282 lines
34 KiB
XML
282 lines
34 KiB
XML
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<mods:title id="C1986C18BB8B8EF1C78745FD3ED76CCC">Review of the Asian Thaumastodinae (Coleoptera, Byrrhoidea, Limnichidae), with a phylogeny of the genera</mods:title>
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<mods:namePart id="27CEDFD437600F674124DA5FC93482C6">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:title id="F0FB871E21DD161D5C88A030BB2B3F59">European Journal of Taxonomy</mods:title>
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<mods:date id="241AEDD13082F56AAA7E0C0D359742EB">2019</mods:date>
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<treatment id="03F9EF199F44FFD1FD89FA9F590B2B09" ID-DOI="http://doi.org/10.5281/zenodo.5386731" ID-GBIF-Taxon="160630992" ID-Zenodo-Dep="5386731" LSID="urn:lsid:plazi:treatment:03F9EF199F44FFD1FD89FA9F590B2B09" httpUri="http://treatment.plazi.org/id/03F9EF199F44FFD1FD89FA9F590B2B09" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
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<subSubSection id="C34A0D849F44FFD0FD89FA9F58F52BEA" box="[619,969,1407,1433]" pageId="18" pageNumber="19" type="nomenclature">
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<paragraph id="8BEF5E0F9F44FFD0FD89FA9F58F52BEA" blockId="18.[619,969,1407,1433]" box="[619,969,1407,1433]" pageId="18" pageNumber="19">
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<heading id="D0A7E9639F44FFD0FD89FA9F58F52BEA" box="[619,969,1407,1433]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
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Genus
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<taxonomicName id="4C50258C9F44FFD0FD5FFA9F58F52BEA" ID-CoL="5R5P" authority="Spilman, 1972" authorityName="Spilman" authorityYear="1972" box="[701,969,1407,1433]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B924821D9F44FFD0FD5FFA9F582B2BEA" bold="true" box="[701,791,1407,1433]" italics="true" pageId="18" pageNumber="19">Mexico</emphasis>
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<bibRefCitation id="EFC123FE9F44FFD0FCFCFA9F58F52BEA" author="Spilman T. J." box="[798,969,1407,1433]" pageId="18" pageNumber="19" pagination="108 - 115" refId="ref15040" refString="Spilman T. J. 1972. A new genus and species of jumping shore beetle from Mexico. Pan-Pacific Entomologist 48: 108 - 115." type="journal article" year="1972">Spilman, 1972</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A0D849F44FFD0FF5FFA2358F428E2" pageId="18" pageNumber="19" type="reference_group">
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<paragraph id="8BEF5E0F9F44FFD0FF5FFA235AC7285B" blockId="18.[189,1398,1474,1576]" pageId="18" pageNumber="19">
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<treatmentCitationGroup id="AB4079219F44FFD0FF5FFA235E4A2871" pageId="18" pageNumber="19">
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<taxonomicName id="4C50258C9F44FFD0FF5FFA23593C2BAE" ID-CoL="5R5P" authority="Spilman, 1972: 113" authorityName="Spilman" authorityPageNumber="113" authorityYear="1972" box="[189,512,1474,1501]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B924821D9F44FFD0FF5FFA235A2F2BAF" box="[189,275,1475,1500]" italics="true" pageId="18" pageNumber="19">Mexico</emphasis>
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<treatmentCitation id="0AF1781E9F44FFD0FEF9FA22593C2BAE" author="Spilman T. J." box="[283,512,1474,1501]" page="113" pageId="18" pageNumber="19" year="1972">
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<bibRefCitation id="EFC123FE9F44FFD0FEF9FA22593C2BAE" author="Spilman T. J." box="[283,512,1474,1501]" pageId="18" pageNumber="19" pagination="108 - 115" refId="ref15040" refString="Spilman T. J. 1972. A new genus and species of jumping shore beetle from Mexico. Pan-Pacific Entomologist 48: 108 - 115." type="journal article" year="1972">Spilman, 1972: 113</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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||
(
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<typeStatus id="54EBE0AD9F44FFD0FDF2FA2359782BAE" box="[528,580,1475,1501]" pageId="18" pageNumber="19">type</typeStatus>
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species:
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<taxonomicName id="4C50258C9F44FFD0FD4DFA235F232BAE" ID-CoL="42KRW" authority="Spilman, 1972" authorityName="Spilman" authorityYear="1972" box="[687,1055,1474,1501]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="litoralis">
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<emphasis id="B924821D9F44FFD0FD4DFA2358562BAF" box="[687,874,1474,1500]" italics="true" pageId="18" pageNumber="19">Mexico litoralis</emphasis>
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<bibRefCitation id="EFC123FE9F44FFD0FC93FA225F232BAE" author="Spilman T. J." box="[881,1055,1474,1501]" pageId="18" pageNumber="19" pagination="108 - 115" refId="ref15040" refString="Spilman T. J. 1972. A new genus and species of jumping shore beetle from Mexico. Pan-Pacific Entomologist 48: 108 - 115." type="journal article" year="1972">Spilman, 1972</bibRefCitation>
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</taxonomicName>
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, by original designation).
