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<document id="71489118DCD6F22D6B038129006DBFB6" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFB1F77E01B010DE4F07CB85" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada tricenaria Köhler 2010, n.sp." docType="treatment" docUuid="33C642CD-AF40-4B3C-B77F-58B905C2563D" docUuidSource="ZooBank" docVersion="3" lastPageNumber="251" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="249" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="996AB77C60A37E73D72795218E218B50">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="B318B2C11B9549AC302232ECC9E010EC">Köhler, Frank</mods:namePart>
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<mods:title id="7E23762E1B7D9FD2B2A596A794629049">Records of the Australian Museum</mods:title>
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<mods:date id="1FA04B5291543B9E135C34F055B601FD">2010</mods:date>
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<mods:number id="DA2866E1573C44C41A1FBA1A31C39BAC">2010-11-24</mods:number>
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<mods:classification id="428D3F598DFF12DC9B0464A37702C0DC">journal article</mods:classification>
<mods:identifier id="BF7A846E40B1107B78A8C4404A2E9E98" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
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<mods:identifier id="102058D668608BF107F3503AECAF8510" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
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<treatment id="03F987C5FFB1F77E01B010DE4F07CB85" ID-DOI="http://doi.org/10.5281/zenodo.5239039" ID-Zenodo-Dep="5239039" LSID="urn:lsid:zoobank.org:act:33C642CD-AF40-4B3C-B77F-58B905C2563D" httpUri="http://treatment.plazi.org/id/03F987C5FFB1F77E01B010DE4F07CB85" lastPageId="34" lastPageNumber="251" pageId="32" pageNumber="249">
<subSubSection id="C34A6558FFB1F77C01B010DE4829CEC0" box="[965,1340,154,181]" pageId="32" pageNumber="249" type="nomenclature">
<paragraph id="8BEF36D3FFB1F77C01B010DE4829CEC0" blockId="32.[965,1340,154,181]" box="[965,1340,154,181]" pageId="32" pageNumber="249">
<heading id="D0A781BFFFB1F77C01B010DE4829CEC0" box="[965,1340,154,181]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="249" reason="2">
<emphasis id="B924EAC1FFB1F77C01B010DE4829CEC0" bold="true" box="[965,1340,154,181]" pageId="32" pageNumber="249">
<taxonomicName id="4C504D50FFB1F77C01B010DE49ECCEC1" ID-CoL="D7S6" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[965,1273,155,181]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="tricenaria" status="sp. nov.">
<emphasis id="B924EAC1FFB1F77C01B010DE49ECCEC1" bold="true" box="[965,1273,155,181]" italics="true" pageId="32" pageNumber="249">Amplirhagada tricenaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB1F77C077510DF4829CEC0" box="[1280,1340,154,180]" pageId="32" pageNumber="249" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6558FFB1F77C013210924805CFDF" pageId="32" pageNumber="249" type="materials_examined">
<paragraph id="8BEF36D3FFB1F77C013210924833CF32" blockId="32.[839,1467,215,326]" pageId="32" pageNumber="249">
<materialsCitation id="3B383C8EFFB1F77C013210924837CF32" collectingDate="1988-07-22" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="Kimberley" latitude="-15.031667" location="Prince Frederick Harbour" longLatPrecision="21" longitude="125.38694" municipality="Bonaparte Archipelago" pageId="32" pageNumber="249" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFB1F77C013210924EF3CE9B" bold="true" box="[839,998,215,239]" pageId="32" pageNumber="249">
