444 lines
68 KiB
XML
444 lines
68 KiB
XML
<document id="F268A935B1BC3C5588CD5F4D2E0DB9D7" ID-DOI="10.1046/j.1096-3642.2003.00046.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5437339" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630436511371" checkinUser="carolina" docAuthor="Huber, Bernhard A." docDate="2003" docId="03F987F6FFEBFFABFF330B8A34A8F8A3" docLanguage="en" docName="j.1096-3642.2003.00046.x.pdf" docOrigin="Zoological Journal of the Linnean Society 137 (2)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2003.00046.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Zatavua madagascariensis Huber 2003" docType="treatment" docVersion="3" lastPageNumber="283" masterDocId="FFC0FF8EFFF8FFBDFFF90D2C3528FFD2" masterDocTitle="Cladistic analysis of Malagasy pholcid spiders reveals generic level endemism: Revision of Zatavua n. gen. and Paramicromerys Millot (Pholcidae, Araneae)" masterLastPageNumber="318" masterPageNumber="261" pageNumber="280" updateTime="1699270910870" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="7B36F2EC3A0A1B43B9763A236435ED7F">
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<mods:title id="43F702A7459D719EAF52C8601DFE74E8">Cladistic analysis of Malagasy pholcid spiders reveals generic level endemism: Revision of Zatavua n. gen. and Paramicromerys Millot (Pholcidae, Araneae)</mods:title>
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<mods:roleTerm id="5F02828454418CEFF07174ECE9119416">Author</mods:roleTerm>
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<mods:namePart id="7BFEE1D54611595276E3D5D320833457">Huber, Bernhard A.</mods:namePart>
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<mods:typeOfResource id="313DEC333C535FB14E1BA9600D7C4EFA">text</mods:typeOfResource>
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<mods:title id="0D9AC60B7F7FB5354A65E9A191AB7067">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="0CDA4EE4DCE8F2731699BB3133DAEEE6">
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<mods:date id="70BA6C43AD100DCF3142AE357F76EB5D">2003</mods:date>
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<mods:detail id="321130EC5D3BED84886D2A84AF823F38" type="pubDate">
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<mods:number id="A322BE7B24D3150BF3A6845B7E308A0B">2003-02-28</mods:number>
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<mods:detail id="43289D954B20E0FCC8F11EB5F83A34ED" type="volume">
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<mods:number id="A26BC4F9DDF0C75DF0C2FC17CACD39DB">137</mods:number>
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<mods:start id="5D91D00759AFD4CA102C05A4E5F15693">261</mods:start>
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<mods:url id="EB76CD2187D5C66CDBFDD1D5D5C099F4">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2003.00046.x</mods:url>
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<mods:classification id="3556564CE1BF86AAB8773B9E23DEC97A">journal article</mods:classification>
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<mods:identifier id="3BAAC2065672DCF2E273B16A51D63099" type="DOI">10.1046/j.1096-3642.2003.00046.x</mods:identifier>
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<mods:identifier id="9EB26844AF58E70F81E17519B71ED3E5" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F987F6FFEBFFABFF330B8A34A8F8A3" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3" httpUri="http://treatment.plazi.org/id/03F987F6FFEBFFABFF330B8A34A8F8A3" lastPageId="22" lastPageNumber="283" pageId="19" pageNumber="280">
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<subSubSection id="C34A656BFFEBFFAEFF330B8A37BDF90C" pageId="19" pageNumber="280" type="nomenclature">
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<paragraph id="8BEF36E0FFEBFFAEFF330B8A37FBF96C" blockId="19.[202,723,1702,1758]" box="[202,723,1702,1726]" pageId="19" pageNumber="280">
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<heading id="D0A7818CFFEBFFAEFF330B8A37FBF96C" box="[202,723,1702,1726]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="280" reason="2">
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<taxonomicName id="4C504D63FFEBFFAEFF330B8A37FBF96C" ID-CoL="5CXMJ" authority="(FAGE, 1945)" authorityName="Huber" authorityYear="2003" baseAuthorityName="Fage" baseAuthorityYear="1945" box="[202,723,1702,1726]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="280" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov.">
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<emphasis id="B924EAF2FFEBFFAEFF330B8A3704F96F" bold="true" box="[202,556,1702,1726]" italics="true" pageId="19" pageNumber="280">ZATAVUA MADAGASCARIENSIS</emphasis>
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<emphasis id="B924EAF2FFEBFFAEFDCD0B8A37FBF96C" bold="true" box="[564,723,1702,1726]" pageId="19" pageNumber="280">
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(
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<bibRefCitation id="EFC14B11FFEBFFAEFDC40B8A3757F96F" author="Fage L" box="[573,639,1702,1726]" pageId="19" pageNumber="280" pagination="301 - 307" refId="ref25029" refString="Fage L. 1945. Arachnides cavernicoles nouveaux de Madagascar. Bulletin du Museum Paris, 2 e Serie 17: 301 - 307." type="journal article" year="1945">FAGE</bibRefCitation>
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, 1945)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BEF36E0FFEBFFAEFEF00BE637BDF90C" blockId="19.[202,723,1702,1758]" box="[265,661,1734,1758]" pageId="19" pageNumber="280">
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<heading id="D0A7818CFFEBFFAEFEF00BE637BDF90C" box="[265,661,1734,1758]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="280" reason="4">
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<emphasis id="B924EAF2FFEBFFAEFEF00BE6345BF90C" bold="true" box="[265,371,1734,1758]" pageId="19" pageNumber="280">N. COMB.</emphasis>
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(
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<figureCitation id="136B2A65FFEBFFAEFE7D0BEA34C9F90C" box="[388,481,1734,1758]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="19" pageNumber="280">FIGS 25</figureCitation>
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,
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<figureCitation id="136B2A65FFEBFFAEFE170BEA3710F90C" box="[494,568,1734,1758]" captionStart="Figures 81–86" captionStartId="18.[142,232,1617,1636]" captionTargetBox="[191,1374,192,1571]" captionTargetId="figure-158@18.[178,1385,168,1584]" captionTargetPageId="18" captionText="Figures 81–86. Zatavua isalo (81–84) and Z. madagascariensis (85, 86). Left male palp in prolateral (81) and retrolateral (82) views, and male chelicerae in frontal (83, 85) and lateral (84, 86) views. ‘e’: embolus; ‘f’: distinctive flap. Scale lines: 0.5 mm (81, 82), 0.3 mm (83–86)." figureDoi="http://doi.org/10.5281/zenodo.5437365" httpUri="https://zenodo.org/record/5437365/files/figure.png" pageId="19" pageNumber="280">85, 86</figureCitation>
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,
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<figureCitation id="136B2A65FFEBFFAEFDBC0BEA37A6F90C" box="[581,654,1734,1758]" captionStart="Figures 87–90" captionStartId="19.[155,245,957,976]" captionTargetBox="[208,1395,200,924]" captionTargetId="figure-456@19.[184,1405,178,977]" captionTargetPageId="19" captionText="Figures 87–90. Zatavua isalo (87–88) and Z. madagascariensis (89, 90). Cleared epigyna in ventral (87, 89) and dorsal (88, 90) views. ‘p’: pockets. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5437367" httpUri="https://zenodo.org/record/5437367/files/figure.png" pageId="19" pageNumber="280">89, 90</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A656BFFEBFFAEFF620BC336AFFB42" pageId="19" pageNumber="280" type="reference_group">
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<paragraph id="8BEF36E0FFEBFFAEFF620BC3340AF8F6" blockId="19.[155,770,1775,1828]" pageId="19" pageNumber="280">
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<treatmentCitationGroup id="AB4011CEFFEBFFAEFF620BC3340AF8F6" pageId="19" pageNumber="280">
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<treatmentCitation id="0AF110F1FFEBFFAEFF620BC337D6F8D6" author="Fage L" box="[155,766,1775,1797]" page="303" pageId="19" pageNumber="280" year="1945">
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<taxonomicName id="4C504D63FFEBFFAEFF620BC337D6F8D6" authority="Fage, 1945: 303" authorityName="Fage" authorityPageNumber="303" authorityYear="1945" box="[155,766,1775,1797]" class="Arachnida" family="Pholcidae" genus="Spermophora" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="280" phylum="Arthropoda" rank="species" species="madagascariensis">
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<emphasis id="B924EAF2FFEBFFAEFF620BC3373EF8D6" box="[155,534,1775,1796]" italics="true" pageId="19" pageNumber="280">Spermophora madagascariensis</emphasis>
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<bibRefCitation id="EFC14B11FFEBFFAEFDD40BC337D6F8D6" author="Fage L" box="[557,766,1775,1797]" pageId="19" pageNumber="280" pagination="301 - 307" refId="ref25029" refString="Fage L. 1945. Arachnides cavernicoles nouveaux de Madagascar. Bulletin du Museum Paris, 2 e Serie 17: 301 - 307." type="journal article" year="1945">Fage, 1945: 303</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, fig. 1(a)–(d).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEF36E0FFEBFFAEFF620A1036AFFB42" blockId="19.[155,771,1852,1904]" lastBlockId="19.[819,1435,1085,1168]" pageId="19" pageNumber="280">
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<emphasis id="B924EAF2FFEBFFAEFF620A1035F3F883" box="[155,219,1852,1873]" italics="true" pageId="19" pageNumber="280">Types</emphasis>
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.