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<taxonomicName id="4C50258C9F44FFD0FF5FFA0859042871" ID-CoL="62FGD" authority="Sato, 1994: 174" authorityName="Sato" authorityPageNumber="174" authorityYear="1994" box="[189,568,1512,1538]" class="Insecta" family="Limnichidae" genus="Babalimnichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B924821D9F44FFD0FF5FFA085A512871" box="[189,365,1512,1538]" italics="true" pageId="18" pageNumber="19">Babalimnichus</emphasis>
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<treatmentCitation id="0AF1781E9F44FFD0FE9AFA0859042871" author="Sato M." box="[376,568,1512,1538]" page="174" pageId="18" pageNumber="19" year="1994">
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<bibRefCitation id="EFC123FE9F44FFD0FE9AFA0859042871" author="Sato M." box="[376,568,1512,1538]" pageId="18" pageNumber="19" pagination="173 - 177" refId="ref14802" refString="Sato M. 1994. Note on the genus Pseudeucinetus Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa Entomological Society 2: 173 - 177." type="journal article" year="1994">Satô, 1994: 174</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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||
(
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<typeStatus id="54EBE0AD9F44FFD0FDA9FA0859432871" box="[587,639,1512,1538]" pageId="18" pageNumber="19">type</typeStatus>
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species:
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<taxonomicName id="4C50258C9F44FFD0FD13FA085FD82871" ID-CoL="K68W" authority="Sato, 1994: 174" authorityName="Sato" authorityPageNumber="174" authorityYear="1994" box="[753,1252,1512,1538]" class="Insecta" family="Limnichidae" genus="Babalimnichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="taiwanus">
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<emphasis id="B924821D9F44FFD0FD13FA085F2A2871" box="[753,1046,1512,1538]" italics="true" pageId="18" pageNumber="19">Babalimnichus taiwanus</emphasis>
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<treatmentCitation id="0AF1781E9F44FFD0FBC3FA085FD82871" author="Sato M." box="[1057,1252,1512,1538]" page="174" pageId="18" pageNumber="19" year="1994">
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<bibRefCitation id="EFC123FE9F44FFD0FBC3FA085FD82871" author="Sato M." box="[1057,1252,1512,1538]" pageId="18" pageNumber="19" pagination="173 - 177" refId="ref14802" refString="Sato M. 1994. Note on the genus Pseudeucinetus Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa Entomological Society 2: 173 - 177." type="journal article" year="1994">Satô, 1994: 174</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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, by original
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</treatmentCitationGroup>
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designation).