<typeStatus id="54EB8871FFB1F77C013210924E96CE9B" box="[839,899,215,239]" pageId="32" pageNumber="249">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFB1F77C018F10924954CE9A" box="[1018,1089,215,238]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="32" pageNumber="249">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFB1F77C06281092483BCE9A" box="[1117,1326,215,238]" country="Australia" name="Western Australia" pageId="32" pageNumber="249">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFB1F77C0748109248A2CE9A" box="[1341,1463,215,238]" pageId="32" pageNumber="249">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFB1F77C013210B14955CF7F" box="[839,1088,244,267]" pageId="32" pageNumber="249">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FFB1F77C063F10B14849CF7F" LSID="urn:lsid:plazi:treatment:03F987C5FFB1F77E01B010DE4F07CB85:8E8F6008FFB1F77C063F10B14849CF7F" box="[1098,1372,244,267]" county="Kimberley" latitude="-15.031667" longLatPrecision="21" longitude="125.38694" municipality="Bonaparte Archipelago" name="Prince Frederick Harbour" pageId="32" pageNumber="249" stateProvince="Western Australia">Prince Frederick Harbour</location>
, north of
<location id="8E8F6008FFB1F77C013211574EDACF5D" LSID="urn:lsid:plazi:treatment:03F987C5FFB1F77E01B010DE4F07CB85:8E8F6008FFB1F77C013211574EDACF5D" box="[839,975,274,297]" county="Kimberley" latitude="-15.031667" longLatPrecision="21" longitude="125.38694" municipality="Bonaparte Archipelago" name="Hunter River" pageId="32" pageNumber="249" stateProvince="Western Australia">Hunter River</location>
mouth,
<geoCoordinate id="EE645014FFB1F77C06571157498FCF5D" box="[1058,1178,274,297]" degrees="15" direction="south" minutes="01" orientation="latitude" pageId="32" pageNumber="249" precision="15" seconds="54" value="-15.031667">15°01'54&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFB1F77C06EA1157483CCF5D" box="[1183,1321,274,297]" degrees="125" direction="east" minutes="23" orientation="longitude" pageId="32" pageNumber="249" precision="15" seconds="13" value="125.38694">125°23'13&quot;E</geoCoordinate>
; KC-080 (leg.
<collectionCode id="ED41AE16FFB1F77C0132116A4E4FCF32" box="[839,858,303,326]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="32" pageNumber="249" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FFB1F77C0111116A4EACCF32" box="[868,953,303,326]" pageId="32" pageNumber="249">Kessner</collectorName>
&amp;
<collectorName id="26A55305FFB1F77C01AF116A4997CF32" box="[986,1154,303,326]" pageId="32" pageNumber="249">A. Longbottom</collectorName>
,
<date id="FFEE1013FFB1F77C06F9116A480CCF32" box="[1164,1305,303,326]" pageId="32" pageNumber="249" value="1988-07-22">
<collectingDate id="EFAAE9FBFFB1F77C06F9116A480CCF32" box="[1164,1305,303,326]" pageId="32" pageNumber="249" value="1988-07-22">22 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFB1F77C0132112D4805CFDF" blockId="32.[839,1467,360,427]" pageId="32" pageNumber="249">
<emphasis id="B924EAC1FFB1F77C0132112D4EC5CF0F" bold="true" box="[839,976,360,379]" pageId="32" pageNumber="249">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFB1F77C01AA112D4834CF0F" accessionNumber="S34614" box="[991,1313,360,379]" collectionCode="WAM" pageId="32" pageNumber="249" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FFB1F77C01AA112D4926CF0F" box="[991,1075,360,379]" pageId="32" pageNumber="249" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFB1F77C064E112D4964CF0F" box="[1083,1137,360,379]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="32" pageNumber="249" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB1F77C060C112D49A8CF0F" box="[1145,1213,360,379]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34614" pageId="32" pageNumber="249" type="EnaNcbi">S34614</accessionNumber>
(Pl. 1.20)
</materialsCitation>
.
<materialsCitation id="3B383C8EFFB1F77C0759112D4819CFDF" accessionNumber="S41476" collectionCode="WAM" pageId="32" pageNumber="249" specimenCode="FMNH 219322" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EB8871FFB1F77C0759112D4896CF0F" box="[1324,1411,360,379]" pageId="32" pageNumber="249" type="paratype">Paratypes</typeStatus>
AMS C463757 (2 preserved specimens),
<specimenCode id="DBF69EA8FFB1F77C06D311C5482DCFE7" box="[1190,1336,384,403]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="32" pageNumber="249" type="Museum">FMNH 219322</specimenCode>
(6 preserved specimens),
<collectionCode id="ED41AE16FFB1F77C01C111DD4EFCCFDF" box="[948,1001,408,427]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="32" pageNumber="249" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFB1F77C019B11DD4925CFDF" box="[1006,1072,408,427]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41476" pageId="32" pageNumber="249" type="EnaNcbi">S41476</accessionNumber>
(4 preserved specimens)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFB1F77E0132118A4F07CB85" lastPageId="34" lastPageNumber="251" pageId="32" pageNumber="249" type="etymology">
<paragraph id="8BEF36D3FFB1F77C0132118A4979CC55" blockId="32.[839,1466,463,545]" pageId="32" pageNumber="249">
<emphasis id="B924EAC1FFB1F77C0132118A4ED7CF93" bold="true" box="[839,962,463,487]" pageId="32" pageNumber="249">Etymology</emphasis>
. From
<taxonomicName id="4C504D50FFB1F77C0661118A496ACF92" authorityName="Köhler" authorityYear="2010" box="[1044,1151,463,486]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="tricenaria">
<emphasis id="B924EAC1FFB1F77C0661118A496ACF92" box="[1044,1151,463,486]" italics="true" pageId="32" pageNumber="249">tricenaria</emphasis>
</taxonomicName>
(Latin = thirty), in reference to manuscript name assigned to this species by Solem (“
<taxonomicName id="4C504D50FFB1F77C0129124F4EE3CC55" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[860,1014,522,545]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B924EAC1FFB1F77C0129124F4EE3CC55" box="[860,1014,522,545]" italics="true" pageId="32" pageNumber="249">Amplirhagada</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB1F77C0189124F493ACC55" box="[1020,1071,522,545]" pageId="32" pageNumber="249" rank="species">n.sp.</taxonomicNameLabel>
30”).
</paragraph>
<paragraph id="8BEF36D3FFB1F77D013212014C83C883" blockId="32.[839,1467,580,955]" lastBlockId="33.[121,747,1731,1783]" lastPageId="33" lastPageNumber="250" pageId="32" pageNumber="249">
<emphasis id="B924EAC1FFB1F77C013212014E6ACC28" bold="true" box="[839,895,580,604]" pageId="32" pageNumber="249">Shell</emphasis>
(
<figureCitation id="136B2A56FFB1F77C01FA12014906CC2F" box="[911,1043,580,603]" captionStart="Figure 49" captionStartId="33.[121,184,1555,1576]" captionTargetBox="[121,1421,151,1543]" captionTargetId="figure-273@33.[121,1421,151,1543]" captionTargetPageId="33" captionText="Figure 49. SEM photographs of Amplirhagada tricenaria n.sp. (AD) Shell, paratypeAMS C463757:(A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to third whorl viewed from above (scale 200 µm); (C) suture between last and penultimate whorl with details of sculpture (scale 100 µm); (D) surface of last whorl, viewed rectangularly from above (Scale 200 µm). (E) Jaw, paratype FMNH 219322 (scale 100 µm). (FG) Radula, paratype FMNH 219332: (F) central and lateral teeth (scale 20 µm); (G) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239187" httpUri="https://zenodo.org/record/5239187/files/figure.png" pageId="32" pageNumber="249">Fig. 49AD</figureCitation>
, Pl. 1.20). Semi-globose, with medium high spire; thin to solid (translucent). Periphery slightly angulate; upper and basal sectors of whorls rounded. Umbilicus forming a chink, 90100 percent concealed by columellar reflection. Background colour dark horn to ochre; sub-sutural and mid-whorl bands diffuse to well defined, yellowish brown, thin to moderately broad, most conspicuous on last whorls; ventral colour horn; outer lip colour same as shell; inner lip yellowish white. Protoconch c.
<quantity id="4CA89B36FFB1F77C073A136A488ACD32" box="[1359,1439,815,838]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="32" pageNumber="249" unit="mm" value="2.6">2.6 mm</quantity>
in diameter, comprising 2 whorls, almost smooth. Teleoconch with pronounced axial growth lines, last whorl almost smooth. Angle of aperture 30°, outer lip rounded, sharp to moderately thick, slightly expanded, slightly reflected, basal node of lip absent or weak, palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<caption id="DF2F665BFFB1F77C013017C949BBC9A2" ID-DOI="http://doi.org/10.5281/zenodo.5239185" ID-Zenodo-Dep="5239185" httpUri="https://zenodo.org/record/5239185/files/figure.png" pageId="32" pageNumber="249" startId="32.[837,900,1932,1953]" targetBox="[858,1453,961,1923]" targetPageId="32">
<paragraph id="8BEF36D3FFB1F77C013017C949BBC9A2" blockId="32.[837,1465,1932,2006]" pageId="32" pageNumber="249">
Figure 48. Interior of penial chamber of
<taxonomicName id="4C504D50FFB1F77C06BD17C948ACC9D5" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1224,1465,1932,1953]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="tricenaria" status="sp. nov.">
<emphasis id="B924EAC1FFB1F77C06BD17C948ACC9D5" box="[1224,1465,1932,1953]" italics="true" pageId="32" pageNumber="249">Amplirhagada tricenaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB1F77C013017E34E60C9CF" box="[837,885,1958,1979]" pageId="32" pageNumber="249" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219332 (22 July, scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFB0F77D020C16564CDCC8E6" ID-DOI="http://doi.org/10.5281/zenodo.5239187" ID-Zenodo-Dep="5239187" httpUri="https://zenodo.org/record/5239187/files/figure.png" pageId="33" pageNumber="250" startId="33.[121,184,1555,1576]" targetBox="[121,1421,151,1543]" targetPageId="33">
<paragraph id="8BEF36D3FFB0F77D020C16564CDCC8E6" blockId="33.[121,1422,1555,1682]" pageId="33" pageNumber="250">
Figure 49. SEM photographs of
<taxonomicName id="4C504D50FFB0F77D03DD16564F82C85C" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[424,663,1555,1576]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="250" phylum="Mollusca" rank="species" species="tricenaria" status="sp. nov.">
<emphasis id="B924EAC1FFB0F77D03DD16564F82C85C" box="[424,663,1555,1576]" italics="true" pageId="33" pageNumber="250">Amplirhagada tricenaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB0F77D00EE16564FDCC85C" box="[667,713,1555,1576]" pageId="33" pageNumber="250" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FFB0F77D00A016564E14C85C" box="[725,769,1555,1576]" italics="true" pageId="33" pageNumber="250">AD</emphasis>
) Shell, paratype AMS C463757: (
<emphasis id="B924EAC1FFB0F77D063616564947C85C" box="[1091,1106,1555,1576]" italics="true" pageId="33" pageNumber="250">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFB0F77D036F166B4C3CC837" box="[282,297,1582,1603]" italics="true" pageId="33" pageNumber="250">B</emphasis>
) sculpture on second to third whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FFB0F77D01A8166B4EF8C837" box="[989,1005,1582,1603]" italics="true" pageId="33" pageNumber="250">C</emphasis>
) suture between last and penultimate whorl with details of sculpture (scale 100 µm); (
<emphasis id="B924EAC1FFB0F77D007B160D4F0AC829" box="[526,543,1608,1629]" italics="true" pageId="33" pageNumber="250">D</emphasis>
) surface of last whorl, viewed rectangularly from above (Scale 200 µm). (
<emphasis id="B924EAC1FFB0F77D0698160D49E9C829" box="[1261,1276,1608,1629]" italics="true" pageId="33" pageNumber="250">E</emphasis>
) Jaw, paratype FMNH 219322 (scale 100 µm). (
<emphasis id="B924EAC1FFB0F77D03C616264CCAC80C" box="[435,479,1635,1656]" italics="true" pageId="33" pageNumber="250">FG</emphasis>
) Radula, paratype FMNH 219332: (
<emphasis id="B924EAC1FFB0F77D014916264E5EC80C" box="[828,843,1635,1656]" italics="true" pageId="33" pageNumber="250">F</emphasis>
) central and lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FFB0F77D06A6162649F1C80C" box="[1235,1252,1635,1656]" italics="true" pageId="33" pageNumber="250">G</emphasis>
) outer lateral and inner marginal teeth (scale 20 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FFB0F77D020C174E4C8DC90E" blockId="33.[121,749,1803,1914]" pageId="33" pageNumber="250">
<emphasis id="B924EAC1FFB0F77D020C174E4C4EC957" bold="true" box="[121,347,1803,1827]" pageId="33" pageNumber="250">Pallial morphology</emphasis>
. Pallial cavity deep, extending one whorl. Pigmentation on mantle mottled, greyish or brownish (probably leached in actual specimens). Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFB0F77E020C17D54F3ACD95" blockId="33.[121,749,1936,2018]" lastBlockId="34.[166,795,765,993]" lastPageId="34" lastPageNumber="251" pageId="33" pageNumber="250">
<emphasis id="B924EAC1FFB0F77D020C17D54C48C9DC" bold="true" box="[121,349,1936,1960]" pageId="33" pageNumber="250">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFB0F77D031E17D54CD6C9D3" box="[363,451,1936,1959]" captionStart="Figure 48" captionStartId="32.[837,900,1932,1953]" captionTargetBox="[858,1453,961,1923]" captionTargetId="figure-494@32.[839,1466,949,1928]" captionTargetPageId="32" captionText="Figure 48. Interior of penial chamber of Amplirhagada tricenaria n.sp., paratype FMNH 219332 (22 July, scale 5 mm). Compare with Fig. 4 for labelling of structures." figureDoi="http://doi.org/10.5281/zenodo.5239185" httpUri="https://zenodo.org/record/5239185/files/figure.png" pageId="33" pageNumber="250">Figs. 48</figureCitation>
,
<figureCitation id="136B2A56FFB0F77D03B817D54CFEC9D3" box="[461,491,1936,1959]" captionStart="Figure 50" captionStartId="34.[166,229,634,655]" captionTargetBox="[171,1462,154,613]" captionTargetPageId="34" captionText="Figure 50. Genitalia of Amplirhagada tricenaria n.sp., paratype AMS C463757 (22 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." figureDoi="http://doi.org/10.5281/zenodo.5239189" httpUri="https://zenodo.org/record/5239189/files/figure.png" pageId="33" pageNumber="250">50</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct. Vas deferens forms simple loop before entering penis. Penial retractor muscle shorter than penis. Penial verge very short (barely visible), slender to spatulate with pointed tip. Penial wall pustules small, elongated, arranged in rows on apical to median portion of penial chamber. Base of inner penial walls with many smooth longitudinal pilasters. Main stimulatory pilaster well-differentiated, elongate to cone-shaped, sculptured by smooth ridges, comprising apical to median portion of penial chamber. Vas deferens entering penial sheath in upper third.Vagina of medium length and thickness, tubular, posteriorly slightly inflated. Inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct moderately thick, internally with smooth longitudinal pilasters. Spermathecal head globular to elongately inflated, connected with oviduct by connective tissue, internally smooth, with thin wall. Free oviduct comprising less than half of anterior part of oviduct, more or less straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<caption id="DF2F665BFFB3F77E02D3123F4C6ACCB0" ID-DOI="http://doi.org/10.5281/zenodo.5239189" ID-Zenodo-Dep="5239189" httpUri="https://zenodo.org/record/5239189/files/figure.png" pageId="34" pageNumber="251" startId="34.[166,229,634,655]" targetBox="[171,1462,154,613]" targetPageId="34">
<paragraph id="8BEF36D3FFB3F77E02D3123F4C6ACCB0" blockId="34.[166,794,634,708]" pageId="34" pageNumber="251">
Figure 50. Genitalia of
<taxonomicName id="4C504D50FFB3F77E03E5123F4F96CCFB" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[400,643,634,655]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="tricenaria" status="sp. nov.">
<emphasis id="B924EAC1FFB3F77E03E5123F4F96CCFB" box="[400,643,634,655]" italics="true" pageId="34" pageNumber="251">Amplirhagada tricenaria</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB3F77E00FE123F4FAECCFB" box="[651,699,634,655]" pageId="34" pageNumber="251" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463757 (22 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFB3F77E0132123F4ED3CCB0" pageId="34" pageNumber="251" startId="34.[839,902,634,655]">
<paragraph id="8BEF36D3FFB3F77E0132123F4ED3CCB0" blockId="34.[839,1466,634,708]" pageId="34" pageNumber="251">
Figure 51. Genitalia of
<taxonomicName id="4C504D50FFB3F77E065D123F49FCCCFB" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1064,1257,634,655]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="regia" status="sp. nov.">
<emphasis id="B924EAC1FFB3F77E065D123F49FCCCFB" box="[1064,1257,634,655]" italics="true" pageId="34" pageNumber="251">Amplirhagada regia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB3F77E069A123F480BCCFB" box="[1263,1310,634,655]" pageId="34" pageNumber="251" rank="species">n.sp.</taxonomicNameLabel>
, paratype WAM
<accessionNumber id="9403AB30FFB3F77E013212D04E85CCDE" box="[839,912,661,682]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36647" pageId="34" pageNumber="251" type="EnaNcbi">S36647</accessionNumber>
(9 August; scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FFB3F77E02D314414CD0CB72" blockId="34.[166,794,1028,1286]" pageId="34" pageNumber="251">
<emphasis id="B924EAC1FFB3F77E02D314414C89CA68" bold="true" box="[166,412,1028,1052]" pageId="34" pageNumber="251">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFB3F77E03DB14404F21CA68" box="[430,564,1029,1052]" captionStart="Figure 49" captionStartId="33.[121,184,1555,1576]" captionTargetBox="[121,1421,151,1543]" captionTargetId="figure-273@33.[121,1421,151,1543]" captionTargetPageId="33" captionText="Figure 49. SEM photographs of Amplirhagada tricenaria n.sp. (AD) Shell, paratypeAMS C463757:(A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second to third whorl viewed from above (scale 200 µm); (C) suture between last and penultimate whorl with details of sculpture (scale 100 µm); (D) surface of last whorl, viewed rectangularly from above (Scale 200 µm). (E) Jaw, paratype FMNH 219322 (scale 100 µm). (FG) Radula, paratype FMNH 219332: (F) central and lateral teeth (scale 20 µm); (G) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239187" httpUri="https://zenodo.org/record/5239187/files/figure.png" pageId="34" pageNumber="251">Fig. 49FG</figureCitation>
). Rectangular. Tooth formula C+1517+3+1823. In average with 152±3.0 rows of teeth, 28.1±0.8 rows per mm (n = 2). Central teeth with sharply pointed, ovate mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with bluntly pointed, ovate mesocones, length equal to base of tooth; ectocones small, endocones vestigial. Marginal teeth with triangular mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.
</paragraph>
<paragraph id="8BEF36D3FFB3F77E02D3156F4F07CB85" blockId="34.[166,795,1322,1521]" pageId="34" pageNumber="251">
<emphasis id="B924EAC1FFB3F77E02D3156F4CB7CB36" bold="true" box="[166,418,1322,1346]" pageId="34" pageNumber="251">Comparative remarks</emphasis>
. This species has a very distinctive colour, similar only to
<taxonomicName id="4C504D50FFB3F77E03EA150D4F4FCB2B" authorityName="Köhler" authorityYear="2010" box="[415,602,1352,1375]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="anderdonensis">
<emphasis id="B924EAC1FFB3F77E03EA150D4F4FCB2B" box="[415,602,1352,1375]" italics="true" pageId="34" pageNumber="251">A. anderdonensis</emphasis>
</taxonomicName>
, which differs by smaller shell size, less globose shell shape, higher position of peripheral angulation. Rather flattened diameter of whorls and very broadly conical shape of shell are diagnostic. Material of the present species has been labelled as “
<taxonomicName id="4C504D50FFB3F77E02C7159F4C64CB85" box="[178,369,1498,1521]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="251" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FFB3F77E02C7159F4C59CB85" box="[178,332,1498,1521]" italics="true" pageId="34" pageNumber="251">Amplirhagada</emphasis>
sp.
</taxonomicName>
30” by Solem.
</paragraph>
</subSubSection>
</treatment>
</document>