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<materialsCitation id="3B383CBDFFEBFFAEFF150A1036ABFB42" collectionCode="MNHN" collectorName="M. R. Decary" country="Madagascar" location="Manombo valley" pageId="19" pageNumber="280" specimenCount="4" specimenCount-female="3" specimenCount-juvenile="1" stateProvince="Toliara" typeStatus="syntype">
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One male,
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<specimenCount id="9D56FD69FFEBFFAEFE970A1034F6F880" box="[366,478,1852,1874]" pageId="19" pageNumber="280" type="female">3 females</specimenCount>
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and
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<specimenCount id="9D56FD69FFEBFFAEFDE60A1037BAF880" box="[543,658,1852,1874]" pageId="19" pageNumber="280" type="juvenile">1 juvenile</specimenCount>
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<typeStatus id="54EB8842FFEBFFAEFD640A103629F883" box="[669,769,1852,1873]" pageId="19" pageNumber="280" type="syntype">syntypes</typeStatus>
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from grotte d’Andoharano (about 22∞50¢S, 43∞40¢E),
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<location id="8E8F603BFFEBFFAEFCCA091136D5FB81" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFEBFFAEFCCA091136D5FB81" box="[819,1021,1085,1107]" country="Madagascar" name="Manombo valley" pageId="19" pageNumber="280" stateProvince="Toliara">Manombo valley</location>
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,
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<collectingRegion id="4994F802FFEBFFAEFBE109113143FB81" box="[1048,1131,1085,1107]" country="Madagascar" name="Toliara" pageId="19" pageNumber="280">Toliara</collectingRegion>
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,
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<collectingCountry id="F3477670FFEBFFAEFB7E0911303CFB81" box="[1159,1300,1085,1107]" name="Madagascar" pageId="19" pageNumber="280">Madagascar</collectingCountry>
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; collection date unknown (
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<collectorName id="26A55336FFEBFFAEFC0A097031B0FBA3" box="[1011,1176,1116,1137]" pageId="19" pageNumber="280">M. R. Decary</collectorName>
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), in
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<collectionCode id="ED41AE25FFEBFFAEFB2E09703003FBA3" box="[1239,1323,1116,1137]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="280">MNHN</collectionCode>
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(MP AR 10389)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A656BFFEBFFABFCCA098534A8F8A3" lastPageId="22" lastPageNumber="283" pageId="19" pageNumber="280" type="diagnosis">
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<paragraph id="8BEF36E0FFEBFFAEFCCA098536E4FA47" blockId="19.[819,1435,1192,1429]" pageId="19" pageNumber="280">
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<emphasis id="B924EAF2FFEBFFAEFCCA0985368BFB6C" box="[819,931,1193,1214]" italics="true" pageId="19" pageNumber="280">Diagnosis</emphasis>
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. Very closely related to
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<taxonomicName id="4C504D63FFEBFFAEFB4B0985302EFB6F" authorityName="Huber" authorityYear="2003" box="[1202,1286,1192,1214]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="280" phylum="Arthropoda" rank="species" species="isalo">
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<emphasis id="B924EAF2FFEBFFAEFB4B0985302EFB6F" box="[1202,1286,1192,1214]" italics="true" pageId="19" pageNumber="280">Z. isalo</emphasis>
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</taxonomicName>
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, with almost identical palps (cf.
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<figureCitation id="136B2A65FFEBFFAEFBF009EB3199FB0F" box="[1033,1201,1223,1245]" captionStart="Figures 81–86" captionStartId="18.[142,232,1617,1636]" captionTargetBox="[191,1374,192,1571]" captionTargetId="figure-158@18.[178,1385,168,1584]" captionTargetPageId="18" captionText="Figures 81–86. Zatavua isalo (81–84) and Z. madagascariensis (85, 86). Left male palp in prolateral (81) and retrolateral (82) views, and male chelicerae in frontal (83, 85) and lateral (84, 86) views. ‘e’: embolus; ‘f’: distinctive flap. Scale lines: 0.5 mm (81, 82), 0.3 mm (83–86)." figureDoi="http://doi.org/10.5281/zenodo.5437365" httpUri="https://zenodo.org/record/5437365/files/figure.png" pageId="19" pageNumber="280">Figs 81 and 82</figureCitation>
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), but with male cheliceral apophyses and pockets on epigynum less widely spread. This species might be a senior synonym of
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<taxonomicName id="4C504D63FFEBFFAEFA7A082936E1FAEB" authority="(Millot)" baseAuthorityName="Millot" baseAuthorityYear="1946" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="280" phylum="Arthropoda" rank="species" species="fagei">
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<emphasis id="B924EAF2FFEBFFAEFA7A08293642FAEA" italics="true" pageId="19" pageNumber="280">Z. fagei</emphasis>
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(Millot)
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</taxonomicName>
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, which was collected by the same collector but not within the cave of Andoharano but apparently outside (‘région de la grotte d’Andoharano’,
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<bibRefCitation id="EFC14B11FFEBFFAEFCCA085336E8FA47" author="Millot J" box="[819,960,1407,1429]" pageId="19" pageNumber="280" pagination="127 - 158" refId="ref25363" refString="Millot J. 1946. Les pholcides de Madagascar (Araneides). Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie 22: 127 - 158." type="journal article" year="1946">Millot, 1946</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BEF36E0FFEBFFA9FCCA088135D5FEA1" blockId="19.[819,1435,1453,1904]" lastBlockId="20.[142,758,196,371]" lastPageId="20" lastPageNumber="281" pageId="19" pageNumber="280">
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<emphasis id="B924EAF2FFEBFFAEFCCA088136C8FA10" box="[819,992,1453,1474]" italics="true" pageId="19" pageNumber="280">
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Male (
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<typeStatus id="54EB8842FFEBFFAEFC85088136FCFA10" box="[892,980,1453,1474]" pageId="19" pageNumber="280" type="syntype">syntype</typeStatus>
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).
