treatments-xml/data/03/F9/87/03F987C5FF94F75B028013D249A3C9A2.xml
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<document id="6F2E4616681311A19025BF43A4C56952" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF94F75B028013D249A3C9A2" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada solemiana Köhler 2010, n.sp." docType="treatment" docUuid="54256420-1589-4DDC-B13C-96C4708A7750" docUuidSource="ZooBank" docVersion="3" lastPageNumber="224" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="222" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6AD4FE19F2DE7AF21E028374C564BCB1">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="76D13519AA3126C2B43AE1C4D4895C78">Köhler, Frank</mods:namePart>
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<mods:title id="5E316FECC1991C98DF25E47242C4B72B">Records of the Australian Museum</mods:title>
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<mods:date id="1D1AB1E0164D04A5704967A86FB8B93B">2010</mods:date>
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<mods:number id="BD6E917BA8539134B14571AC2E8F3EC3">2010-11-24</mods:number>
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<mods:classification id="46B777EA623FFC9BA9EA661F647BAEA2">journal article</mods:classification>
<mods:identifier id="6D57FC897B9001EDE598DC939E7A25B6" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
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<treatment id="03F987C5FF94F75B028013D249A3C9A2" ID-DOI="http://doi.org/10.5281/zenodo.5239013" ID-Zenodo-Dep="5239013" LSID="urn:lsid:zoobank.org:act:54256420-1589-4DDC-B13C-96C4708A7750" httpUri="http://treatment.plazi.org/id/03F987C5FF94F75B028013D249A3C9A2" lastPageId="7" lastPageNumber="224" pageId="5" pageNumber="222">
<subSubSection id="C34A6558FF94F759028013D24F64CDC5" box="[245,625,919,945]" pageId="5" pageNumber="222" type="nomenclature">
<paragraph id="8BEF36D3FF94F759028013D24F64CDC5" blockId="5.[245,625,919,945]" box="[245,625,919,945]" pageId="5" pageNumber="222">
<heading id="D0A781BFFF94F759028013D24F64CDC5" box="[245,625,919,945]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="222" reason="2">
<emphasis id="B924EAC1FF94F759028013D24F64CDC5" bold="true" box="[245,625,919,945]" pageId="5" pageNumber="222">
<taxonomicName id="4C504D50FF94F759028013D24F39CDC5" ID-CoL="D7RZ" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[245,556,919,945]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="222" phylum="Mollusca" rank="species" species="solemiana" status="sp. nov.">
<emphasis id="B924EAC1FF94F759028013D24F39CDC5" bold="true" box="[245,556,919,945]" italics="true" pageId="5" pageNumber="222">Amplirhagada solemiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF94F759004013D24F64CDC5" box="[565,625,919,945]" pageId="5" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6558FF94F759020C13964CF3CBD3" pageId="5" pageNumber="222" type="materials_examined">
<paragraph id="8BEF36D3FF94F759020C13964C30CA09" blockId="5.[121,749,979,1149]" pageId="5" pageNumber="222">
<materialsCitation id="3B383C8EFF94F759020C13964C37CA09" collectingDate="2008-02-13" collectionCode="V" county="northwestern Kimberley" latitude="-14.305" location="Admiralty Gulf" longLatPrecision="21" longitude="125.99305" municipality="Bonaparte Archipelago" pageId="5" pageNumber="222" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF94F759020C13964C06CD9F" bold="true" box="[121,275,979,1003]" pageId="5" pageNumber="222">
<typeStatus id="54EB8871FF94F759020C13964DA6CD9F" box="[121,179,979,1003]" pageId="5" pageNumber="222">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF94F759035313914C7FCD9F" box="[294,362,980,1003]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="5" pageNumber="222">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF94F75903F013914F5ACD9F" box="[389,591,980,1003]" country="Australia" name="Western Australia" pageId="5" pageNumber="222">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF94F759002813914DF9CA7C" pageId="5" pageNumber="222">northwestern Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF94F759028C13B44CE3CA7C" box="[249,502,1009,1032]" pageId="5" pageNumber="222">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FF94F759007713B44FBBCA7C" LSID="urn:lsid:plazi:treatment:03F987C5FF94F75B028013D249A3C9A2:8E8F6008FF94F759007713B44FBBCA7C" box="[514,686,1009,1032]" county="northwestern Kimberley" latitude="-14.305" longLatPrecision="21" longitude="125.99305" municipality="Bonaparte Archipelago" name="Admiralty Gulf" pageId="5" pageNumber="222" stateProvince="Western Australia">Admiralty Gulf</location>
, west coast of
<location id="8E8F6008FF94F75902AC144B4CD9CA51" LSID="urn:lsid:plazi:treatment:03F987C5FF94F75B028013D249A3C9A2:8E8F6008FF94F75902AC144B4CD9CA51" box="[217,460,1038,1061]" county="northwestern Kimberley" latitude="-14.305" longLatPrecision="21" longitude="125.99305" municipality="Bonaparte Archipelago" name="Middle Osborn Island" pageId="5" pageNumber="222" stateProvince="Western Australia">Middle Osborn Island</location>
,
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<geoCoordinate id="EE645014FF94F7590028144B4FF2CA51" box="[605,743,1038,1061]" degrees="125" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="222" precision="15" seconds="35" value="125.99305">125°59'35&quot;E</geoCoordinate>
; KIS 2030.
