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<document id="73B2B5C42F67FF6EDAA112C506E2DA8D" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFB3F778033716784F24CA55" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada regia Köhler 2010, n.sp." docType="treatment" docUuid="9D681E48-6B8B-4F2C-9914-8FC5EC4E5BF8" docUuidSource="ZooBank" docVersion="3" lastPageNumber="253" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="251" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CCA481F6489BDD34CE1F9F0470907FBC">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="D1EFF91C59AA37142F5C290AA3A4725B" type="personal">
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<mods:namePart id="10B4E1C444B053289929DA2F75BB87D8">Köhler, Frank</mods:namePart>
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<mods:typeOfResource id="1B1424E4DB62CC28109AD3444F7A2D1D">text</mods:typeOfResource>
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<mods:title id="5A62BD216062606B20AC39EE9302B279">Records of the Australian Museum</mods:title>
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<mods:part id="F3E6E900050000D89438EF4EC235EE60">
<mods:date id="3C7553CB1711C7877B301834C90FAD1D">2010</mods:date>
<mods:detail id="9882E0A269E4090D7EA0149CFCB89C85" type="pubDate">
<mods:number id="0515BD45DC60EBE92223BD69F052B89B">2010-11-24</mods:number>
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<mods:number id="5068C7AFF52935C4DF3DD8222943E3EA">62</mods:number>
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<mods:number id="3929B95925C39D442CE7056217002B43">2</mods:number>
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<mods:url id="482C548E43F5823B36B5B47A9E9CD480">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<mods:classification id="A0E6189F881F211DBC5AB090231A65BE">journal article</mods:classification>
<mods:identifier id="60F48055C8A48BC993F504DE67BB817C" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="30F366315CC533125E28E92FF4253C9C" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="A8E276945720EB38607EF7441A7BC3D9" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="82423D5C73759683F14D9FE6F74A3BB1" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FFB3F778033716784F24CA55" LSID="urn:lsid:zoobank.org:act:9D681E48-6B8B-4F2C-9914-8FC5EC4E5BF8" httpUri="http://treatment.plazi.org/id/03F987C5FFB3F778033716784F24CA55" lastPageId="36" lastPageNumber="253" pageId="34" pageNumber="251">
<subSubSection id="C34A6558FFB3F77E033716784F6BC823" box="[322,638,1597,1623]" pageId="34" pageNumber="251" type="nomenclature">
<paragraph id="8BEF36D3FFB3F77E033716784F6BC823" blockId="34.[322,638,1597,1623]" box="[322,638,1597,1623]" pageId="34" pageNumber="251">
<heading id="D0A781BFFFB3F77E033716784F6BC823" box="[322,638,1597,1623]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="251" reason="2">
<emphasis id="B924EAC1FFB3F77E033716784F6BC823" bold="true" box="[322,638,1597,1623]" pageId="34" pageNumber="251">
<taxonomicName id="4C504D50FFB3F77E033716784F2FC823" ID-CoL="D7RX" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[322,570,1597,1623]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="regia" status="sp. nov.">
<emphasis id="B924EAC1FFB3F77E033716784F2FC823" bold="true" box="[322,570,1597,1623]" italics="true" pageId="34" pageNumber="251">Amplirhagada regia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB3F77E003716784F6BC823" box="[578,638,1597,1623]" pageId="34" pageNumber="251" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6558FFB3F77E02D3163F4C82C99D" pageId="34" pageNumber="251" type="materials_examined">
<paragraph id="8BEF36D3FFB3F77E02D3163F4C6FC972" blockId="34.[166,794,1658,1798]" pageId="34" pageNumber="251">
<materialsCitation id="3B383C8EFFB3F77E02D3163F4C63C972" collectingDate="2007-08-09" collectorName="Dry &amp; M. Shea" county="Kimberley" latitude="-15.070833" location="Boongaree Island" longLatPrecision="21" longitude="125.187225" municipality="Bonaparte Archipelago" pageId="34" pageNumber="251" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFB3F77E02D3163F4C50C8E6" bold="true" box="[166,325,1658,1682]" pageId="34" pageNumber="251">
<typeStatus id="54EB8871FFB3F77E02D3163F4DF7C8E6" box="[166,226,1658,1682]" pageId="34" pageNumber="251">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFB3F77E032C163F4CB4C8E5" box="[345,417,1658,1681]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="34" pageNumber="251">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFB3F77E03C9163F4F98C8E5" box="[444,653,1658,1681]" country="Australia" name="Western Australia" pageId="34" pageNumber="251">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFB3F77E00E9163F4E03C8E5" box="[668,790,1658,1681]" pageId="34" pageNumber="251">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFB3F77E02D316D24C8EC8DA" box="[166,411,1687,1710]" pageId="34" pageNumber="251">Bonaparte Archipelago</collectingMunicipality>
, central section of
<location id="8E8F6008FFB3F77E002916D24E03C8DA" LSID="urn:lsid:plazi:treatment:03F987C5FFB3F778033716784F24CA55:8E8F6008FFB3F77E002916D24E03C8DA" box="[604,790,1687,1710]" county="Kimberley" latitude="-15.070833" longLatPrecision="21" longitude="125.187225" municipality="Bonaparte Archipelago" name="Boongaree Island" pageId="34" pageNumber="251" stateProvince="Western Australia">Boongaree Island</location>
,
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<geoCoordinate id="EE645014FFB3F77E035716F04CBEC8B8" box="[290,427,1717,1740]" degrees="125" direction="east" minutes="11" orientation="longitude" pageId="34" pageNumber="251" precision="15" seconds="14" value="125.187225">125°11'14&quot;E</geoCoordinate>
; KIS 1-30.
<collectorName id="26A55305FFB3F77E006A16F04F5DC8B8" box="[543,584,1717,1740]" pageId="34" pageNumber="251">Dry</collectorName>
vine thicket on west facing sandstone scree below escarpment, under rocks (leg.
<collectorName id="26A55305FFB3F77E02D316AA4C14C972" box="[166,257,1775,1798]" pageId="34" pageNumber="251">M. Shea</collectorName>
,
<date id="FFEE1013FFB3F77E037916AA4C7BC972" box="[268,366,1775,1798]" pageId="34" pageNumber="251" value="2007-08-09">
<collectingDate id="EFAAE9FBFFB3F77E037916AA4C7BC972" box="[268,366,1775,1798]" pageId="34" pageNumber="251" value="2007-08-09">9.8.2007</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFB3F77E02D3176D4F3FC9EF" blockId="34.[166,794,1832,1947]" pageId="34" pageNumber="251">
<emphasis id="B924EAC1FFB3F77E02D3176D4C3AC94F" bold="true" box="[166,303,1832,1851]" pageId="34" pageNumber="251">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFB3F77E034B176D4F6AC94F" accessionNumber="S34615" box="[318,639,1832,1851]" collectionCode="WAM" pageId="34" pageNumber="251" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FFB3F77E034B176D4C87C94F" box="[318,402,1832,1851]" pageId="34" pageNumber="251" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFB3F77E03EF176D4CC5C94F" box="[410,464,1832,1851]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E03AD176D4F08C94F" box="[472,541,1832,1851]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34615" pageId="34" pageNumber="251" type="EnaNcbi">S34615</accessionNumber>
(Pl. 1.21)
</materialsCitation>
.
<materialsCitation id="3B383C8EFFB3F77E00FE176D4C09C9F7" accessionNumber="S36647, S36449" collectionCode="WAM" latitude="-15.066667" longLatPrecision="21" longitude="125.186386" pageId="34" pageNumber="251" specimenCount="3" typeStatus="paratype">
<typeStatus id="54EB8871FFB3F77E00FE176D4FF7C94F" box="[651,738,1832,1851]" pageId="34" pageNumber="251" type="paratype">Paratypes</typeStatus>
AMS C463705 (4 preserved specimens, as
<typeStatus id="54EB8871FFB3F77E03AE17054F3CC927" box="[475,553,1856,1875]" pageId="34" pageNumber="251" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FFB3F77E004017054F7CC927" box="[565,617,1856,1875]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E001817054FBBC927" box="[621,686,1856,1875]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36647" pageId="34" pageNumber="251" type="EnaNcbi">S36647</accessionNumber>
(3 preserved specimens, as
<typeStatus id="54EB8871FFB3F77E0369171D4C7CC91F" box="[284,361,1880,1899]" pageId="34" pageNumber="251" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FFB3F77E0301171D4CB2C91F" box="[372,423,1880,1899]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E03D9171D4CF9C91F" box="[428,492,1880,1899]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36449" pageId="34" pageNumber="251" type="EnaNcbi">S36449</accessionNumber>
(6 preserved specimens,
<geoCoordinate id="EE645014FFB3F77E00CF171D4E0CC91F" box="[698,793,1880,1899]" degrees="15" direction="south" minutes="04" orientation="latitude" pageId="34" pageNumber="251" precision="15" seconds="00" value="-15.066667">15°04'00&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFB3F77E02D317354C00C9F7" box="[166,277,1904,1923]" degrees="125" direction="east" minutes="11" orientation="longitude" pageId="34" pageNumber="251" precision="15" seconds="11" value="125.186386">125°11'11&quot;E</geoCoordinate>
)
</materialsCitation>
, AMS C463704 (6 shells,
<geoCoordinate id="EE645014FFB3F77E007517354F74C9F7" box="[512,609,1904,1923]" degrees="15" direction="south" minutes="04" orientation="latitude" pageId="34" pageNumber="251" precision="15" seconds="31" value="-15.075277">15°04'31&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFB3F77E001217354FC3C9F7" box="[615,726,1904,1923]" degrees="125" direction="east" minutes="11" orientation="longitude" pageId="34" pageNumber="251" precision="15" seconds="07" value="125.18528">125°11'07&quot;E</geoCoordinate>
),
<materialsCitation id="3B383C8EFFB3F77E009317354F32C9EF" accessionNumber="S36480" collectionCode="WAM" latitude="-15.075277" longLatPrecision="21" longitude="125.18528" pageId="34" pageNumber="251" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED41AE16FFB3F77E009317354E0FC9F7" box="[742,794,1904,1923]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E02D317CD4DF2C9EF" box="[166,231,1928,1947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36480" pageId="34" pageNumber="251" type="EnaNcbi">S36480</accessionNumber>
(12 shells,
<geoCoordinate id="EE645014FFB3F77E033F17CD4CBEC9EF" box="[330,427,1928,1947]" degrees="15" direction="south" minutes="04" orientation="latitude" pageId="34" pageNumber="251" precision="15" seconds="31" value="-15.075277">15°04'31&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFB3F77E03C517CD4F35C9EF" box="[432,544,1928,1947]" degrees="125" direction="east" minutes="11" orientation="longitude" pageId="34" pageNumber="251" precision="15" seconds="07" value="125.18528">125°11'07&quot;E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<caption id="DF2F665BFFB3F77E013217E54941C99F" ID-DOI="http://doi.org/10.5281/zenodo.5239191" ID-Zenodo-Dep="5239191" httpUri="https://zenodo.org/record/5239191/files/figure.png" pageId="34" pageNumber="251" startId="34.[839,902,1952,1973]" targetBox="[853,1462,788,1941]" targetPageId="34">
<paragraph id="8BEF36D3FFB3F77E013217E54941C99F" blockId="34.[839,1468,1952,2027]" pageId="34" pageNumber="251">
Figure 52. Interior of penial chamber of
<taxonomicName id="4C504D50FFB3F77E06B717E54896C9C1" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1218,1411,1952,1973]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="regia" status="sp. nov.">
<emphasis id="B924EAC1FFB3F77E06B717E54896C9C1" box="[1218,1411,1952,1973]" italics="true" pageId="34" pageNumber="251">Amplirhagada regia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB3F77E07F217E548A2C9C1" box="[1415,1463,1952,1973]" pageId="34" pageNumber="251" rank="species">n.sp.</taxonomicNameLabel>
, paratype WAM
<accessionNumber id="9403AB30FFB3F77E01AE17FE4931C9A4" box="[987,1060,1979,2000]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36647" pageId="34" pageNumber="251" type="EnaNcbi">S36647</accessionNumber>
(9 August; scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FFB3F77E02D317F84C82C99D" blockId="34.[166,794,1981,2025]" pageId="34" pageNumber="251">
<emphasis id="B924EAC1FFB3F77E02D317F84CA2C9A4" bold="true" box="[166,439,1981,2000]" pageId="34" pageNumber="251">Additional, non-type material</emphasis>
.
<collectionCode id="ED41AE16FFB3F77E03B617FB4CE2C9A5" box="[451,503,1982,2001]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E038817FB4F54C9A5" box="[509,577,1982,2001]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36479" pageId="34" pageNumber="251" type="EnaNcbi">S36479</accessionNumber>
,
<collectionCode id="ED41AE16FFB3F77E003F17FB4F6BC9A5" box="[586,638,1982,2001]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E00F117FB4FD2C9A5" box="[644,711,1982,2001]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36481" pageId="34" pageNumber="251" type="EnaNcbi">S36481</accessionNumber>
5,
<collectionCode id="ED41AE16FFB3F77E009317FB4E0FC9A5" box="[742,794,1982,2001]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="34" pageNumber="251" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB3F77E02D317934DFDC99D" box="[166,232,2006,2025]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36648" pageId="34" pageNumber="251" type="EnaNcbi">S36648</accessionNumber>
(Boongaree Island).
</paragraph>
</subSubSection>
<caption id="DF2F665BFFB2F77F020C15884C87C838" ID-DOI="http://doi.org/10.5281/zenodo.5239193" ID-Zenodo-Dep="5239193" httpUri="https://zenodo.org/record/5239193/files/figure.png" pageId="35" pageNumber="252" startId="35.[121,183,1485,1506]" targetBox="[121,1421,151,1471]" targetPageId="35">
<paragraph id="8BEF36D3FFB2F77F020C15884C87C838" blockId="35.[121,1422,1485,1612]" pageId="35" pageNumber="252">
Figure 53. SEM photographs of
<taxonomicName id="4C504D50FFB2F77F03D015884F76CB96" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[421,611,1485,1506]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="252" phylum="Mollusca" rank="species" species="regia" status="sp. nov.">
<emphasis id="B924EAC1FFB2F77F03D015884F76CB96" box="[421,611,1485,1506]" italics="true" pageId="35" pageNumber="252">Amplirhagada regia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB2F77F001215884F80CB96" box="[615,661,1485,1506]" pageId="35" pageNumber="252" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FFB2F77F00D515884FD9CB96" box="[672,716,1485,1506]" italics="true" pageId="35" pageNumber="252">AD</emphasis>
) Shell, paratype AMS C463704: (
<emphasis id="B924EAC1FFB2F77F067F1588490CCB96" box="[1034,1049,1485,1506]" italics="true" pageId="35" pageNumber="252">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFB2F77F02AA15A24DFBCB88" box="[223,238,1511,1532]" italics="true" pageId="35" pageNumber="252">B</emphasis>
) sculpture on second to forth whorl viewed from above (scale 1 mm); (
<emphasis id="B924EAC1FFB2F77F01D515A24EA5CB88" box="[928,944,1511,1532]" italics="true" pageId="35" pageNumber="252">C</emphasis>
) surface of last whorl (scale 1 mm); (
<emphasis id="B924EAC1FFB2F77F076915A24838CB88" box="[1308,1325,1511,1532]" italics="true" pageId="35" pageNumber="252">D</emphasis>
) close-up showing axial growth lines on last whorl (Scale 100 µm). (
<emphasis id="B924EAC1FFB2F77F00D616474FA7C863" box="[675,690,1538,1559]" italics="true" pageId="35" pageNumber="252">E</emphasis>
) Jaw, paratype WAM
<accessionNumber id="9403AB30FFB2F77F01F116474EDBC863" box="[900,974,1538,1559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36647" pageId="35" pageNumber="252" type="EnaNcbi">S36647</accessionNumber>
(scale 100 µm). (
<emphasis id="B924EAC1FFB2F77F06061647498AC863" box="[1139,1183,1538,1559]" italics="true" pageId="35" pageNumber="252">EG</emphasis>
) Radula, paratype WAM
<accessionNumber id="9403AB30FFB2F77F020C16594DD3C845" box="[121,198,1564,1585]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36647" pageId="35" pageNumber="252" type="EnaNcbi">S36647</accessionNumber>
: (
<emphasis id="B924EAC1FFB2F77F02A216584DF3C846" box="[215,230,1565,1586]" italics="true" pageId="35" pageNumber="252">E</emphasis>
) close-up of lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FFB2F77F000116584F96C846" box="[628,643,1565,1586]" italics="true" pageId="35" pageNumber="252">F</emphasis>
) close-up of central and inner lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FFB2F77F06CC165849DFC846" box="[1209,1226,1565,1586]" italics="true" pageId="35" pageNumber="252">G</emphasis>
) close-up of middle marginal teeth (scale 10 µm).
</paragraph>
</caption>
<subSubSection id="C34A6558FFB2F778020C16334F24CA55" lastPageId="36" lastPageNumber="253" pageId="35" pageNumber="252" type="etymology">
<paragraph id="8BEF36D3FFB2F77F020C16334F75C8DE" blockId="35.[121,748,1654,1706]" pageId="35" pageNumber="252">
<emphasis id="B924EAC1FFB2F77F020C16334DE6C8FA" bold="true" box="[121,243,1654,1678]" pageId="35" pageNumber="252">Etymology</emphasis>
. From
<taxonomicName id="4C504D50FFB2F77F034816334C66C8F9" authorityName="Köhler" authorityYear="2010" box="[317,371,1654,1677]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="252" phylum="Mollusca" rank="species" species="regia">
<emphasis id="B924EAC1FFB2F77F034816334C66C8F9" box="[317,371,1654,1677]" italics="true" pageId="35" pageNumber="252">regia</emphasis>
</taxonomicName>
(Latin = royal), referring to fact that this species is among the largest of the genus.
</paragraph>
<paragraph id="8BEF36D3FFB2F77F020C168B4CBFC891" blockId="35.[121,426,1742,1765]" box="[121,426,1742,1765]" pageId="35" pageNumber="252">Sealing strategy. Free sealer.</paragraph>
<paragraph id="8BEF36D3FFB2F77F020C174D485AC976" blockId="35.[121,749,1800,2000]" lastBlockId="35.[794,1421,1654,1794]" pageId="35" pageNumber="252">
<emphasis id="B924EAC1FFB2F77F020C174D4DA4C954" bold="true" box="[121,177,1800,1824]" pageId="35" pageNumber="252">Shell</emphasis>
(
<figureCitation id="136B2A56FFB2F77F02CA174C4C54C954" box="[191,321,1801,1824]" captionStart="Figure 53" captionStartId="35.[121,183,1485,1506]" captionTargetBox="[121,1421,151,1471]" captionTargetId="figure-9@35.[121,1421,151,1471]" captionTargetPageId="35" captionText="Figure 53. SEM photographs of Amplirhagada regia n.sp.(AD) Shell, paratypeAMS C463704: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to forth whorl viewed from above (scale 1 mm); (C) surface of last whorl (scale 1 mm); (D) close-up showing axial growth lines on last whorl (Scale 100 µm). (E) Jaw, paratype WAM S36647 (scale 100 µm). (EG) Radula, paratype WAM S36647: (E) close-up of lateral teeth (scale 10 µm); (F) close-up of central and inner lateral teeth (scale 10 µm); (G) close-up of middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239193" httpUri="https://zenodo.org/record/5239193/files/figure.png" pageId="35" pageNumber="252">Fig. 53AD</figureCitation>
, Pl. 1.21). Globose with medium high to high spire; solid to thick. Periphery angulated; upper sector of whorls rather flattened, basal sector rounded. Umbilicus forming chink or narrowly winding opening, 60100 percent concealed by columellar reflection. Background colour uniform, creamish horn; outer lip purple; inner lip pale purple. Protoconch c.
<quantity id="4CA89B36FFB2F77F031D17FC4CAFC9A4" box="[360,442,1977,2000]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="35" pageNumber="252" unit="mm" value="2.2">2.2 mm</quantity>
in diameter, comprising 1.5 whorls, sculpture almost smooth. Teleoconch smooth except for faint axial growth lines. Angle of aperture 45; outer lip simple rounded; moderate; slightly expanded; not reflected, or slightly reflected; basal node of lip present; weak; palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<paragraph id="8BEF36D3FFB2F77F016F17604854C90C" blockId="35.[794,1421,1829,1912]" pageId="35" pageNumber="252">
<emphasis id="B924EAC1FFB2F77F016F17604EE9C949" bold="true" box="[794,1020,1829,1853]" pageId="35" pageNumber="252">Pallial morphology</emphasis>
. Pallial cavity deep, extending one whorl. Pigmentation on mantle comprising brownish grey spots. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFB2F778016F17DE4F52CCF3" blockId="35.[794,1421,1947,2000]" lastBlockId="36.[166,795,154,647]" lastPageId="36" lastPageNumber="253" pageId="35" pageNumber="252">
<emphasis id="B924EAC1FFB2F77F016F17DE4EEBC9C7" bold="true" box="[794,1022,1947,1971]" pageId="35" pageNumber="252">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFB2F77F067E17DE4998C9C6" box="[1035,1165,1947,1970]" captionStart-0="Figure 51" captionStart-1="Figure 52" captionStartId-0="34.[839,902,634,655]" captionStartId-1="34.[839,902,1952,1973]" captionTargetBox-1="[853,1462,788,1941]" captionTargetId-1="figure-337@34.[839,1466,775,1942]" captionTargetPageId-1="34" captionText-0="Figure 51. Genitalia of Amplirhagada regia n.sp., paratype WAM S36647 (9August; scale 10 mm).Compare with Fig. 3 for labelling of structures." captionText-1="Figure 52. Interior of penial chamber of Amplirhagada regia n.sp., paratype WAM S36647 (9August; scale 5 mm).Compare with Fig. 4 for labelling of structures." figureDoi-1="http://doi.org/10.5281/zenodo.5239191" httpUri-1="https://zenodo.org/record/5239191/files/figure.png" pageId="35" pageNumber="252">Figs. 5152</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than or equal to penis length. Penial sheath evenly thin. Penial verge short (&lt;
<emphasis id="B924EAC1FFB5F778038B10904F13CE97" bold="true" box="[510,518,213,227]" pageId="36" pageNumber="253">
<superScript id="7C259B9BFFB5F778038B10904F13CE97" attach="none" box="[510,518,213,227]" fontSize="6" pageId="36" pageNumber="253">1</superScript>
</emphasis>
<emphasis id="B924EAC1FFB5F778007F10994F07CE9E" bold="true" box="[522,530,220,234]" pageId="36" pageNumber="253">
<subScript id="17D43496FFB5F778007F10994F07CE9E" attach="left" box="[522,530,220,234]" fontSize="6" pageId="36" pageNumber="253">8</subScript>
</emphasis>
penial chamber), slender to spatulate, with pointed tip. Penial wall pustules absent. Main stimulatory pilaster absent. Two longitudinal pilasters present at basal portion of penial chamber, two additional weakly developed pilasters visible at apical portion. Vas deferens entering penial sheath in upper third. Vagina short to moderately long, tubular. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately wide, internally with smooth longitudinal pilasters. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue, internally entirely smooth, with delicate wall. Free oviduct comprising about half of anterior part of oviduct, more or less straight. Spermoviduct of same length as anterior part of oviduct. Talon embedded in albumen gland close to anterior end of albumen gland.
</paragraph>
<paragraph id="8BEF36D3FFB5F77802D312EF4F8CCDD8" blockId="36.[166,794,682,940]" pageId="36" pageNumber="253">
<emphasis id="B924EAC1FFB5F77802D312EF4C89CCB6" bold="true" box="[166,412,682,706]" pageId="36" pageNumber="253">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFB5F77803DB12EF4F21CCB5" box="[430,564,682,705]" captionStart="Figure 53" captionStartId="35.[121,183,1485,1506]" captionTargetBox="[121,1421,151,1471]" captionTargetId="figure-9@35.[121,1421,151,1471]" captionTargetPageId="35" captionText="Figure 53. SEM photographs of Amplirhagada regia n.sp.(AD) Shell, paratypeAMS C463704: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to forth whorl viewed from above (scale 1 mm); (C) surface of last whorl (scale 1 mm); (D) close-up showing axial growth lines on last whorl (Scale 100 µm). (E) Jaw, paratype WAM S36647 (scale 100 µm). (EG) Radula, paratype WAM S36647: (E) close-up of lateral teeth (scale 10 µm); (F) close-up of central and inner lateral teeth (scale 10 µm); (G) close-up of middle marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239193" httpUri="https://zenodo.org/record/5239193/files/figure.png" pageId="36" pageNumber="253">Fig. 53FH</figureCitation>
). Rectangular. Tooth formula C+20+3+20. In average with 147±2.5 rows of teeth, 29.6±1.7 rows per mm (n = 2). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of tooth; ectocones and endocones vestigial. Marginal teeth with elongate mesocones; ectocones shorter and narrower than mesocones; endocones greatly reduced in size.
</paragraph>
<paragraph id="8BEF36D3FFB5F77802D3138A4F24CA55" blockId="36.[166,793,975,1057]" pageId="36" pageNumber="253">
<emphasis id="B924EAC1FFB5F77802D3138A4CB5CD93" bold="true" box="[166,416,975,999]" pageId="36" pageNumber="253">Comparative remarks</emphasis>
. Diagnostic features are the broadly conical to dome-shaped shell, smooth inner penial wall with lack of pustulation and main pilaster.
</paragraph>
</subSubSection>
</treatment>
</document>