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<emphasis id="B924821D9F44FFD0FE68F9ED5AC7285B" bold="true" box="[394,507,1549,1576]" pageId="18" pageNumber="19">Syn. nov.</emphasis>
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</paragraph>
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<paragraph id="8BEF5E0F9F44FFD0FF5FF9B259312818" blockId="18.[189,968,1617,1681]" box="[189,525,1617,1644]" pageId="18" pageNumber="19">
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||
<treatmentCitationGroup id="AB4079219F44FFD0FF5FF9B259312818" box="[189,525,1617,1644]" pageId="18" pageNumber="19">
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||
<taxonomicName id="4C50258C9F44FFD0FF5FF9B259342818" ID-CoL="5R5P" authority="- Skelley 2005: 119" authorityName="- Skelley" authorityPageNumber="119" authorityYear="2005" box="[189,520,1617,1644]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B924821D9F44FFD0FF5FF9B25A2F2818" box="[189,275,1618,1643]" italics="true" pageId="18" pageNumber="19">Mexico</emphasis>
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||
–
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<treatmentCitation id="0AF1781E9F44FFD0FED3F9B159342818" author="Skelley P. E." box="[305,520,1617,1644]" page="119" pageId="18" pageNumber="19" year="2005">
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||
<bibRefCitation id="EFC123FE9F44FFD0FED3F9B159342818" author="Skelley P. E." box="[305,520,1617,1644]" pageId="18" pageNumber="19" pagination="119 - 122" refId="ref14836" refString="Skelley P. E. 2005. A new species of ' jumping shore beetle' in the genus Mexico Spilman from the Bahamas (Coleoptera: Limnichidae: Thaumastodinae). Insecta Mundi 19 (1 - 2): 119 - 122." type="journal article" year="2005">Skelley 2005: 119</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BEF5E0F9F44FFD0FF5FF99758F428E2" blockId="18.[189,968,1617,1681]" box="[189,968,1654,1681]" pageId="18" pageNumber="19">
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||
<treatmentCitationGroup id="AB4079219F44FFD0FF5FF99758F428E2" box="[189,968,1654,1681]" pageId="18" pageNumber="19">
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||
<taxonomicName id="4C50258C9F44FFD0FF5FF99759E528E2" ID-CoL="62FGD" authority="- Yoshitomi & Sato 2001: 472" authorityName="Yoshitomi & Sato" authorityPageNumber="472" authorityYear="2001" box="[189,729,1655,1681]" class="Insecta" family="Limnichidae" genus="Babalimnichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis id="B924821D9F44FFD0FF5FF9975A5128E2" box="[189,365,1655,1681]" italics="true" pageId="18" pageNumber="19">Babalimnichus</emphasis>
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–
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<treatmentCitation id="0AF1781E9F44FFD0FE6BF99759E528E2" author="Yoshitomi H. & Sato M." box="[393,729,1655,1681]" page="472" pageId="18" pageNumber="19" year="2001">
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||
<bibRefCitation id="EFC123FE9F44FFD0FE6BF99759E528E2" author="Yoshitomi H. & Sato M." box="[393,729,1655,1681]" pageId="18" pageNumber="19" pagination="471 - 474" refId="ref15361" refString="Yoshitomi H. & Sato M. 2001. Description of the larva of Babalimnichus masamii M. Sato (Coleoptera: Limnichidae, Thaumastodinae). Coleopterist's Bulletin 55: 471 - 474." type="journal article" year="2001">Yoshitomi & Satô 2001: 472</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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[larval description].