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</emphasis>
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Total length 2.8, carapace width 1.4. Legs 1 and 2 missing (leg 1 according to original description
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<quantity id="4CA89B05FFEBFFAEFC4508C6312FF9D2" box="[956,1031,1514,1536]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="280" unit="cm" value="2.8">2.8 cm</quantity>
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[mm was a lapsus!]), tibia 3: 3.3, tibia 4: 5.0. Habitus as in
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<taxonomicName id="4C504D63FFEBFFAEFBA70B26319AF9CC" authorityName="Huber" authorityYear="2003" box="[1118,1202,1545,1567]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="280" phylum="Arthropoda" rank="species" species="isalo">
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<emphasis id="B924EAF2FFEBFFAEFBA70B26319AF9CC" box="[1118,1202,1545,1567]" italics="true" pageId="19" pageNumber="280">Z. isalo</emphasis>
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||
</taxonomicName>
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(cf.
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<figureCitation id="136B2A65FFEBFFAEFB1B0B2530A6F9CD" box="[1250,1422,1545,1567]" captionStart="Figures 1–18" captionStartId="3.[155,245,1182,1201]" captionTargetBox="[155,1435,198,1150]" captionTargetId="figure-400@3.[155,1435,198,1150]" captionTargetPageId="3" captionText="Figures 1–18. Zatavua habitus, males in dorsal and lateral views. 1–2, Z. griswoldi. 3–4, Z. analalava. 5–6, Z. mahafaly. 7–8, Z. vohiparara. 9–10, Z. zanahary. 11–12, Z. isalo. 13–14, Z. tamatave. 15–16, Z. voahangyae. 17–18, Z. talatakely." figureDoi="http://doi.org/10.5281/zenodo.5437341" httpUri="https://zenodo.org/record/5437341/files/figure.png" pageId="19" pageNumber="280">Figs 11 and 12</figureCitation>
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). Carapace pale ochre-yellow with pair of slightly darker marks parallel to lateral margins; sternum whitish with heart-shaped ochre yellow mark; legs pale ochre-yellow, with slightly darker rings on femora and tibiae (subdistally) and in patella area; opisthosoma monochromous grey. Ocular area elevated and clearly separated from carapace; distance PME-PME 100 Mm; diameter PME 100 Mm; distance PME-ALE 30 Mm. Thoracic furrow distinct but shallow. Sternum wider than long (0.88/0.72). Clypeus unmodified. Chelicerae as in
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<figureCitation id="136B2A65FFEBFFAEFC380A7631A6F8A2" box="[961,1166,1882,1904]" captionStart="Figures 81–86" captionStartId="18.[142,232,1617,1636]" captionTargetBox="[191,1374,192,1571]" captionTargetId="figure-158@18.[178,1385,168,1584]" captionTargetPageId="18" captionText="Figures 81–86. Zatavua isalo (81–84) and Z. madagascariensis (85, 86). Left male palp in prolateral (81) and retrolateral (82) views, and male chelicerae in frontal (83, 85) and lateral (84, 86) views. ‘e’: embolus; ‘f’: distinctive flap. Scale lines: 0.5 mm (81, 82), 0.3 mm (83–86)." figureDoi="http://doi.org/10.5281/zenodo.5437365" httpUri="https://zenodo.org/record/5437365/files/figure.png" pageId="19" pageNumber="280">Figures 85 and 86</figureCitation>
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||
, without modified hairs on tips of apophyses; distance between tips of apophyses 0.45. Palps as in
|
||
<taxonomicName id="4C504D63FFECFFA9FE690DCF34CFFF2A" authorityName="Huber" authorityYear="2003" box="[400,487,227,248]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="isalo">
|
||
<emphasis id="B924EAF2FFECFFA9FE690DCF34CFFF2A" box="[400,487,227,248]" italics="true" pageId="20" pageNumber="281">Z. isalo</emphasis>
|
||
</taxonomicName>
|
||
, but flat cuticular flap retrolaterally on procursus (‘f’ in
|
||
<figureCitation id="136B2A65FFECFFA9FDD40C2D3752FEC5" box="[557,634,257,279]" captionStart="Figures 81–86" captionStartId="18.[142,232,1617,1636]" captionTargetBox="[191,1374,192,1571]" captionTargetId="figure-158@18.[178,1385,168,1584]" captionTargetPageId="18" captionText="Figures 81–86. Zatavua isalo (81–84) and Z. madagascariensis (85, 86). Left male palp in prolateral (81) and retrolateral (82) views, and male chelicerae in frontal (83, 85) and lateral (84, 86) views. ‘e’: embolus; ‘f’: distinctive flap. Scale lines: 0.5 mm (81, 82), 0.3 mm (83–86)." figureDoi="http://doi.org/10.5281/zenodo.5437365" httpUri="https://zenodo.org/record/5437365/files/figure.png" pageId="20" pageNumber="281">Fig. 82</figureCitation>
|
||
) narrower in present species. Legs without spines, without curved hairs, few vertical hairs (most hairs missing, however).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770CA7379FFE0C" blockId="20.[142,758,395,479]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770CA735F7FE72" box="[142,223,395,416]" italics="true" pageId="20" pageNumber="281">Female</emphasis>
|
||
. In general similar to male. Tibia
|
||
<quantity id="4CA89B05FFECFFA9FD8E0CA7378DFE73" box="[631,677,395,417]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="281" unit="in" value="1.0">1 in</quantity>
|
||
a
|
||
<typeStatus id="54EB8842FFECFFA9FD3C0CA035EAFE6D" pageId="20" pageNumber="281" type="syntype">syntype</typeStatus>
|
||
: 7.3. Epigynum as in
|
||
<figureCitation id="136B2A65FFECFFA9FE450C86371DFE12" box="[444,565,426,448]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">Figures 25</figureCitation>
|
||
and
|
||
<figureCitation id="136B2A65FFECFFA9FD970C8637A3FE6D" box="[622,651,426,447]" captionStart="Figures 87–90" captionStartId="19.[155,245,957,976]" captionTargetBox="[208,1395,200,924]" captionTargetId="figure-456@19.[184,1405,178,977]" captionTargetPageId="19" captionText="Figures 87–90. Zatavua isalo (87–88) and Z. madagascariensis (89, 90). Cleared epigyna in ventral (87, 89) and dorsal (88, 90) views. ‘p’: pockets. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5437367" httpUri="https://zenodo.org/record/5437367/files/figure.png" pageId="20" pageNumber="281">89</figureCitation>
|
||
; distance between pockets 0.34. Dorsal view as in
|
||
<figureCitation id="136B2A65FFECFFA9FD9A0CE5379BFE0C" box="[611,691,457,479]" captionStart="Figures 87–90" captionStartId="19.[155,245,957,976]" captionTargetBox="[208,1395,200,924]" captionTargetId="figure-456@19.[184,1405,178,977]" captionTargetPageId="19" captionText="Figures 87–90. Zatavua isalo (87–88) and Z. madagascariensis (89, 90). Cleared epigyna in ventral (87, 89) and dorsal (88, 90) views. ‘p’: pockets. Scale lines: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5437367" httpUri="https://zenodo.org/record/5437367/files/figure.png" pageId="20" pageNumber="281">Fig. 90</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770CDB37C6FDDF" blockId="20.[142,750,503,525]" box="[142,750,503,525]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770CDB3431FDDE" box="[142,281,503,524]" italics="true" pageId="20" pageNumber="281">Distribution</emphasis>