<location id="8E8F6008FF94F759028B14694C29CA37" LSID="urn:lsid:plazi:treatment:03F987C5FF94F75B028013D249A3C9A2:8E8F6008FF94F759028B14694C29CA37" box="[254,316,1068,1091]" county="northwestern Kimberley" latitude="-14.305" longLatPrecision="21" longitude="125.99305" municipality="Bonaparte Archipelago" name="Small" pageId="5" pageNumber="222" stateProvince="Western Australia">Small</location>
vine thicket at base of hill, west facing slopes.
<location id="8E8F6008FF94F75902BE140C4C3BCA14" LSID="urn:lsid:plazi:treatment:03F987C5FF94F75B028013D249A3C9A2:8E8F6008FF94F75902BE140C4C3BCA14" box="[203,302,1097,1120]" county="northwestern Kimberley" latitude="-14.305" longLatPrecision="21" longitude="125.99305" municipality="Bonaparte Archipelago" name="Common" pageId="5" pageNumber="222" stateProvince="Western Australia">Common</location>
on trees and bushes (leg.
<collectionCode id="ED41AE16FF94F759003B140C4F74CA14" box="[590,609,1097,1120]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="222" type="Museum">V</collectionCode>
. Kessner,
<date id="FFEE1013FF94F75900A7140C4C0FCA09" pageId="5" pageNumber="222" value="2008-02-13">
<collectingDate id="EFAAE9FBFF94F75900A7140C4C0FCA09" pageId="5" pageNumber="222" value="2008-02-13">13 February 2008</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF94F759020C14DA4C6FCB36" blockId="5.[121,749,1183,1346]" pageId="5" pageNumber="222">
<emphasis id="B924EAC1FF94F759020C14DA4C11CAC6" bold="true" box="[121,260,1183,1202]" pageId="5" pageNumber="222">Type material</emphasis>
.
<materialsCitation id="3B383C8EFF94F759036114DA4F45CAC6" accessionNumber="S34602" box="[276,592,1183,1202]" collectionCode="WAM" pageId="5" pageNumber="222" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FF94F759036114DA4C7CCAC6" box="[276,361,1183,1202]" pageId="5" pageNumber="222" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FF94F759030414DA4CB2CAC6" box="[369,423,1183,1202]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F75903C514DA4CE3CAC6" box="[432,502,1183,1202]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34602" pageId="5" pageNumber="222" type="EnaNcbi">S34602</accessionNumber>
(Pl. 1.3)
</materialsCitation>
.
<materialsCitation id="3B383C8EFF94F759002914DA4DEBCA8E" accessionNumber="S36860" collectionCode="WAM" latitude="-14.3103895" longLatPrecision="1" longitude="125.98844" pageId="5" pageNumber="222" specimenCount="3" typeStatus="paratype">
<typeStatus id="54EB8871FF94F759002914DA4FA0CAC6" box="[604,693,1183,1202]" pageId="5" pageNumber="222" type="paratype">Paratypes</typeStatus>
AMS C463681 (8 preserved specimens, as
<typeStatus id="54EB8871FF94F75903AE14F24F3ACABE" box="[475,559,1207,1226]" pageId="5" pageNumber="222" type="holotype">holotype</typeStatus>
),
<collectionCode id="ED41AE16FF94F759003414F24F62CABE" box="[577,631,1207,1226]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F75900F514F24FD0CABE" box="[640,709,1207,1226]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36860" pageId="5" pageNumber="222" type="EnaNcbi">S36860</accessionNumber>
(20 preserved specimens, as
<typeStatus id="54EB8871FF94F759033F148A4C8DCA96" box="[330,408,1231,1250]" pageId="5" pageNumber="222" type="holotype">holotype</typeStatus>
),AMS C463683 (4 shells,
<geoCoordinate id="EE645014FF94F759000E148A4FF8CA96" box="[635,749,1231,1250]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="5" pageNumber="222" precision="1" seconds="37.4" value="-14.3103895">14°18'37.4&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF94F759020C14A24DEDCA8E" box="[121,248,1255,1274]" degrees="125" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="222" precision="1" seconds="18.4" value="125.98844">125°59'18.4&quot;E</geoCoordinate>
)
</materialsCitation>
,
<materialsCitation id="3B383C8EFF94F759037D14A24FFDCA8E" accessionNumber="S36572" box="[264,744,1255,1274]" collectionCode="WAM" latitude="-14.3103895" longLatPrecision="1" longitude="125.98844" pageId="5" pageNumber="222" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED41AE16FF94F759037D14A24C29CA8E" box="[264,316,1255,1274]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759033614A24C91CA8E" box="[323,388,1255,1274]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36572" pageId="5" pageNumber="222" type="EnaNcbi">S36572</accessionNumber>
(10 shells,
<geoCoordinate id="EE645014FF94F759039E14A24F48CA8E" box="[491,605,1255,1274]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="5" pageNumber="222" precision="1" seconds="37.4" value="-14.3103895">14°18'37.4&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF94F759001614A24FF6CA8E" box="[611,739,1255,1274]" degrees="125" direction="east" minutes="59" orientation="longitude" pageId="5" pageNumber="222" precision="1" seconds="18.4" value="125.98844">125°59'18.4&quot;E</geoCoordinate>
)
</materialsCitation>
, AMS C463682 (2 preserved specimens, east coast,