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||
</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C34A0D849F44FFD0FF5FF95A5A542922" pageId="18" pageNumber="19" type="diagnosis">
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||
<paragraph id="8BEF5E0F9F44FFD0FF5FF95A5A0A28A7" blockId="18.[189,310,1722,1748]" box="[189,310,1722,1748]" pageId="18" pageNumber="19">
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||
<heading id="D0A7E9639F44FFD0FF5FF95A5A0A28A7" bold="true" box="[189,310,1722,1748]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
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||
<emphasis id="B924821D9F44FFD0FF5FF95A5A0A28A7" bold="true" box="[189,310,1722,1748]" pageId="18" pageNumber="19">Diagnosis</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BEF5E0F9F44FFD0FF5FF90B5A542922" blockId="18.[189,1399,1771,1873]" pageId="18" pageNumber="19">Body oblong. Gena relatively narrow; shallow antennal groove present. Antennae stout and short, 11-segmented; antennal insertions situated near mandibles. Legs relatively short and stout; tarsal formula 4-4-4.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34A0D849F44FFD1FF5FF89A582C2C41" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="8BEF5E0F9F44FFD0FF5FF89A5A1329E7" blockId="18.[189,303,1914,1940]" box="[189,303,1914,1940]" pageId="18" pageNumber="19">
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||
<heading id="D0A7E9639F44FFD0FF5FF89A5A1329E7" bold="true" box="[189,303,1914,1940]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
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||
<emphasis id="B924821D9F44FFD0FF5FF89A5A1329E7" bold="true" box="[189,303,1914,1940]" pageId="18" pageNumber="19">Remarks</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BEF5E0F9F44FFD0FF5FF84B58642998" blockId="18.[189,1398,1963,2027]" pageId="18" pageNumber="19">
|
||
As already mentioned by
|
||
<bibRefCitation id="EFC123FE9F44FFD0FE0FF84B59A229B5" author="Skelley P. E." box="[493,670,1963,1990]" pageId="18" pageNumber="19" pagination="119 - 122" refId="ref14836" refString="Skelley P. E. 2005. A new species of ' jumping shore beetle' in the genus Mexico Spilman from the Bahamas (Coleoptera: Limnichidae: Thaumastodinae). Insecta Mundi 19 (1 - 2): 119 - 122." type="journal article" year="2005">Skelley (2005)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C50258C9F44FFD0FD48F84B58DC29B6" authority="Sato, 1994" authorityName="Sato" authorityYear="1994" box="[682,992,1963,1989]" class="Insecta" family="Limnichidae" genus="Babalimnichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B924821D9F44FFD0FD48F84B586629B6" box="[682,858,1963,1989]" italics="true" pageId="18" pageNumber="19">Babalimnichus</emphasis>
|
||
<bibRefCitation id="EFC123FE9F44FFD0FC80F84B58DC29B6" author="Sato M." box="[866,992,1963,1989]" pageId="18" pageNumber="19" pagination="173 - 177" refId="ref14802" refString="Sato M. 1994. Note on the genus Pseudeucinetus Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa Entomological Society 2: 173 - 177." type="journal article" year="1994">Satô, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
cannot be separated from
|
||
<taxonomicName id="4C50258C9F44FFD0FAFBF84C5E5329B6" authorityName="- Skelley" authorityYear="2005" box="[1305,1391,1964,1989]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924821D9F44FFD0FAFBF84C5E5329B6" box="[1305,1391,1964,1989]" italics="true" pageId="18" pageNumber="19">Mexico</emphasis>
|
||
</taxonomicName>
|
||
, and it should be treated as a junior synonym of the latter.