|
||
. Known only from
|
||
<typeStatus id="54EB8842FFECFFA9FE0E0CDB3700FDDE" box="[503,552,503,524]" pageId="20" pageNumber="281">type</typeStatus>
|
||
locality (Map 1).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770F0934CFFD8B" blockId="20.[142,758,549,601]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770F09344EFDE8" box="[142,358,549,570]" italics="true" pageId="20" pageNumber="281">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B383CBDFFECFFA9FE5C0F0934CBFD8B" country="Madagascar" location="Manombo valley" pageId="20" pageNumber="281" specimenCount="1" stateProvince="Toliara">
|
||
<collectingCountry id="F3477670FFECFFA9FE5C0F093748FDE8" box="[421,608,549,570]" name="Madagascar" pageId="20" pageNumber="281">MADAGASCAR</collectingCountry>
|
||
:
|
||
<emphasis id="B924EAF2FFECFFA9FD650F0937DEFDE8" box="[668,758,549,570]" italics="true" pageId="20" pageNumber="281">
|
||
<collectingRegion id="4994F802FFECFFA9FD650F0937D8FDE8" box="[668,752,549,570]" country="Madagascar" name="Toliara" pageId="20" pageNumber="281">Toliara</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<location id="8E8F603BFFECFFA9FF770F6F3466FD8B" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFECFFA9FF770F6F3466FD8B" box="[142,334,579,601]" country="Madagascar" name="Manombo valley" pageId="20" pageNumber="281" stateProvince="Toliara">Manombo valley</location>
|
||
:
|
||
<typeStatus id="54EB8842FFECFFA9FEA00F6834BEFD8B" box="[345,406,580,601]" pageId="20" pageNumber="281">types</typeStatus>
|
||
above
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF0B0FA037BAFD71" blockId="20.[242,658,652,708]" box="[242,658,652,676]" pageId="20" pageNumber="281">
|
||
<taxonomicName id="4C504D63FFECFFA9FF0B0FA034CCFD71" authority="Huber, 2003" authorityName="Huber" authorityYear="2003" box="[242,484,652,676]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="mahafaly" status="sp. nov.">
|
||
<emphasis id="B924EAF2FFECFFA9FF0B0FA034CCFD71" bold="true" box="[242,484,652,676]" italics="true" pageId="20" pageNumber="281">ZATAVUA MAHAFALY</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B924EAF2FFECFFA9FE1D0FA037BAFD71" bold="true" box="[484,658,652,676]" pageId="20" pageNumber="281">
|
||
,
|
||
<taxonomicNameLabel id="A2175789FFECFFA9FE0D0FBC37BAFD71" box="[500,658,656,675]" pageId="20" pageNumber="281" rank="species">NEW SPECIES</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FEBD0F803768FD16" blockId="20.[242,658,652,708]" box="[324,576,684,708]" pageId="20" pageNumber="281">
|
||
(
|
||
<figureCitation id="136B2A65FFECFFA9FEB50F803490FD16" box="[332,440,684,708]" captionStart="Figures 1–18" captionStartId="3.[155,245,1182,1201]" captionTargetBox="[155,1435,198,1150]" captionTargetId="figure-400@3.[155,1435,198,1150]" captionTargetPageId="3" captionText="Figures 1–18. Zatavua habitus, males in dorsal and lateral views. 1–2, Z. griswoldi. 3–4, Z. analalava. 5–6, Z. mahafaly. 7–8, Z. vohiparara. 9–10, Z. zanahary. 11–12, Z. isalo. 13–14, Z. tamatave. 15–16, Z. voahangyae. 17–18, Z. talatakely." figureDoi="http://doi.org/10.5281/zenodo.5437341" httpUri="https://zenodo.org/record/5437341/files/figure.png" pageId="20" pageNumber="281">FIGS 5, 6</figureCitation>
|
||
,
|
||
<figureCitation id="136B2A65FFECFFA9FE3C0F8034CCFD16" box="[453,484,684,708]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">27</figureCitation>
|
||
,
|
||
<figureCitation id="136B2A65FFECFFA9FE080F803710FD16" box="[497,568,684,708]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">91–95</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770FF137BCFC9C" blockId="20.[142,758,733,847]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770FF135E3FD20" box="[142,203,733,754]" italics="true" pageId="20" pageNumber="281">Type.</emphasis>
|
||
<materialsCitation id="3B383CBDFFECFFA9FF2F0FF137B8FC9C" collectingDate="1992-09-15" collectingDateMax="1992-09-16" collectingDateMin="1992-09-15" collectionCode="CAS" collectorName="V. & B. Roth" country="Madagascar" county="Mahafaly" location="Male" municipality="Lac Tsimanampetsoa" pageId="20" pageNumber="281" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
|
||
<location id="8E8F603BFFECFFA9FF2F0FF13427FD21" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFECFFA9FF2F0FF13427FD21" box="[214,271,733,755]" country="Madagascar" county="Mahafaly" municipality="Lac Tsimanampetsoa" name="Male" pageId="20" pageNumber="281" stateProvince="Toliara">Male</location>
|
||
<typeStatus id="54EB8842FFECFFA9FEE20FF13454FD21" box="[283,380,733,755]" pageId="20" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
from near
|
||
<location id="8E8F603BFFECFFA9FDF00FF13777FD21" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFECFFA9FDF00FF13777FD21" box="[521,607,733,755]" country="Madagascar" county="Mahafaly" municipality="Lac Tsimanampetsoa" name="Eloetse" pageId="20" pageNumber="281" stateProvince="Toliara">Eloetse</location>
|
||
, by
|
||
<collectingMunicipality id="6B8BAC9AFFECFFA9FD6F0FF13406FCC3" pageId="20" pageNumber="281">Lac Tsimanampetsoa</collectingMunicipality>
|
||
(24∞10¢S, 43∞45¢E),
|
||
<collectingCounty id="628E4E6CFFECFFA9FDD80FD737A5FCC3" box="[545,653,763,785]" pageId="20" pageNumber="281">Mahafaly</collectingCounty>
|
||
,
|
||
<collectingRegion id="4994F802FFECFFA9FD640FD737D9FCC3" box="[669,753,763,785]" country="Madagascar" name="Toliara" pageId="20" pageNumber="281">Toliara</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3477670FFECFFA9FF770E363433FCE2" box="[142,283,794,816]" name="Madagascar" pageId="20" pageNumber="281">Madagascar</collectingCountry>
|
||
;
|
||
<date id="FFEE1020FFECFFA9FEDF0E363718FCFD" box="[294,560,794,816]" pageId="20" pageNumber="281" value="1992-09-15" valueMax="1992-09-16" valueMin="1992-09-15">
|
||
<collectingDate id="EFAAE9C8FFECFFA9FEDF0E363718FCFD" box="[294,560,794,816]" pageId="20" pageNumber="281" value="1992-09-15" valueMax="1992-09-16" valueMin="1992-09-15">September 15–16, 1992</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26A55336FFECFFA9FDC70E3637E4FCE2" box="[574,716,794,816]" pageId="20" pageNumber="281">V. & B. Roth</collectorName>
|
||
), in back of shelter cave, shaded to dark; in
|
||