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<geoCoordinate id="EE645014FF94F759020C15524C17CB5E" box="[121,258,1303,1322]" degrees="126" direction="east" minutes="02" orientation="longitude" pageId="5" pageNumber="222" precision="1" seconds="06.7" value="126.035194">126°02'06.7&quot;E</geoCoordinate>
),
<materialsCitation id="3B383C8EFF94F759036115524C60CB36" accessionNumber="S36859" collectionCode="WAM" latitude="-14.314861" longLatPrecision="1" longitude="126.035194" pageId="5" pageNumber="222" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED41AE16FF94F759036115524C5FCB5E" box="[276,330,1303,1322]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759032615524C8DCB5E" box="[339,408,1303,1322]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36859" pageId="5" pageNumber="222" type="EnaNcbi">S36859</accessionNumber>
(6 preserved specimens, east coast,
<geoCoordinate id="EE645014FF94F759020C156A4DFFCB36" box="[121,234,1327,1346]" degrees="14" direction="south" minutes="18" orientation="latitude" pageId="5" pageNumber="222" precision="1" seconds="53.5" value="-14.314861">14°18'53.5&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FF94F759029A156A4C65CB36" box="[239,368,1327,1346]" degrees="126" direction="east" minutes="02" orientation="longitude" pageId="5" pageNumber="222" precision="1" seconds="06.7" value="126.035194">126°02'06.7&quot;E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<caption id="DF2F665BFF94F759016F15544EAFCBE4" ID-DOI="http://doi.org/10.5281/zenodo.5239094" ID-Zenodo-Dep="5239094" httpUri="https://zenodo.org/record/5239094/files/figure.png" pageId="5" pageNumber="222" startId="5.[794,857,1297,1318]" targetBox="[817,1389,140,1272]" targetPageId="5">
<paragraph id="8BEF36D3FF94F759016F15544EAFCBE4" blockId="5.[794,1422,1297,1424]" pageId="5" pageNumber="222">
Figure 4. Interior of penial chamber of
<taxonomicName id="4C504D50FF94F75906FB15544841CB52" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1166,1364,1297,1318]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="222" phylum="Mollusca" rank="species" species="euroa" status="sp. nov.">
<emphasis id="B924EAC1FF94F75906FB15544841CB52" box="[1166,1364,1297,1318]" italics="true" pageId="5" pageNumber="222">Amplirhagada euroa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF94F759072F1554489CCB52" box="[1370,1417,1297,1318]" pageId="5" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463680 (7 August, scale 5 mm). Labelling of structures: iw, inner penial wall; lp, longitudinal (main) pilaster; rm, penial retractor muscle; sh, penial sheath; vd, vas deferens; vg, penial verge.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF94F759020C15214CF3CBD3" blockId="5.[121,749,1380,1447]" pageId="5" pageNumber="222">
<emphasis id="B924EAC1FF94F759020C15214C92CB03" bold="true" box="[121,391,1380,1399]" pageId="5" pageNumber="222">Additional, non-type material</emphasis>
.
<collectionCode id="ED41AE16FF94F75903E415214CD0CB03" box="[401,453,1380,1399]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F75903BF15214F19CB03" box="[458,524,1380,1399]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36471" pageId="5" pageNumber="222" type="EnaNcbi">S36471</accessionNumber>
,
<collectionCode id="ED41AE16FF94F759006015214F5CCB03" box="[533,585,1380,1399]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759003B15214F84CB03" box="[590,657,1380,1399]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36566" pageId="5" pageNumber="222" type="EnaNcbi">S36566</accessionNumber>
71,
<collectionCode id="ED41AE16FF94F75900CD15214FF9CB03" box="[696,748,1380,1399]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759020C15394DA8CBFB" box="[121,189,1404,1423]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36573" pageId="5" pageNumber="222" type="EnaNcbi">S36573</accessionNumber>
,
<collectionCode id="ED41AE16FF94F75902B115394DEDCBFB" box="[196,248,1404,1423]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759028915394C2ACBFB" box="[252,319,1404,1423]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36585" pageId="5" pageNumber="222" type="EnaNcbi">S36585</accessionNumber>
99,
<collectionCode id="ED41AE16FF94F759031215394C8ECBFB" box="[359,411,1404,1423]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F75903EA15394CF6CBFB" box="[415,483,1404,1423]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36858" pageId="5" pageNumber="222" type="EnaNcbi">S36858</accessionNumber>
,
<collectionCode id="ED41AE16FF94F759039F15394F0BCBFB" box="[490,542,1404,1423]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F759005615394F70CBFB" box="[547,613,1404,1423]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36861" pageId="5" pageNumber="222" type="EnaNcbi">S36861</accessionNumber>
,
<collectionCode id="ED41AE16FF94F759001815394FB4CBFB" box="[621,673,1404,1423]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="5" pageNumber="222" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF94F75900D315394FFCCBFB" box="[678,745,1404,1423]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36971" pageId="5" pageNumber="222" type="EnaNcbi">S36971</accessionNumber>
, AMS C4636845 (Middle Osborn Island).
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF94F75B020C158E49A3C9A2" lastPageId="7" lastPageNumber="224" pageId="5" pageNumber="222" type="etymology">
<paragraph id="8BEF36D3FF94F759020C158E4F48C874" blockId="5.[121,748,1483,1536]" pageId="5" pageNumber="222">
<emphasis id="B924EAC1FF94F759020C158E4DE6CB97" bold="true" box="[121,243,1483,1507]" pageId="5" pageNumber="222">Etymology</emphasis>
. Named in honour of Alan Solem, in recognition of his achievements in camaenid systematics.
</paragraph>
<paragraph id="8BEF36D3FF94F759020C16614CA7C84F" blockId="5.[121,434,1572,1595]" box="[121,434,1572,1595]" pageId="5" pageNumber="222">Sealing strategy. Rock sealer.</paragraph>
<paragraph id="8BEF36D3FF94F759020C161B4922C951" blockId="5.[121,749,1630,1829]" lastBlockId="5.[794,1422,1513,1829]" pageId="5" pageNumber="222">
<emphasis id="B924EAC1FF94F759020C161B4DA4C802" bold="true" box="[121,177,1630,1654]" pageId="5" pageNumber="222">Shell</emphasis>
(
<figureCitation id="136B2A56FF94F75902CB161B4C3BC801" box="[190,302,1630,1653]" captionStart="Figure 5" captionStartId="5.[121,183,1873,1894]" captionText="Figure 5 (facing page).SEM photographs of Amplirhagada solemiana n.sp.(AE). Shell, paratypeAMS C463683:(A) apical whorl viewed from above (scale 200 µm); (B) first four whorls viewed from above (scale 1 mm); (C) shell, lateral view (scale 1 mm); (D) details of axial sculpture on last whorl, lateral view (scale 1 mm); (E) close-up of axial lirae on last whorl (scale 100 µm). (FI) Radula, paratype AMS C463681: (F) rows of central and lateral teeth (Scale 20 µm); (G) details if central and lateral teeth (scale 20 µm); (H) outer lateral and inner marginal teeth (scale 10 µm); (I) middle and outer marginal teeth (scale 10 µm). (J) jaw, paratype AMS C463681 (scale 100 µm)." pageId="5" pageNumber="222">Fig. 5AE</figureCitation>
, Pl. 1.34). Semi-globose with moderately high spire. Thin to solid, translucent. Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls well rounded. Umbilicus open, forming a chink, to 8090 percent concealed by columellar reflection. Background colour horn to yellowish brown, with brown, well defined to diffuse, moderately broad, sub-sutural and mid-whorl bands, visible on entire shell; ventral and outer lip colour horn; inner lip translucent, pale. Protoconch c.
<quantity id="4CA89B36FF94F759071916434E56C84E" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="222" unit="mm" value="1.8">1.8 mm</quantity>
in diameter, comprising about 1.5 whorls, sculptured by comparatively strong axial ribs. Teleoconch sculptured by coarse, regular lirae, rounded in cross-section; spaces equal to thickness of lirae; sculpture evenly distributed across whorls of shell and across whorl diameter, height of lirae reduced underneath suture. Angle of aperture 45°; outer lip sharp to moderately thick, rounded, slightly expanded, slightly reflected; basal and palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<caption id="DF2F665BFF94F759020C17144893C9A5" pageId="5" pageNumber="222" startId="5.[121,183,1873,1894]">
<paragraph id="8BEF36D3FF94F759020C17144893C9A5" blockId="5.[121,1422,1873,2001]" pageId="5" pageNumber="222">
Figure 5 (facing page). SEM photographs of
<taxonomicName id="4C504D50FF94F759006F17174E1FC913" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[538,778,1874,1895]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="222" phylum="Mollusca" rank="species" species="solemiana" status="sp. nov.">
<emphasis id="B924EAC1FF94F759006F17174E1FC913" box="[538,778,1874,1895]" italics="true" pageId="5" pageNumber="222">Amplirhagada solemiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF94F759017B17144E29C912" box="[782,828,1873,1894]" pageId="5" pageNumber="222" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FF94F759013217174E65C913" box="[839,880,1874,1895]" italics="true" pageId="5" pageNumber="222">AE</emphasis>
). Shell, paratype AMS C463683: (
<emphasis id="B924EAC1FF94F75906C0171749D1C913" box="[1205,1220,1874,1895]" italics="true" pageId="5" pageNumber="222">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF94F75903FF17294C8CC9F5" box="[394,409,1900,1921]" italics="true" pageId="5" pageNumber="222">B</emphasis>
) first four whorls viewed from above (scale 1 mm); (
<emphasis id="B924EAC1FF94F75901F917294E89C9F5" box="[908,924,1900,1921]" italics="true" pageId="5" pageNumber="222">C</emphasis>
) shell, lateral view (scale 1 mm); (
<emphasis id="B924EAC1FF94F7590691172949E0C9F5" box="[1252,1269,1900,1921]" italics="true" pageId="5" pageNumber="222">D</emphasis>
) details of axial sculpture on last whorl, lateral view (scale 1 mm); (
<emphasis id="B924EAC1FF94F759001D17C24F62C9E8" box="[616,631,1927,1948]" italics="true" pageId="5" pageNumber="222">E</emphasis>
) close-up of axial lirae on last whorl (scale 100 µm). (
<emphasis id="B924EAC1FF94F75906F417C249B1C9E8" box="[1153,1188,1927,1948]" italics="true" pageId="5" pageNumber="222">FI</emphasis>
) Radula, paratype AMS C463681: (
<emphasis id="B924EAC1FF94F759029317E74DE0C9C3" box="[230,245,1954,1975]" italics="true" pageId="5" pageNumber="222">F</emphasis>
) rows of central and lateral teeth (Scale 20 µm); (
<emphasis id="B924EAC1FF94F75900A317E74FF2C9C3" box="[726,743,1954,1975]" italics="true" pageId="5" pageNumber="222">G</emphasis>
) details if central and lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FF94F75906A417E749F7C9C3" box="[1233,1250,1954,1975]" italics="true" pageId="5" pageNumber="222">H</emphasis>
) outer lateral and inner marginal teeth (scale 10 µm); (
<emphasis id="B924EAC1FF94F75903A217F94CCAC9A5" box="[471,479,1980,2001]" italics="true" pageId="5" pageNumber="222">I</emphasis>
) middle and outer marginal teeth (scale 10 µm). (
<emphasis id="B924EAC1FF94F75901C917F94ED2C9A5" box="[956,967,1980,2001]" italics="true" pageId="5" pageNumber="222">J</emphasis>
) jaw, paratype AMS C463681 (scale 100 µm).
</paragraph>
</caption>
<caption id="DF2F665BFF96F75B020C11BA4C66CC3E" ID-DOI="http://doi.org/10.5281/zenodo.5239096" ID-Zenodo-Dep="5239096" httpUri="https://zenodo.org/record/5239096/files/figure.png" pageId="7" pageNumber="224" startId="7.[121,184,511,532]" targetBox="[127,742,152,497]" targetPageId="7">
<paragraph id="8BEF36D3FF96F75B020C11BA4C66CC3E" blockId="7.[121,748,511,586]" pageId="7" pageNumber="224">
Figure 6. Genitalia of
<taxonomicName id="4C504D50FF96F75B032E12454F46CC61" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[347,595,512,533]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana" status="sp. nov.">
<emphasis id="B924EAC1FF96F75B032E12454F46CC61" box="[347,595,512,533]" italics="true" pageId="7" pageNumber="224">Amplirhagada solemiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF96F75B002911BA4F99CC60" box="[604,652,511,532]" pageId="7" pageNumber="224" rank="species">n.sp.</taxonomicNameLabel>
, paratype WAM
<accessionNumber id="9403AB30FF96F75B02CF125F4C11CC5B" box="[186,260,538,559]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S36581" pageId="7" pageNumber="224" type="EnaNcbi">S36581</accessionNumber>
(13 February, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF96F75B020C12D34C67CD71" blockId="7.[121,749,662,773]" pageId="7" pageNumber="224">
<emphasis id="B924EAC1FF96F75B020C12D34C4ECCDA" bold="true" box="[121,347,662,686]" pageId="7" pageNumber="224">Pallial morphology</emphasis>
. Pallial cavity deep, extending one whorl. Pigmentation on mantle consists of sparsely spaced dark grey-black spots. Kidney extending about half of pallial cavity or slightly more.
</paragraph>
<paragraph id="8BEF36D3FF96F75B020C136C4CE8CBDC" blockId="7.[121,749,809,1448]" pageId="7" pageNumber="224">
<emphasis id="B924EAC1FF96F75B020C136C4C4ACD35" bold="true" box="[121,351,809,833]" pageId="7" pageNumber="224">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF96F75B0318136C4CC3CD34" box="[365,470,809,832]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="7.[121,184,511,532]" captionStartId-1="7.[794,857,1252,1273]" captionTargetBox-0="[127,742,152,497]" captionTargetBox-1="[829,1376,143,1246]" captionTargetId-0="figure-370@7.[121,748,142,506]" captionTargetId-1="figure-420@7.[829,1400,138,1247]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figure 6. Genitalia of Amplirhagada solemiana n.sp., paratype WAM S36581 (13 February, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 7. Interior of penial chamber of Amplirhagada solemiana n.sp., paratypeAMS C463681 (13 February,scale 3 mm).Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239096" figureDoi-1="http://doi.org/10.5281/zenodo.5239099" httpUri-0="https://zenodo.org/record/5239096/files/figure.png" httpUri-1="https://zenodo.org/record/5239099/files/figure.png" pageId="7" pageNumber="224">Figs. 67</figureCitation>
). Penis straight to slightly curved; same length as anterior part of oviduct or slightly longer.Vas deferens forms simple loop before entering penis. Penial retractor muscle about as long as penis. Penial verge long to very long (c.
<emphasis id="B924EAC1FF96F75B032F13DB4C77CDD8" bold="true" box="[346,354,926,940]" pageId="7" pageNumber="224">
<superScript id="7C259B9BFF96F75B032F13DB4C77CDD8" attach="right" box="[346,354,926,940]" fontSize="6" pageId="7" pageNumber="224">1</superScript>
</emphasis>
<emphasis id="B924EAC1FF96F75B031313E04C7BCDC7" bold="true" box="[358,366,933,947]" pageId="7" pageNumber="224">
<subScript id="17D43496FF96F75B031313E04C7BCDC7" attach="left" box="[358,366,933,947]" fontSize="6" pageId="7" pageNumber="224">3</subScript>
</emphasis>
to ½ of penial chamber), slender to spatulate with pointed tip. Pustulation comprising entire length of inner penial wall; pustules small to moderate in size, densely arranged in rows, some of which form four to five corrugated longitudinal pilasters along entire length of penial chamber; main stimulatory pilaster not differentiated. Vas deferens entering penial sheath apically. Vagina of medium length, tubular or posteriorly inflated. Inner vaginal wall densely ciliated, ciliae arranged to form smooth longitudinal pilasters, pilasters may be weakly developed. Spermathecal duct of medium thickness; internally with smooth longitudinal pilasters. Spermathecal head elongately inflated, connected with oviduct by connective tissue; inside entirely smooth; wall moderately thick. Length of free oviduct equivalent to about half of anterior part of oviduct; coiled underneath entrance to spermoviduct. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland close to anterior end.
</paragraph>
<caption id="DF2F665BFF96F75B016F14A14996CB5B" ID-DOI="http://doi.org/10.5281/zenodo.5239099" ID-Zenodo-Dep="5239099" httpUri="https://zenodo.org/record/5239099/files/figure.png" pageId="7" pageNumber="224" startId="7.[794,857,1252,1273]" targetBox="[829,1376,143,1246]" targetPageId="7">
<paragraph id="8BEF36D3FF96F75B016F14A14996CB5B" blockId="7.[794,1421,1252,1327]" pageId="7" pageNumber="224">
Figure 7. Interior of penial chamber of
<taxonomicName id="4C504D50FF96F75B06EC14A14898CA8D" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1177,1421,1252,1273]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana" status="sp. nov.">
<emphasis id="B924EAC1FF96F75B06EC14A14898CA8D" box="[1177,1421,1252,1273]" italics="true" pageId="7" pageNumber="224">Amplirhagada solemiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF96F75B016F14BA4E5DCB60" box="[794,840,1279,1300]" pageId="7" pageNumber="224" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463681 (13 February, scale 3 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FF96F75B020C158E4F9FC869" blockId="7.[121,749,1483,1741]" pageId="7" pageNumber="224">
<emphasis id="B924EAC1FF96F75B020C158E4C62CB97" bold="true" box="[121,375,1483,1507]" pageId="7" pageNumber="224">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FF96F75B03FE15894CE8CB97" box="[395,509,1484,1507]" captionStart="Figure 5" captionStartId="5.[121,183,1873,1894]" captionText="Figure 5 (facing page).SEM photographs of Amplirhagada solemiana n.sp.(AE). Shell, paratypeAMS C463683:(A) apical whorl viewed from above (scale 200 µm); (B) first four whorls viewed from above (scale 1 mm); (C) shell, lateral view (scale 1 mm); (D) details of axial sculpture on last whorl, lateral view (scale 1 mm); (E) close-up of axial lirae on last whorl (scale 100 µm). (FI) Radula, paratype AMS C463681: (F) rows of central and lateral teeth (Scale 20 µm); (G) details if central and lateral teeth (scale 20 µm); (H) outer lateral and inner marginal teeth (scale 10 µm); (I) middle and outer marginal teeth (scale 10 µm). (J) jaw, paratype AMS C463681 (scale 100 µm)." pageId="7" pageNumber="224">Fig. 5FI</figureCitation>
). Rectangular. Tooth formula C+912+34+1921. Average number of rows of teeth 131±22 with 29.9±0.6 rows per mm (n = 3).
</paragraph>
<paragraph id="8BEF36D3FF96F75B02E216614FF1C8B9" blockId="7.[121,749,1483,1741]" pageId="7" pageNumber="224">Central teeth with bluntly pointed, ovate mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with bluntly pointed ovate mesocones, length equal to base of tooth; small ectocones; endocones absent. Marginal teeth multicuspic, mesocone and endocone similar in length, ectocone smaller than endocone, occasionally subdivided.</paragraph>
<paragraph id="8BEF36D3FF96F75B020C16B449A3C9A2" blockId="7.[121,749,1777,2005]" lastBlockId="7.[794,1422,1396,2006]" pageId="7" pageNumber="224">
<emphasis id="B924EAC1FF96F75B020C16B44C61C97D" bold="true" box="[121,372,1777,1801]" pageId="7" pageNumber="224">Comparative remarks</emphasis>
. Shells are close to
<taxonomicName id="4C504D50FF96F75B002016B44C84C951" authority="(E. A. Smith, 1894)" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B002016B44DD6C952" italics="true" pageId="7" pageNumber="224">Amplirhagada imitata</emphasis>
(E. A.
<bibRefCitation id="EFC14B22FF96F75B037F174B4C9FC951" author="Smith, E" box="[266,394,1806,1829]" pageId="7" pageNumber="224" pagination="84 - 99" refId="ref39111" refString="Smith, E. A., 1894. On the land-shells of Western Australia. Proceedings of the Malacological Society of London 1: 84 - 99." type="journal article" year="1894">Smith, 1894</bibRefCitation>
)
</taxonomicName>
in overall shape, colouration, and sculpture, but see
<figureCitation id="136B2A56FF96F75B034317694C60C937" box="[310,373,1836,1859]" captionStart="Figure 8" captionStartId="8.[166,229,764,785]" captionTargetBox="[170,794,145,747]" captionTargetId="figure-254@8.[167,794,142,748]" captionTargetPageId="8" captionText="Figure 8. Comparison of A. solemiana and A. indistincta by means of shell parameters. Scatter-plot showing the ratio of shell height (H) versus diameter (D)." figureDoi="http://doi.org/10.5281/zenodo.5239103" httpUri="https://zenodo.org/record/5239103/files/figure.png" pageId="7" pageNumber="224">Fig. 8</figureCitation>
. The
<typeStatus id="54EB8871FF96F75B03C517694CC8C937" box="[432,477,1836,1859]" pageId="7" pageNumber="224">type</typeStatus>
locality of
<taxonomicName id="4C504D50FF96F75B002017694FA8C937" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" box="[597,701,1836,1859]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B002017694FA8C937" box="[597,701,1836,1859]" italics="true" pageId="7" pageNumber="224">A. imitata</emphasis>
</taxonomicName>
was restricted to Baudin Island by
<bibRefCitation id="EFC14B22FF96F75B03C5170C4F59C914" author="Solem, A" box="[432,588,1865,1888]" pageId="7" pageNumber="224" pagination="147 - 320" refId="ref39183" refString="Solem, A., 1981 a. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Supplement 11: 147 - 320." type="journal article" year="1981">Solem (1981a)</bibRefCitation>
.
<taxonomicName id="4C504D50FF96F75B0023170C4DD6C90A" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B0023170C4DD6C90A" italics="true" pageId="7" pageNumber="224">Amplirhagada imitata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF96F75B028917224C92C90A" authorityName="Köhler" authorityYear="2010" box="[252,391,1895,1918]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana">
<emphasis id="B924EAC1FF96F75B028917224C92C90A" box="[252,391,1895,1918]" italics="true" pageId="7" pageNumber="224">A. solemiana</emphasis>
</taxonomicName>
exhibit the most sharply defined and prominent axial sculpture among all congeners except for
<taxonomicName id="4C504D50FF96F75B02EB17E44C3CC9CC" authorityName="Köhler" authorityYear="2010" box="[158,297,1953,1976]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="indistincta">
<emphasis id="B924EAC1FF96F75B02EB17E44C3CC9CC" box="[158,297,1953,1976]" italics="true" pageId="7" pageNumber="224">A. indistincta</emphasis>
</taxonomicName>
. The anatomy of
<taxonomicName id="4C504D50FF96F75B039417E44F5DC9CC" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" box="[481,584,1953,1976]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B039417E44F5DC9CC" box="[481,584,1953,1976]" italics="true" pageId="7" pageNumber="224">A. imitata</emphasis>
</taxonomicName>
is unknown for only dry shells were available to
<bibRefCitation id="EFC14B22FF96F75B03A417FB4F7AC9A1" author="Solem, A" box="[465,623,1982,2005]" pageId="7" pageNumber="224" pagination="147 - 320" refId="ref39183" refString="Solem, A., 1981 a. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Supplement 11: 147 - 320." type="journal article" year="1981">Solem (1981a)</bibRefCitation>
. In between Baudin Island (
<taxonomicName id="4C504D50FF96F75B01B51531493FCBFF" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" box="[960,1066,1396,1419]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B01B51531493FCBFF" box="[960,1066,1396,1419]" italics="true" pageId="7" pageNumber="224">A. imitata</emphasis>
</taxonomicName>
) and the two Osborn Islands (
<taxonomicName id="4C504D50FF96F75B070315314E91CBDD" authorityName="Köhler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana">
<emphasis id="B924EAC1FF96F75B070315314E91CBDD" italics="true" pageId="7" pageNumber="224">A. solemiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF96F75B01C715D7492CCBDD" authorityName="Köhler" authorityYear="2010" box="[946,1081,1426,1449]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="indistincta">
<emphasis id="B924EAC1FF96F75B01C715D7492CCBDD" box="[946,1081,1426,1449]" italics="true" pageId="7" pageNumber="224">A. indistincta</emphasis>
</taxonomicName>
) there are numerous other islands that are occupied by
<taxonomicName id="4C504D50FF96F75B018115EA49C9CBB2" box="[1012,1244,1455,1478]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FF96F75B018115EA499ECBB2" box="[1012,1163,1455,1478]" italics="true" pageId="7" pageNumber="224">Amplirhagada</emphasis>
species
</taxonomicName>
(e.g., Kingsmill, Corneille, Fenelon, and Descartes Islands) (
<figureCitation id="136B2A56FF96F75B06AC15894801CB97" box="[1241,1300,1484,1507]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="7" pageNumber="224">Fig. 1</figureCitation>
). Given the usually very restricted occurrence of
<taxonomicName id="4C504D50FF96F75B06D615AF4898C875" box="[1187,1421,1514,1537]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FF96F75B06D615AF482FC875" box="[1187,1338,1514,1537]" italics="true" pageId="7" pageNumber="224">Amplirhagada</emphasis>
species
</taxonomicName>
and the marked genetic differentiation of
<taxonomicName id="4C504D50FF96F75B06A01642484BC86A" authorityName="Köhler" authorityYear="2010" box="[1237,1374,1543,1566]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana">
<emphasis id="B924EAC1FF96F75B06A01642484BC86A" box="[1237,1374,1543,1566]" italics="true" pageId="7" pageNumber="224">A. solemiana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D50FF96F75B016F16614EBCC84F" authorityName="Köhler" authorityYear="2010" box="[794,937,1572,1595]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="indistincta">
<emphasis id="B924EAC1FF96F75B016F16614EBCC84F" box="[794,937,1572,1595]" italics="true" pageId="7" pageNumber="224">A. indistincta</emphasis>
</taxonomicName>
, it is considered very unlikely that either of them could be conspecific with
<taxonomicName id="4C504D50FF96F75B06F3160749E0C82D" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" box="[1158,1269,1602,1625]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B06F3160749E0C82D" box="[1158,1269,1602,1625]" italics="true" pageId="7" pageNumber="224">A. imitata</emphasis>
</taxonomicName>
from Baudin Island, which is located in a considerable distance (c.
<quantity id="4CA89B36FF96F75B073E161A4898C802" box="[1355,1421,1631,1654]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="224" unit="km" value="50.0">50 km</quantity>
NW of Osborn Islands). Based on shell features only,
<bibRefCitation id="EFC14B22FF96F75B073F16394E7EC8C5" author="Solem, A" pageId="7" pageNumber="224" pagination="147 - 320" refId="ref39183" refString="Solem, A., 1981 a. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). II. Taxa from the Kimberley, Amplirhagada Iredale 1933. Records of the Western Australian Museum Supplement 11: 147 - 320." type="journal article" year="1981">Solem (1981a)</bibRefCitation>
also synonymized
<taxonomicName id="4C504D50FF96F75B064516DF4841C8C5" authority="Iredale, 1938" authorityName="Iredale" authorityYear="1938" box="[1072,1364,1690,1713]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="burrowsena">
<emphasis id="B924EAC1FF96F75B064516DF49D2C8C5" box="[1072,1223,1690,1713]" italics="true" pageId="7" pageNumber="224">A. burrowsena</emphasis>
<bibRefCitation id="EFC14B22FF96F75B06B916DF4841C8C5" author="Iredale, T" box="[1228,1364,1690,1713]" pageId="7" pageNumber="224" pagination="83 - 124" refId="ref38394" refString="Iredale, T., 1938. A basic list of the land Mollusca of Australia - Part III. Australian Zoologist 9: 83 - 124." type="journal article" year="1938">Iredale, 1938</bibRefCitation>
</taxonomicName>
from Vansittart Bay (c.
<quantity id="4CA89B36FF96F75B01A316F2490DC8BA" box="[982,1048,1719,1742]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="224" unit="km" value="80.0">80 km</quantity>
E of Baudin Is) with
<taxonomicName id="4C504D50FF96F75B068F16F24874C8BA" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" box="[1274,1377,1719,1742]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">
<emphasis id="B924EAC1FF96F75B068F16F24874C8BA" box="[1274,1377,1719,1742]" italics="true" pageId="7" pageNumber="224">A. imitata</emphasis>
</taxonomicName>
and reported this species to further occur in rainforest patches on the mainland (RFS 03/3, 11/1). These reports, however, almost certainly refer to extralimital populations. Species such as
<emphasis id="B924EAC1FF96F75B0703174A4836C937" italics="true" pageId="7" pageNumber="224">
<taxonomicName id="4C504D50FF96F75B0703174A4E70C937" baseAuthorityName="E. A. Smith" baseAuthorityYear="1894" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="imitata">A. imitata</taxonomicName>
,
<taxonomicName id="4C504D50FF96F75B011817694EE0C937" authorityName="Köhler" authorityYear="2010" box="[877,1013,1836,1859]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="solemiana">A. solemiana</taxonomicName>
,
<taxonomicName id="4C504D50FF96F75B018817694992C937" authorityName="Köhler" authorityYear="2010" box="[1021,1159,1836,1859]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="indistincta">A. indistincta</taxonomicName>
,
<taxonomicName id="4C504D50FF96F75B06FA17694836C937" authorityName="Iredale" authorityYear="1938" box="[1167,1315,1836,1859]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="burrowsena">A. burrowsena</taxonomicName>
</emphasis>
and likely an undescribed species from the rainforest patches on the mainland may possibly form a group of sibling species that are not clearly differentiated by means of their shell morphology as is also known from the species group of
<taxonomicName id="4C504D50FF96F75B069917E74E88C9A2" authority="Solem, 1981" authorityName="Solem" authorityYear="1981" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="224" phylum="Mollusca" rank="species" species="mitchelliana">
<emphasis id="B924EAC1FF96F75B069917E74898C9CD" box="[1260,1421,1954,1977]" italics="true" pageId="7" pageNumber="224">A. mitchelliana</emphasis>
Solem, 1981
</taxonomicName>
from the Mitchell Plateau.
</paragraph>
</subSubSection>
</treatment>
</document>