|
||
</paragraph>
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||
<paragraph id="8BEF5E0F9F45FFD1FF5FFEEB58642F3A" blockId="19.[189,1398,267,329]" pageId="19" pageNumber="20">
|
||
The male genital structures are very peculiar, i.e., with very short lateral lobes with apical setae, longer median lobe and the presence of ventral plates (
|
||
<figureCitation id="136B428A9F45FFD1FD0EFECE58752F3A" box="[748,841,302,329]" captionStart="Fig" captionStartId="2.[189,232,1894,1920]" captionTargetBox="[189,1398,774,1835]" captionTargetId="figure@2.[189,1398,756,1850]" captionTargetPageId="2" captionText="Fig. 1. Aedeagus, showing the terminology and the method of measurements. A. Mexico Spilman, 1972, left lateral view. B. Acontosceles Champion, 1924, left lateral view. C. Pseudeucinetus Heller, 1921, ventral view. Abbreviations: bap = basal piece; lal = lateral lobe; mel = median lobe; vep = ventral plate; BL = length of basal piece; LL = length of lateral lobe; ML = length of median lobe." figureDoi="http://doi.org/10.5281/zenodo.3564534" httpUri="https://zenodo.org/record/3564534/files/figure.png" pageId="19" pageNumber="20">Fig. 1A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFE905ABD2FF9" blockId="19.[189,385,368,394]" box="[189,385,368,394]" pageId="19" pageNumber="20">
|
||
<heading id="D0A7E9639F45FFD1FF5FFE905ABD2FF9" bold="true" box="[189,385,368,394]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
||
<emphasis id="B924821D9F45FFD1FF5FFE905ABD2FF9" bold="true" box="[189,385,368,394]" pageId="19" pageNumber="20">Biological notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFE7E5A2F2F8C" blockId="19.[189,1398,414,511]" pageId="19" pageNumber="20">
|
||
The habitat of this genus (
|
||
<figureCitation id="136B428A9F45FFD1FE14FE7E59022FCA" box="[502,574,414,441]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[189,1399,873,1724]" captionTargetId="figure@6.[189,1399,873,1724]" captionTargetPageId="6" captionText="Fig. 4. Habitats (A–C) and living adults (D–F) of Asian Thaumastodinae Champion, 1924. A. Dondong River, Pengkol, Yogyakarta special Region, Java, Indonesia (type locality of Acontosceles javanicus Yoshitomi & Putra, 2011 (F) and Pseudeucinetus javanicus Yoshitomi & Putra, 2010 (D)). B. Nusa Dua, Bali, Indonesia (type locality of Mexico baliensis sp. nov.). C. Hirashima, Ogasawara, Japan (type locality of Mexico ogasawaraensis sp. nov.). D. Type locality of Pseudeucinetus javanicus Yoshitomi & Putra, 2010. E. Mexico masamii (Satô, 1994) in Shikinejima, Izu Islands. F. Type locality of Acontosceles javanicus Yoshitomi & Putra, 2011." figureDoi="http://doi.org/10.5281/zenodo.3564540" httpUri="https://zenodo.org/record/3564540/files/figure.png" pageId="19" pageNumber="20">Fig. 4</figureCitation>
|
||
B–C) is rocky seashores. The adults (
|
||
<figureCitation id="136B428A9F45FFD1FC1EFE7E5F6A2FCA" box="[1020,1110,414,441]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[189,1399,873,1724]" captionTargetId="figure@6.[189,1399,873,1724]" captionTargetPageId="6" captionText="Fig. 4. Habitats (A–C) and living adults (D–F) of Asian Thaumastodinae Champion, 1924. A. Dondong River, Pengkol, Yogyakarta special Region, Java, Indonesia (type locality of Acontosceles javanicus Yoshitomi & Putra, 2011 (F) and Pseudeucinetus javanicus Yoshitomi & Putra, 2010 (D)). B. Nusa Dua, Bali, Indonesia (type locality of Mexico baliensis sp. nov.). C. Hirashima, Ogasawara, Japan (type locality of Mexico ogasawaraensis sp. nov.). D. Type locality of Pseudeucinetus javanicus Yoshitomi & Putra, 2010. E. Mexico masamii (Satô, 1994) in Shikinejima, Izu Islands. F. Type locality of Acontosceles javanicus Yoshitomi & Putra, 2011." figureDoi="http://doi.org/10.5281/zenodo.3564540" httpUri="https://zenodo.org/record/3564540/files/figure.png" pageId="19" pageNumber="20">Fig. 4E</figureCitation>
|
||
) are found in the splash zone of wet rocks, actively flying when disturbed. The larval stage was described by
|
||