<collectionCode id="ED41AE25FFECFFA9FDA30E1537B8FC9C" box="[602,656,825,846]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="281" type="Museum">CAS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770E4B34CDFC49" blockId="20.[142,758,871,923]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770E4B342DFCAE" box="[142,261,871,892]" italics="true" pageId="20" pageNumber="281">Etymology</emphasis>
|
||
. Named for the
|
||
<typeStatus id="54EB8842FFECFFA9FE370E4B34D7FCAE" box="[462,511,871,892]" pageId="20" pageNumber="281">type</typeStatus>
|
||
locality. The specific name is a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF770E983407FBB1" blockId="20.[142,758,947,1123]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770E9835D6FC1B" box="[142,254,948,969]" italics="true" pageId="20" pageNumber="281">Diagnosis</emphasis>
|
||
. Large species with globular opisthosoma, easily distinguished from known congeners by the widely spread male cheliceral apophyses (
|
||
<figureCitation id="136B2A65FFECFFA9FD940EDD3795FBD5" box="[621,701,1009,1031]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Fig. 93</figureCitation>
|
||
), the globular male palpal tibia (
|
||
<figureCitation id="136B2A65FFECFFA9FE3509233753FBF7" box="[460,635,1039,1061]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Figs 91 and 92</figureCitation>
|
||
), the massive curved procursus (
|
||
<figureCitation id="136B2A65FFECFFA9FE5B090234D9FB96" box="[418,497,1070,1092]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Fig. 92</figureCitation>
|
||
), and the wide epigynum (
|
||
<figureCitation id="136B2A65FFECFFA9FF2B0961340AFBB1" box="[210,290,1101,1123]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">Fig. 27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FF77095734EEF8A3" blockId="20.[142,758,1147,1905]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FF770957346DFB42" box="[142,325,1147,1168]" italics="true" pageId="20" pageNumber="281">
|
||
Male (
|
||
<typeStatus id="54EB8842FFECFFA9FF2E09573411FB42" box="[215,313,1147,1168]" pageId="20" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Total length 4.8, carapace width 2.1. Leg 1: 50.2 (13.5 + 0.8 + 14.0 + 20.2 + 1.7), tibia 2: 9.5, tibia 3: 6.5, tibia 4: 8.5; tibia 1 L/d: 78. Habitus as in
|
||
<figureCitation id="136B2A65FFECFFA9FF7709FB3469FB3F" box="[142,321,1239,1261]" captionStart="Figures 1–18" captionStartId="3.[155,245,1182,1201]" captionTargetBox="[155,1435,198,1150]" captionTargetId="figure-400@3.[155,1435,198,1150]" captionTargetPageId="3" captionText="Figures 1–18. Zatavua habitus, males in dorsal and lateral views. 1–2, Z. griswoldi. 3–4, Z. analalava. 5–6, Z. mahafaly. 7–8, Z. vohiparara. 9–10, Z. zanahary. 11–12, Z. isalo. 13–14, Z. tamatave. 15–16, Z. voahangyae. 17–18, Z. talatakely." figureDoi="http://doi.org/10.5281/zenodo.5437341" httpUri="https://zenodo.org/record/5437341/files/figure.png" pageId="20" pageNumber="281">Figures 5 and 6</figureCitation>
|
||
. Carapace pale ochre with light brown mark posteriorly; sternum pale ochre with dark margin. Legs ochre to light brown, patella area darker. Opisthosoma pale ochre. Ocular area slightly elevated, not clearly separated from carapace; distance PME-PME 150 Mm; diameter PME 110 Mm; distance PME-ALE 40 Mm. Thoracic furrow absent, only shallow indentation behind ocular area. Clypeus not modified. Sternum wider than long (1.3/1.0). Chelicerae as in
|
||
<figureCitation id="136B2A65FFECFFA9FF5408C73435F9D2" box="[173,285,1515,1537]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Figure 93</figureCitation>
|
||
, proximal apophyses directed backwards as in
|
||
<taxonomicName id="4C504D63FFECFFA9FF250B2634AFF9CC" authorityName="Huber" authorityYear="2003" box="[220,391,1545,1567]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="voahangyae">
|
||
<emphasis id="B924EAF2FFECFFA9FF250B2634AFF9CC" box="[220,391,1545,1567]" italics="true" pageId="20" pageNumber="281">Z. voahangyae</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="136B2A65FFECFFA9FE3F0B25373DF9CD" box="[454,533,1545,1567]" captionStart="Figures 60–65" captionStartId="15.[155,245,1561,1580]" captionTargetBox="[174,1413,155,1523]" captionTargetId="figure-176@15.[174,1413,149,1541]" captionTargetPageId="15" captionText="Figures 60–65. Zatavua voahangyae. Left male palp in prolateral (60) and retrolateral (61) views, male chelicerae in lateral (62) frontal (63) views, and cleared epigynum in ventral (64) and dorsal (65) views. ‘e’: embolus; ‘p’: pocket; ‘s’: spine on embolus; ‘to’: tarsal organ. Scale lines: 0.5 mm (60–61), 0.2 mm (62–63), 0.3 mm (64–65)." figureDoi="http://doi.org/10.5281/zenodo.5437359" httpUri="https://zenodo.org/record/5437359/files/figure.png" pageId="20" pageNumber="281">Fig. 62</figureCitation>
|
||
), distal apophyses widely spread, with sclerotized teeth at tip, without modified hairs; distance between outer tips of apophyses 1.95. Palps as in
|
||
<figureCitation id="136B2A65FFECFFA9FE6A0B493744F9A8" box="[403,620,1637,1659]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Figures 91 and 92</figureCitation>
|
||
, trochanter with small sclerotized hump ventrally, femur with distinct hump proximo-ventrally, tibia almost globular, procursus massive with complex tip. Embolus (‘e’ in
|
||
<figureCitation id="136B2A65FFECFFA9FF770BCC35F3F924" box="[142,219,1760,1782]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Fig. 91</figureCitation>
|
||
) partly sclerotized, with subdistal conical projection but without spine. Legs without spines, without curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 5%; tarsus 1 with>10 very indistinct pseudosegments.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0DE83652FEA1" blockId="20.[806,1422,196,371]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0DE8365FFF0B" box="[806,887,196,217]" italics="true" pageId="20" pageNumber="281">Female</emphasis>
|
||
. In general similar to male, but sternum with light brown marks. Tibia 1: 11.5, tibia 2: 7.3, tibia 3: 5.5, tibia 4: 7.3. Epigynum as in
|
||
<figureCitation id="136B2A65FFECFFA9FB630C2D303BFEC4" box="[1178,1299,257,279]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">Figures 27</figureCitation>
|
||
and
|
||
<figureCitation id="136B2A65FFECFFA9FAB40C2D3042FEC4" box="[1357,1386,257,278]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">94</figureCitation>
|
||
; at lateral extremes with sclerotized ridges (‘r’ in
|
||
<figureCitation id="136B2A65FFECFFA9FAC30C0C30A2FEE7" box="[1338,1418,288,310]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Fig. 94</figureCitation>
|
||
; distance between ridges 1.62). Dorsal view as in
|
||
<figureCitation id="136B2A65FFECFFA9FCDF0C71365EFEA1" box="[806,886,349,371]" captionStart="Figures 91–95" captionStartId="21.[155,245,1825,1844]" captionTargetBox="[202,1392,179,1792]" captionTargetId="figure-76@21.[180,1408,176,1814]" captionTargetPageId="21" captionText="Figures 91–95. Zatavua mahafaly. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93)." figureDoi="http://doi.org/10.5281/zenodo.5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="20" pageNumber="281">Fig. 95</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0CA736AFFE12" blockId="20.[806,1420,395,448]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0CA73699FE72" box="[806,945,395,416]" italics="true" pageId="20" pageNumber="281">Distribution</emphasis>
|
||
. Known only from
|
||
<typeStatus id="54EB8842FFECFFA9FB100CA03032FE73" box="[1257,1306,396,417]" pageId="20" pageNumber="281">type</typeStatus>
|
||
locality (Map 1).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0CF43055FDDF" blockId="20.[806,1422,472,525]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0CF43137FE3F" box="[806,1055,472,493]" italics="true" pageId="20" pageNumber="281">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B383CBDFFECFFA9FBB70CF43051FDDF" country="Madagascar" location="Mahafaly" pageId="20" pageNumber="281" specimenCount="1" specimenCount-female="1" stateProvince="Toliara">
|
||
<collectingCountry id="F3477670FFECFFA9FBB70CF43021FE3F" box="[1102,1289,472,493]" name="Madagascar" pageId="20" pageNumber="281">MADAGASCAR</collectingCountry>
|
||
:
|
||
<emphasis id="B924EAF2FFECFFA9FACD0CF430A6FE3F" box="[1332,1422,472,493]" italics="true" pageId="20" pageNumber="281">
|
||
<collectingRegion id="4994F802FFECFFA9FACD0CF430A0FE3F" box="[1332,1416,472,493]" country="Madagascar" name="Toliara" pageId="20" pageNumber="281">Toliara</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<location id="8E8F603BFFECFFA9FCDF0CDB36BEFDDF" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFECFFA9FCDF0CDB36BEFDDF" box="[806,918,503,525]" country="Madagascar" name="Mahafaly" pageId="20" pageNumber="281" stateProvince="Toliara">Mahafaly</location>
|
||
:
|
||
<typeStatus id="54EB8842FFECFFA9FC580CDB36FAFDDE" box="[929,978,503,524]" pageId="20" pageNumber="281">type</typeStatus>
|
||
above, together with
|
||
<specimenCount id="9D56FD69FFECFFA9FB370CDB31D9FDDE" box="[1230,1265,502,524]" pageId="20" pageNumber="281" type="female">1♀</specimenCount>
|
||
, same data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FC520F133022FD84" blockId="20.[939,1290,575,631]" box="[939,1290,575,599]" pageId="20" pageNumber="281">
|
||
<taxonomicName id="4C504D63FFECFFA9FC520F133174FD84" authority="Huber, 2003" authorityName="Huber" authorityYear="2003" box="[939,1116,575,599]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="kely" status="sp. nov.">
|
||
<emphasis id="B924EAF2FFECFFA9FC520F133174FD84" bold="true" box="[939,1116,575,599]" italics="true" pageId="20" pageNumber="281">ZATAVUA KELY</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B924EAF2FFECFFA9FBA50F133022FD84" bold="true" box="[1116,1290,575,599]" pageId="20" pageNumber="281">
|
||
,
|
||
<taxonomicNameLabel id="A2175789FFECFFA9FB920F6F3022FD84" box="[1131,1290,579,598]" pageId="20" pageNumber="281" rank="species">NEW SPECIES</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FC0B0F7331E9FDA5" blockId="20.[939,1290,575,631]" box="[1010,1217,607,631]" pageId="20" pageNumber="281">
|
||
(
|
||
<figureCitation id="136B2A65FFECFFA9FC020F733170FDA5" box="[1019,1112,607,631]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">FIGS 28</figureCitation>
|
||
,
|
||
<figureCitation id="136B2A65FFECFFA9FB9D0F733191FDA5" box="[1124,1209,607,631]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">96–101</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0FBD31BEFD30" blockId="20.[806,1422,656,739]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0FBD364BFD74" box="[806,867,657,678]" italics="true" pageId="20" pageNumber="281">Type.</emphasis>
|
||
<materialsCitation id="3B383CBDFFECFFA9FC920FBC31BAFD30" collectingDate="1992-08-20" collectingDateMax="1992-08-26" collectingDateMin="1992-08-20" collectionCode="CAS" collectorName="V. & B. Roth" country="Madagascar" location="Male" municipality="English Camp" pageId="20" pageNumber="281" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
|
||
<location id="8E8F603BFFECFFA9FC920FBC368CFD74" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFECFFA9FC920FBC368CFD74" box="[875,932,656,678]" country="Madagascar" municipality="English Camp" name="Male" pageId="20" pageNumber="281" stateProvince="Antsiranana">Male</location>
|
||
<typeStatus id="54EB8842FFECFFA9FC540FBC3126FD74" box="[941,1038,656,678]" pageId="20" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
from
|
||
<collectingMunicipality id="6B8BAC9AFFECFFA9FBAC0FBC31D2FD77" box="[1109,1274,656,678]" pageId="20" pageNumber="281">English Camp</collectingMunicipality>
|
||
(12∞54¢34≤S, 49∞06¢36≤E),
|
||
<collectingRegion id="4994F802FFECFFA9FC390F83317AFD17" box="[960,1106,687,709]" country="Madagascar" name="Antsiranana" pageId="20" pageNumber="281">Antsiranana</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3477670FFECFFA9FB9C0F8331DAFD17" box="[1125,1266,687,709]" name="Madagascar" pageId="20" pageNumber="281">Madagascar</collectingCountry>
|
||
;
|
||
<date id="FFEE1020FFECFFA9FAFD0F8336AFFD30" pageId="20" pageNumber="281" value="1992-08-20" valueMax="1992-08-26" valueMin="1992-08-20">
|
||
<collectingDate id="EFAAE9C8FFECFFA9FAFD0F8336AFFD30" pageId="20" pageNumber="281" value="1992-08-20" valueMax="1992-08-26" valueMin="1992-08-20">August 20– 26, 1992</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="26A55336FFECFFA9FC6F0FE13101FD31" box="[918,1065,717,739]" pageId="20" pageNumber="281">V. & B. Roth</collectorName>
|
||
); in
|
||
<collectionCode id="ED41AE25FFECFFA9FBA50FE131BAFD30" box="[1116,1170,717,738]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="281" type="Museum">CAS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0FD736EBFC9D" blockId="20.[806,1421,763,847]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0FD736B5FCC2" box="[806,925,763,784]" italics="true" pageId="20" pageNumber="281">Etymology</emphasis>
|
||
. ‘Kely’ is Malagasy for small and refers to the size of this species. The word is here used as noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF0E4B36A3FB81" blockId="20.[806,1422,871,1107]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0E4B36BEFCAE" box="[806,918,871,892]" italics="true" pageId="20" pageNumber="281">Diagnosis</emphasis>
|
||
. Small species with globular opisthosoma, easily distinguished from known congeners by the widely spread male cheliceral apophyses (
|
||
<figureCitation id="136B2A65FFECFFA9FAE10E88304FFC6B" box="[1304,1383,932,954]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Fig. 99</figureCitation>
|
||
;
|
||
<taxonomicName id="4C504D63FFECFFA9FA8E0E8936BAFC0A" authorityName="Huber" authorityYear="2003" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="mahafaly">
|
||
<emphasis id="B924EAF2FFECFFA9FA8E0E8936BAFC0A" italics="true" pageId="20" pageNumber="281">Z. mahafaly</emphasis>
|
||
</taxonomicName>
|
||
has similar chelicerae but is much larger), the simple spine-like procursus (‘pr’ in
|
||
<figureCitation id="136B2A65FFECFFA9FB050ECD3063FC25" box="[1276,1355,993,1015]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Fig. 97</figureCitation>
|
||
), and the epigynum with lateral sclerotized structures (‘r’ in
|
||
<figureCitation id="136B2A65FFECFFA9FCDF093336AAFBE6" box="[806,898,1055,1077]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Fig. 101</figureCitation>
|
||
; receptacles?) shining through cuticle (
|
||
<figureCitation id="136B2A65FFECFFA9FCD709113656FB81" box="[814,894,1085,1107]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFA9FCDF094736FAF8F6" blockId="20.[806,1422,1131,1828]" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF094736F5FB52" box="[806,989,1131,1152]" italics="true" pageId="20" pageNumber="281">
|
||
Male (
|
||
<typeStatus id="54EB8842FFECFFA9FC96094736F8FB52" box="[879,976,1131,1152]" pageId="20" pageNumber="281" type="holotype">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Total length 1.3, carapace width 0.6. Leg 1: 7.7 (2.0 + 0.2 + 2.2 + 2.6 + 0.7), tibia 2: 1.2, tibia 3: 0.8, tibia 4: 1.5; tibia 1 L/d: 37. Habitus similar to
|
||
<taxonomicName id="4C504D63FFECFFA9FA8E09853674FB0E" authorityName="Huber" authorityYear="2003" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="281" phylum="Arthropoda" rank="species" species="isalo">
|
||
<emphasis id="B924EAF2FFECFFA9FA8E09853674FB0E" italics="true" pageId="20" pageNumber="281">Z. isalo</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation id="136B2A65FFECFFA9FC7509EB311FFB0F" box="[908,1079,1223,1245]" captionStart="Figures 1–18" captionStartId="3.[155,245,1182,1201]" captionTargetBox="[155,1435,198,1150]" captionTargetId="figure-400@3.[155,1435,198,1150]" captionTargetPageId="3" captionText="Figures 1–18. Zatavua habitus, males in dorsal and lateral views. 1–2, Z. griswoldi. 3–4, Z. analalava. 5–6, Z. mahafaly. 7–8, Z. vohiparara. 9–10, Z. zanahary. 11–12, Z. isalo. 13–14, Z. tamatave. 15–16, Z. voahangyae. 17–18, Z. talatakely." figureDoi="http://doi.org/10.5281/zenodo.5437341" httpUri="https://zenodo.org/record/5437341/files/figure.png" pageId="20" pageNumber="281">Figs 11 and 12</figureCitation>
|
||
). Prosoma and legs ochre-yellow. Opisthosoma ochre-grey, with slightly darker spots shining through cuticle. Ocular area slightly elevated, not clearly separated from carapace; distance PME- PME 50 Mm; diameter PME 50 Mm; distance PME-ALE 20 Mm. Thoracic furrow present, but shallow and indistinct. Clypeus not modified. Sternum wider than long (0.44/0.32). Chelicerae as in
|
||
<figureCitation id="136B2A65FFECFFA9FB9708B23068FA61" box="[1134,1344,1438,1460]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Figures 98 and 99</figureCitation>
|
||
, distal apophyses widely spread, without modified hairs but striated cuticle at tips of apophyses; distance between tips of apophyses 0.45. Palps as in
|
||
<figureCitation id="136B2A65FFECFFA9FB4E08D630A1F9DD" box="[1207,1417,1530,1552]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Figures 96 and 97</figureCitation>
|
||
; trochanter with small conical projection ventrally, procursus very simple, ending in sclerotized spine that is hidden in fold of embolus (‘e’ in
|
||
<figureCitation id="136B2A65FFECFFA9FB3D0B7A303CF9BE" box="[1220,1300,1622,1644]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Fig. 97</figureCitation>
|
||
). Embolus mostly sclerotized, with curved terminal apophysis (‘a’ in
|
||
<figureCitation id="136B2A65FFECFFA9FCB80BBF36B8F97B" box="[833,912,1683,1705]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="20" pageNumber="281">Fig. 96</figureCitation>
|
||
); sperm duct apparently opens at basis of apophysis. Legs without spines, without curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 not seen; tarsus 1 with>10 pseudosegments (only distally fairly distinct).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFECFFABFCDF0A10371EF905" blockId="20.[806,1422,1852,1905]" lastBlockId="22.[142,758,1668,1751]" lastPageId="22" lastPageNumber="283" pageId="20" pageNumber="281">
|
||
<emphasis id="B924EAF2FFECFFA9FCDF0A10365FF883" box="[806,887,1852,1873]" italics="true" pageId="20" pageNumber="281">Female</emphasis>
|
||
. In general similar to male. Tibia 1:
|
||
<quantity id="4CA89B05FFECFFA9FAEC0A10307CF880" box="[1301,1364,1852,1874]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="20" pageNumber="281" unit="in" value="1.8">1.8 in</quantity>
|
||
both females seen. Epigynum as in
|
||
<figureCitation id="136B2A65FFECFFA9FB500A7731D0F8A2" box="[1193,1272,1883,1905]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="20" pageNumber="281">Fig. 28</figureCitation>
|
||
, protruding; with pair of lateral pockets (‘p’ in
|
||
<figureCitation id="136B2A65FFEEFFABFDDC0BA83757F948" box="[549,639,1668,1690]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="22" pageNumber="283">Fig. 101</figureCitation>
|
||
) and oval structures that shine through cuticle (
|
||
<figureCitation id="136B2A65FFEEFFABFD900B8F3794F96A" box="[617,700,1699,1721]" captionStart="Figures 19–28" captionStartId="4.[142,232,1020,1039]" captionTargetBox="[144,1422,197,987]" captionTargetId="figure-471@4.[142,1422,197,987]" captionTargetPageId="4" captionText="Figures 19–28. Zatavua, female opisthosomata, ventral views with epigynum. 19, Z. griswoldi. 20, Z. vohiparara. 21, Z. tamatave. 22, Z. analalava. 23, Z. zanahary. 24, Z. voahangyae. 25, Z. madagascariensis. 26, Z. isalo. 27, Z. mahafaly. 28, Z. kely." figureDoi="http://doi.org/10.5281/zenodo.5437343" httpUri="https://zenodo.org/record/5437343/files/figure.png" pageId="22" pageNumber="283">Figs 28</figureCitation>
|
||
and
|
||
<figureCitation id="136B2A65FFEEFFABFF770BED3592F905" box="[142,186,1729,1751]" captionStart="Figures 96–101" captionStartId="22.[142,232,1505,1524]" captionTargetBox="[183,1379,198,1461]" captionTargetId="figure-80@22.[175,1387,171,1472]" captionTargetPageId="22" captionText="Figures 96–101. Zatavua kely. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="22" pageNumber="283">100</figureCitation>
|
||
). Distance between pockets 0.35.
|
||
</paragraph>
|
||
<caption id="DF2F6668FFEDFFA8FF620A0D37A1F8BD" ID-DOI="http://doi.org/10.5281/zenodo.5437369" ID-Zenodo-Dep="5437369" httpUri="https://zenodo.org/record/5437369/files/figure.png" pageId="21" pageNumber="282" startId="21.[155,245,1825,1844]" targetBox="[202,1392,179,1792]" targetPageId="21">
|
||
<paragraph id="8BEF36E0FFEDFFA8FF620A0D37A1F8BD" blockId="21.[155,1434,1825,1904]" pageId="21" pageNumber="282">
|
||
<emphasis id="B924EAF2FFEDFFA8FF620A0D3469F8E6" bold="true" box="[155,321,1825,1844]" pageId="21" pageNumber="282">Figures 91–95.</emphasis>
|
||
<taxonomicName id="4C504D63FFEDFFA8FEBE0A0E3721F8E7" authorityName="Huber" authorityYear="2003" box="[327,521,1826,1845]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="282" phylum="Arthropoda" rank="species" species="mahafaly">
|
||
<emphasis id="B924EAF2FFEDFFA8FEBE0A0E3721F8E7" box="[327,521,1826,1845]" italics="true" pageId="21" pageNumber="282">Zatavua mahafaly</emphasis>
|
||
</taxonomicName>
|
||
. Left male palp in prolateral (91) and retrolateral (92) views, male chelicerae in frontal view (93), and cleared epigynum in ventral (94) and dorsal (95) views. ‘e’: embolus; ‘r’: sclerotized ridge; ‘to’: tarsal organ. Scale lines: 1 mm (91, 92, 94, 95), 0.5 mm (93).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2F6668FFEEFFABFF7708CD310DF9FD" ID-DOI="http://doi.org/10.5281/zenodo.5437371" ID-Zenodo-Dep="5437371" httpUri="https://zenodo.org/record/5437371/files/figure.png" pageId="22" pageNumber="283" startId="22.[142,232,1505,1524]" targetBox="[183,1379,198,1461]" targetPageId="22">
|
||
<paragraph id="8BEF36E0FFEEFFABFF7708CD310DF9FD" blockId="22.[142,1422,1505,1583]" pageId="22" pageNumber="283">
|
||
<emphasis id="B924EAF2FFEEFFABFF7708CD3469FA26" bold="true" box="[142,321,1505,1524]" pageId="22" pageNumber="283">Figures 96–101.</emphasis>
|
||
<taxonomicName id="4C504D63FFEEFFABFEBE08CE34E7FA26" authorityName="Huber" authorityYear="2003" box="[327,463,1505,1524]" class="Arachnida" family="Pholcidae" genus="Zatavua" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="283" phylum="Arthropoda" rank="species" species="kely">
|
||
<emphasis id="B924EAF2FFEEFFABFEBE08CE34E7FA26" box="[327,463,1505,1524]" italics="true" pageId="22" pageNumber="283">Zatavua kely</emphasis>
|
||
</taxonomicName>
|
||
. Left male palp in prolateral (96) and retrolateral (97) views, male chelicerae in lateral (98) and frontal (99) views, and cleared epigynum in ventral (100) and dorsal (101) views. ‘a’: apophysis on embolus; ‘e’: embolus; ‘p’: pocket; ‘pr’: procursus; ‘r’: putative receptacle; ‘to’: tarsal organ. Scale lines: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEF36E0FFEEFFABFF770BC337C6F8D7" blockId="22.[142,750,1775,1797]" box="[142,750,1775,1797]" pageId="22" pageNumber="283">
|
||
<emphasis id="B924EAF2FFEEFFABFF770BC33431F8D6" box="[142,281,1775,1796]" italics="true" pageId="22" pageNumber="283">Distribution</emphasis>
|
||
. Known only from
|
||
<typeStatus id="54EB8842FFEEFFABFE0E0BDC3700F8D7" box="[503,552,1776,1797]" pageId="22" pageNumber="283">type</typeStatus>
|
||
locality (Map 1).
|
||
</paragraph>
|
||
<paragraph id="8BEF36E0FFEEFFABFF770A3134A8F8A3" blockId="22.[142,758,1821,1905]" pageId="22" pageNumber="283">
|
||
<emphasis id="B924EAF2FFEEFFABFF770A31344EF8E0" box="[142,358,1821,1842]" italics="true" pageId="22" pageNumber="283">Material examined</emphasis>
|
||
.
|
||
<materialsCitation id="3B383CBDFFEEFFABFE7F0A313454F8A3" country="Madagascar" location="English Camp" pageId="22" pageNumber="283" specimenCount="2" specimenCount-female="2" stateProvince="Antsiranana">
|
||
<collectingCountry id="F3477670FFEEFFABFE7F0A313769F8E0" box="[390,577,1821,1842]" name="Madagascar" pageId="22" pageNumber="283">MADAGASCAR</collectingCountry>
|
||
:
|
||
<emphasis id="B924EAF2FFEEFFABFDA60A3237DDF8E1" box="[607,757,1822,1843]" italics="true" pageId="22" pageNumber="283">
|
||
<collectingRegion id="4994F802FFEEFFABFDA60A3237D8F8E1" box="[607,752,1822,1843]" country="Madagascar" name="Antsiranana" pageId="22" pageNumber="283">Antsiranana</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
<location id="8E8F603BFFEEFFABFF770A10341BF883" LSID="urn:lsid:plazi:treatment:03F987F6FFEBFFABFF330B8A34A8F8A3:8E8F603BFFEEFFABFF770A10341BF883" box="[142,307,1852,1874]" country="Madagascar" name="English Camp" pageId="22" pageNumber="283" stateProvince="Antsiranana">English Camp</location>
|
||
:
|
||
<typeStatus id="54EB8842FFEEFFABFEC40A103446F883" box="[317,366,1852,1873]" pageId="22" pageNumber="283">type</typeStatus>
|
||
above, together with
|
||
<specimenCount id="9D56FD69FFEEFFABFD980A1037ADF883" box="[609,645,1851,1873]" pageId="22" pageNumber="283" type="female">2♀</specimenCount>
|
||
and some juveniles, same data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |