris margaretarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657EEFFDC27032AA3577FFF6BFA6D" pageId="2" pageNumber="582" rank="species">sp. n.</taxonomicNameLabel>
A, holotype from Yunnan, China (NHMW); B, paratype from Arunachal province, India (MMB); C, paratype from Kambaiti Pass, Mianmar (NHRS).
</paragraph>
</caption>
<subSubSection id="C35B650CFFDC2703280655E3FC4BFA7E" pageId="2" pageNumber="582" type="description">
<paragraph id="8BFE3687FFDC2703280655E3FD18F7E4" blockId="2.[113,752,2033,2090]" pageId="2" pageNumber="582">
<emphasis id="B935EA95FFDC2703280655E3FECDF7C7" bold="true" box="[113,293,2034,2058]" pageId="2" pageNumber="582">Measurements.</emphasis>
(
<typeStatus id="54FA8825FFDC2703294F55E0FE76F7C4" box="[312,414,2033,2057]" pageId="2" pageNumber="582" type="holotype">Holotype</typeStatus>
[minimummaximum]): total body length: 4.98 (4.934.98); head length: 0.79 (0.650.79);
</paragraph>
<paragraph id="8BFE3687FFDC27032B5C56CAFC4BFA7E" blockId="2.[811,1451,1243,1459]" pageId="2" pageNumber="582">head width: 2.23 (2.142.23); antennal segments (I-II-III- IV): 0.33-0.60-0.56-0.70 (0.28-0.51-0.51-0.65 0.33-0.60- 0.56-0.70); labial segments (I-II-III-IV): 0.60-0.42-0.65-0.47 (0.51-0.37-0.56-0.42 0.60-0.42-0.65-0.47); pronotum length: 1.30 (1.161.30); pronotum width: 2.28 (2.092.28); scutellum length: 1.21 (1.071.21); scutellum width: 1.3 (1.211.3).</paragraph>
</subSubSection>
<subSubSection id="C35B650CFFDC27032B5A57CBFA7AF93E" pageId="2" pageNumber="582" type="diagnosis">
<paragraph id="8BFE3687FFDC27032B5A57CBFA7AF93E" blockId="2.[813,1453,1498,1779]" pageId="2" pageNumber="582">
<emphasis id="B935EA95FFDC27032B5A57CBFC49FA3F" bold="true" box="[813,929,1498,1522]" pageId="2" pageNumber="582">Diagnosis.</emphasis>
This new species differs in colouration patterns and punctation from other closely related representatives in the region: Head with variably extended blackish markings always exceeding the level of ocelli. Femora with fuscous annulation apically. Punctuation finer and sparser than
<taxonomicName id="4C414D04FFDC27032B5A546AFC5DF95E" authorityName="Jakovlev" authorityYear="1904" box="[813,949,1659,1683]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B935EA95FFDC27032B5A546AFC5DF95E" box="[813,949,1659,1683]" italics="true" pageId="2" pageNumber="582">G. chinensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C414D04FFDC27032BB1546AFB84F95E" baseAuthorityName="Fieber" baseAuthorityYear="1844" box="[966,1132,1659,1683]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="ochropterus">
<emphasis id="B935EA95FFDC27032BB1546AFB84F95E" box="[966,1132,1659,1683]" italics="true" pageId="2" pageNumber="582">G. ochropterus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C414D04FFDC27032CD9546AFAFDF95F" baseAuthorityName="Uhler" baseAuthorityYear="1860" box="[1198,1301,1659,1682]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="582" phylum="Arthropoda" rank="species" species="varius">
<emphasis id="B935EA95FFDC27032CD9546AFAFDF95F" box="[1198,1301,1659,1682]" italics="true" pageId="2" pageNumber="582">G. varius</emphasis>
</taxonomicName>
contrastingly punctation of hemelytra deeper, Cu-M region with sparse, irregular punctures. Apical margin of corium S-shaped, unlikely to relatives where it is simply concave (
<figureCitation id="137A2A02FFDC27032D3554CAFA6BF93E" box="[1346,1411,1755,1779]" captionStart="Fig" captionStartId="1.[835,870,826,848]" captionTargetBox="[835,1474,189,808]" captionTargetId="figure-730@1.[835,1474,189,808]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Geocoris margaretarum new species (holotype, NHMW). A, dorsal view; B, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5360044" httpUri="https://zenodo.org/record/5360044/files/figure.png" pageId="2" pageNumber="582">Fig. 1</figureCitation>
.)
</paragraph>
</subSubSection>
<subSubSection id="C35B650CFFDC27032B5A550AFC7FF83E" pageId="2" pageNumber="582" type="etymology">
<paragraph id="8BFE3687FFDC27032B5A550AFC7FF83E" blockId="2.[812,1452,1819,2035]" pageId="2" pageNumber="582">
<emphasis id="B935EA95FFDC27032B5A550AFC5FF8FE" bold="true" box="[813,951,1819,1843]" pageId="2" pageNumber="582">Etymology.</emphasis>
The name of the species comes from the feminine first name Margaret (Margit in Hungarian) as a tribute to the birthday of Margit Kiss (mother of the author) and also commemorates the life of Margit Kiss (19322017, grandmother of the author) expressing the gratefulness towards them for the love and support received throughout the years.
</paragraph>
</subSubSection>
<caption id="DF3E660FFFDD270228FF566EFD30FB58" ID-DOI="http://doi.org/10.5281/zenodo.5360050" ID-Zenodo-Dep="5360050" box="[136,728,1151,1173]" httpUri="https://zenodo.org/record/5360050/files/figure.png" pageId="3" pageNumber="583" startId="3.[136,171,1151,1173]" targetBox="[136,1474,189,1135]" targetPageId="3">
<paragraph id="8BFE3687FFDD270228FF566EFD30FB58" blockId="3.[136,728,1151,1173]" box="[136,728,1151,1173]" pageId="3" pageNumber="583">
Fig. 4. Distribution of
<taxonomicName id="4C414D04FFDD2702291E566EFDBCFB59" authority="Kóbor, 2018" authorityName="Kóbor" authorityYear="2018" box="[361,596,1151,1172]" class="Insecta" family="Lygaeidae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="583" phylum="Arthropoda" rank="species" species="margaretarum" status="sp. nov.">
<emphasis id="B935EA95FFDD2702291E566EFDBCFB59" box="[361,596,1151,1172]" italics="true" pageId="3" pageNumber="583">Geocoris margaretarum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657EEFFDD27022A2B566EFD3CFB58" box="[604,724,1151,1173]" pageId="3" pageNumber="583" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<subSubSection id="C35B650CFFDD270228FF56DAFD9BFA49" pageId="3" pageNumber="583" type="distribution">
<paragraph id="8BFE3687FFDD270228FF56DAFD9BFA49" blockId="3.[134,776,1227,1412]" pageId="3" pageNumber="583">
<emphasis id="B935EA95FFDD270228FF56DAFEFEFB2E" bold="true" box="[136,278,1227,1251]" pageId="3" pageNumber="583">Distribution.</emphasis>
According to the available data this new species is distributed in North-East
<collectingCountry id="F3567617FFDD270229CD56FDFE19FAC9" box="[442,497,1260,1284]" name="India" pageId="3" pageNumber="583">India</collectingCountry>
(
<collectingRegion id="4985F865FFDD27022A7656FDFD60FACE" box="[513,648,1260,1284]" country="India" name="Assam" pageId="3" pageNumber="583">Assam state</collectingRegion>
,
<collectingRegion id="4985F865FFDD27022AE156FDFF09FAE9" country="India" name="Arunachal Pradesh" pageId="3" pageNumber="583">Arunachal Pradesh</collectingRegion>
) and South-East
<collectingCountry id="F3567617FFDD270229DD571DFE03FAE9" box="[426,491,1292,1316]" name="China" pageId="3" pageNumber="583">China</collectingCountry>
(
<collectingRegion id="4985F865FFDD2702298A571DFD2FFAE9" box="[509,711,1292,1316]" country="China" name="Yunnan" pageId="3" pageNumber="583">Yunnan province</collectingRegion>
). The distributional area of the species is in the
<collectingCountry id="F3567617FFDD27022A23573DFD75FA8E" box="[596,669,1324,1347]" name="Myanmar" pageId="3" pageNumber="583">Burma</collectingCountry>
monsoon forest (Indomalayan realm) biogeographical province according to
<bibRefCitation id="EFD04B76FFDD27022960577DFE30FA49" author="Udvardy MDF" box="[279,472,1388,1412]" pageId="3" pageNumber="583" refId="ref4869" refString="Udvardy MDF (1975) A Classification of the Biogeographical Provinces of World. IUCN Occasional Paper, 18, 49 pp." type="book" year="1975">Udvardys (1975)</bibRefCitation>
classification.
</paragraph>
</subSubSection>
</treatment>
</document>