<bibRefCitation id="EFC123FE9F45FFD1FB41FE215A332F8C" author="Yoshitomi H. & Sato M." pageId="19" pageNumber="20" pagination="471 - 474" refId="ref15361" refString="Yoshitomi H. & Sato M. 2001. Description of the larva of Babalimnichus masamii M. Sato (Coleoptera: Limnichidae, Thaumastodinae). Coleopterist's Bulletin 55: 471 - 474." type="journal article" year="2001">Yoshitomi & Satô (2001)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFDC6582C2C41" blockId="19.[189,784,548,576]" box="[189,784,548,576]" pageId="19" pageNumber="20">
|
||
<emphasis id="B924821D9F45FFD1FF5FFDC6582C2C41" bold="true" box="[189,784,548,576]" pageId="19" pageNumber="20">
|
||
Key to the males of the species of Asian
|
||
<taxonomicName id="4C50258C9F45FFD1FD4FFDC6583B2C33" authorityName="- Skelley" authorityYear="2005" box="[685,775,550,576]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B924821D9F45FFD1FD4FFDC6583B2C33" bold="true" box="[685,775,550,576]" italics="true" pageId="19" pageNumber="20">Mexico</emphasis>
|
||
</taxonomicName>
|
||
<superScript id="7C25F3479F45FFD1FCE5FDC4582C2C41" attach="left" box="[775,784,548,562]" fontSize="6" pageId="19" pageNumber="20">*</superScript>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34A0D849F45FFD1FF5FFDB55E4B2B65" pageId="19" pageNumber="20" type="key">
|
||
<key id="FFC1093E9F45FFD1FF5FFDB55E4B2B65" pageId="19" pageNumber="20">
|
||
<keyStep id="30A446AA9F45FFD1FF5FFDB55E4B2CC6" pageId="19" pageNumber="20">
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFDB55E4B2CE1" blockId="19.[189,1399,596,693]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFDB55E4B2CE1" pageId="19" pageNumber="20">
|
||
1. Lateral lobes long, exceeding proximal half of median lobe, with very short apical setae;
|
||
<collectingCountry id="F3471E9F9F45FFD1FB13FDB55A032CE1" name="Papua New Guinea" pageId="19" pageNumber="20">Papua New Guinea</collectingCountry>
|
||
...............................................................................................................
|
||
<taxonomicName id="4C50258C9F45FFD1FB97FD985E292CE1" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[1141,1301,632,658]" class="Insecta" family="Limnichidae" genus="Pseudeucinetus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
|
||
<emphasis id="B924821D9F45FFD1FB97FD985E292CE1" bold="true" box="[1141,1301,632,658]" italics="true" pageId="19" pageNumber="20">M. papuanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FAFEFD985E4B2CE1" box="[1308,1399,632,658]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF23FD7B5E4B2CC6" blockId="19.[189,1399,596,693]" box="[193,1399,667,693]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF23FD7B5E4B2CC6" box="[193,1399,667,693]" pageId="19" pageNumber="20">– Lateral lobes short, reaching about proximal third of median lobe, with long apical setae .............. 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFD3C5E4B2D6A" blockId="19.[189,1399,732,794]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFD3C5E4B2D6A" pageId="19" pageNumber="20">
|
||
<keyStep id="30A446AA9F45FFD1FF5FFD3C5E4B2D6A" pageId="19" pageNumber="20">2. Ventral plates almost of same proportion on right and left................................................................ 4 – Apical part of left ventral plates distinctly curved interiorly............................................................. 3</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30A446AA9F45FFD1FF5FFCA15E4B2D9B" pageId="19" pageNumber="20">
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFCA15E4B2D0D" blockId="19.[189,1399,833,1000]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFCA15E4B2D0D" pageId="19" pageNumber="20">
|
||
3. Left ventral plates with thumb-like projection on inner margin of apex; median lobe rather slender;
|
||
<collectingCountry id="F3471E9F9F45FFD1FF08FC845A7C2D0D" box="[234,320,868,894]" name="Taiwan" pageId="19" pageNumber="20">Taiwan</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3471E9F9F45FFD1FE9BFC845A832D0D" box="[377,447,868,894]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
|
||
, Yonaguni-jima..............................................
|
||
<taxonomicName id="4C50258C9F45FFD1FC31FC845FD22D0D" ID-CoL="K68W" authority="Sato, 1994" authorityName="Sato" authorityYear="1994" box="[979,1262,868,894]" class="Insecta" family="Limnichidae" genus="Babalimnichus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="taiwanus" status="comb. nov.">
|
||
<emphasis id="B924821D9F45FFD1FC31FC845F552D0D" bold="true" box="[979,1129,868,894]" italics="true" pageId="19" pageNumber="20">M. taiwanus</emphasis>
|
||
<bibRefCitation id="EFC123FE9F45FFD1FB92FC845FD22D0D" author="Sato M." box="[1136,1262,868,894]" pageId="19" pageNumber="20" pagination="173 - 177" refId="ref14802" refString="Sato M. 1994. Note on the genus Pseudeucinetus Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa Entomological Society 2: 173 - 177." type="journal article" year="1994">Satô, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FB17FC845E4B2D0D" box="[1269,1399,868,894]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF23FC675E4B2D9B" blockId="19.[189,1399,833,1000]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF23FC675E4B2D9B" pageId="19" pageNumber="20">
|
||
– Left ventral plates long and curved interiorly in apical part, without thumb-like projection; median lobe wide, expanded in apical quarter;
|
||
<collectingCountry id="F3471E9F9F45FFD1FBEDFC4B5F692DB6" box="[1039,1109,939,965]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
|
||
, Honshu to Ishigakijima ............................................................................................
|
||
<taxonomicName id="4C50258C9F45FFD1FC24FC2E5FD22D94" authority="(Sato, 1994)" authorityName="Yoshitomi" authorityYear="2019" baseAuthorityName="Sato" baseAuthorityYear="1994" box="[966,1262,973,1000]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="masamii" status="comb. nov.">
|
||
<emphasis id="B924821D9F45FFD1FC24FC2E5F6A2D94" bold="true" box="[966,1110,973,1000]" italics="true" pageId="19" pageNumber="20">M. masamii</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC123FE9F45FFD1FB87FC2D5FD92D94" author="Sato M." box="[1125,1253,973,999]" pageId="19" pageNumber="20" pagination="173 - 177" refId="ref14802" refString="Sato M. 1994. Note on the genus Pseudeucinetus Heller and its new relative (Coleoptera, Limnichidae). Special Bulletin of the Essa Entomological Society 2: 173 - 177." type="journal article" year="1994">Satô, 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FB17FC2D5E4B2D9B" box="[1269,1399,973,1000]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFBEF5E4B2A3F" blockId="19.[189,1399,1038,1100]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFBEF5E4B2A3F" pageId="19" pageNumber="20">
|
||
<keyStep id="30A446AA9F45FFD1FF5FFBEF5E4B2A3F" pageId="19" pageNumber="20">
|
||
4. Apical part of ventral plates separated from median lobe;
|
||
<collectingCountry id="F3471E9F9F45FFD1FC6DFBEE58EC2A5B" box="[911,976,1038,1064]" name="Palau" pageId="19" pageNumber="20">Palau</collectingCountry>
|
||
....................
|
||
<taxonomicName id="4C50258C9F45FFD1FB89FBEF5E292A5A" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[1131,1301,1038,1065]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="palauensis" status="sp. nov.">
|
||
<emphasis id="B924821D9F45FFD1FB89FBEF5E292A5A" bold="true" box="[1131,1301,1038,1065]" italics="true" pageId="19" pageNumber="20">M. palauensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FAFEFBEF5E4B2A5A" box="[1308,1399,1039,1065]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– Apical part of ventral plates connected with median lobe................................................................. 5
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFB935E4B2AC3" blockId="19.[189,1399,1139,1201]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFB935E4B2AC3" pageId="19" pageNumber="20">
|
||
<keyStep id="30A446AA9F45FFD1FF5FFB935E4B2AC3" pageId="19" pageNumber="20">
|
||
5. Apex of median lobe nipple-like; Borneo .......................................................
|
||
<taxonomicName id="4C50258C9F45FFD1FB88FB935E292AFD" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[1130,1301,1139,1166]" class="Insecta" family="Limnichidae" genus="Acontosceles" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="B924821D9F45FFD1FB88FB935E292AFD" bold="true" box="[1130,1301,1139,1166]" italics="true" pageId="19" pageNumber="20">M. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FAFEFB935E4B2AFE" box="[1308,1399,1139,1165]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– Apex of median lobe simply pointed................................................................................................. 6
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFB375E4B2B65" blockId="19.[189,1399,1239,1302]" pageId="19" pageNumber="20">
|
||
<keyLead id="30A1FD3A9F45FFD1FF5FFB375E4B2B65" pageId="19" pageNumber="20">
|
||
<keyStep id="30A446AA9F45FFD1FF5FFB375E4B2B65" pageId="19" pageNumber="20">
|
||
6. Lateral lobe robust, with 2–3 apical setae;
|
||
<collectingRegion id="499490ED9F45FFD1FD3FFB3758322A82" box="[733,782,1239,1265]" country="Indonesia" name="Bali" pageId="19" pageNumber="20">Bali</collectingRegion>
|
||
..................................................
|
||
<taxonomicName id="4C50258C9F45FFD1FB60FB385E292A81" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[1154,1301,1239,1266]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="baliensis" status="sp. nov.">
|
||
<emphasis id="B924821D9F45FFD1FB60FB385E292A81" bold="true" box="[1154,1301,1239,1266]" italics="true" pageId="19" pageNumber="20">M. baliensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FAFEFB385E4B2A81" box="[1308,1399,1240,1266]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
– Lateral lobe slender, with 4 apical setae; Ogasawara Islands..................
|
||
<taxonomicName id="4C50258C9F45FFD1FBCFFB1C5E292B66" authority="Yoshitomi, 2019" authorityName="Yoshitomi" authorityYear="2019" box="[1069,1301,1275,1302]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ogasawaraensis" status="sp. nov.">
|
||
<emphasis id="B924821D9F45FFD1FBCFFB1C5E292B66" bold="true" box="[1069,1301,1275,1302]" italics="true" pageId="19" pageNumber="20">M. ogasawaraensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FAFEFB1C5E4B2B65" box="[1308,1399,1276,1302]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="C34A0D849F45FFD1FF5FFADB590B2B09" pageId="19" pageNumber="20" type="reference_group">
|
||
<paragraph id="8BEF5E0F9F45FFD1FF5FFADB590B2B09" blockId="19.[189,1398,1339,1402]" pageId="19" pageNumber="20">
|
||
<superScript id="7C25F3479F45FFD1FF5FFADB5BFA2B3A" attach="left" box="[189,198,1339,1353]" fontSize="6" pageId="19" pageNumber="20">*</superScript>
|
||
<taxonomicName id="4C50258C9F45FFD1FF2BFADD59C72B24" authority="(Hernando & Ribera, 2003)" authorityName="Yoshitomi" authorityYear="2019" baseAuthorityName="Hernando & Ribera" baseAuthorityYear="2003" box="[201,763,1340,1367]" class="Insecta" family="Limnichidae" genus="Mexico" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="splendens" status="comb. nov.">
|
||
<emphasis id="B924821D9F45FFD1FF2BFADD5AA32B25" box="[201,415,1340,1366]" italics="true" pageId="19" pageNumber="20">Mexico splendens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC123FE9F45FFD1FE50FADC59CF2B24" author="Hernando C. & Ribera I." box="[434,755,1340,1367]" pageId="19" pageNumber="20" pagination="269 - 273" refId="ref14267" refString="Hernando C. & Ribera I. 2003. Babalimnichus splendens sp. n., a new jumping shore beetle from the Tonga Islands (Polynesia) (Coleoptera: Limnichidae, Thaumastodinae). Koleopterologische Rundschau 73: 269 - 273." type="journal article" year="2003">Hernando & Ribera, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2173F669F45FFD1FCE5FADC58B12B24" box="[775,909,1340,1367]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is omitted, because this species is only known from a female specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |