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<document id="D6575F87A941357AF6BE963E1C27DE03" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FF8FF77C01DD145D4FD6C9AD" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada anderdonensis Köhler 2010, n.sp." docType="treatment" docUuid="2BA494DB-7453-4613-98D4-658575F73813" docUuidSource="ZooBank" docVersion="3" lastPageNumber="249" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="247" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="B78558DA2C49A4594DC97DD78D28DF7D">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
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<mods:namePart id="4B92FEB097929F3EC1DC4C7FF1FC268B">Köhler, Frank</mods:namePart>
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<mods:title id="7CC46B1FA3D8ACDFE09A25F497ECE22B">Records of the Australian Museum</mods:title>
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<mods:date id="1BFA24716ABA1EBB6002FB4ED0D3396C">2010</mods:date>
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<mods:number id="45BEC968D1E0716F2DC77EBC71DECEDC">2010-11-24</mods:number>
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<mods:number id="9E1226DC3201744655200B61001E0A06">62</mods:number>
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<mods:url id="621D75CBA237384E77110D95CD817806">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
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<mods:identifier id="2782EBF0A431BE8126C40511CFA20C0D" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
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<treatment id="03F987C5FF8FF77C01DD145D4FD6C9AD" LSID="urn:lsid:zoobank.org:act:2BA494DB-7453-4613-98D4-658575F73813" httpUri="http://treatment.plazi.org/id/03F987C5FF8FF77C01DD145D4FD6C9AD" lastPageId="32" lastPageNumber="249" pageId="30" pageNumber="247">
<subSubSection id="C34A6558FF8FF74201DD145D484CCA46" box="[936,1369,1048,1074]" pageId="30" pageNumber="247" type="nomenclature">
<paragraph id="8BEF36D3FF8FF74201DD145D484CCA46" blockId="30.[936,1369,1048,1074]" box="[936,1369,1048,1074]" pageId="30" pageNumber="247">
<heading id="D0A781BFFF8FF74201DD145D484CCA46" box="[936,1369,1048,1074]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="247" reason="2">
<emphasis id="B924EAC1FF8FF74201DD145D484CCA46" bold="true" box="[936,1369,1048,1074]" pageId="30" pageNumber="247">
<taxonomicName id="4C504D50FF8FF74201DD145D4803CA46" ID-CoL="D7PY" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[936,1302,1048,1074]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="anderdonensis" status="sp. nov.">
<emphasis id="B924EAC1FF8FF74201DD145D4803CA46" bold="true" box="[936,1302,1048,1074]" italics="true" pageId="30" pageNumber="247">Amplirhagada anderdonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8FF7420768145D484CCA46" box="[1309,1369,1048,1074]" pageId="30" pageNumber="247" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6558FF8FF74201321410497BCB32" pageId="30" pageNumber="247" type="materials_examined">
<paragraph id="8BEF36D3FF8FF742013214104967CA95" blockId="30.[839,1467,1109,1249]" pageId="30" pageNumber="247">
<materialsCitation id="3B383C8EFF8FF74201321410497BCA95" collectingDate="1988-07-22" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="Kimberley" latitude="-14.963611" location="Prince Frederick Harbour" municipality="Bonaparte Archipelago" pageId="30" pageNumber="247" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FF8FF742013214104EF3CA19" bold="true" box="[839,998,1109,1133]" pageId="30" pageNumber="247">
<typeStatus id="54EB8871FF8FF742013214104E96CA19" box="[839,899,1109,1133]" pageId="30" pageNumber="247">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FF8FF742018F14104954CA18" box="[1018,1089,1109,1132]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="30" pageNumber="247">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FF8FF74206281410483BCA18" box="[1117,1326,1109,1132]" country="Australia" name="Western Australia" pageId="30" pageNumber="247">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFF8FF7420748141048A2CA18" box="[1341,1463,1109,1132]" pageId="30" pageNumber="247">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FF8FF74201321437492FCAFD" box="[839,1082,1138,1161]" pageId="30" pageNumber="247">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FF8FF74206371437485ECAFD" LSID="urn:lsid:plazi:treatment:03F987C5FF8FF77C01DD145D4FD6C9AD:8E8F6008FF8FF74206371437485ECAFD" box="[1090,1355,1138,1161]" county="Kimberley" latitude="-14.963611" municipality="Bonaparte Archipelago" name="Prince Frederick Harbour" pageId="30" pageNumber="247" stateProvince="Western Australia">Prince Frederick Harbour</location>
,
<location id="8E8F6008FF8FF742072414374E82CAD3" LSID="urn:lsid:plazi:treatment:03F987C5FF8FF77C01DD145D4FD6C9AD:8E8F6008FF8FF742072414374E82CAD3" county="Kimberley" latitude="-14.963611" municipality="Bonaparte Archipelago" name="Anderdon Islands" pageId="30" pageNumber="247" stateProvince="Western Australia">Anderdon Islands</location>
, unnamed island c.
<locationDeviation id="9A5D2FF1FF8FF742060914D548A2CAD3" box="[1148,1463,1168,1191]" location="Cape Torrens" pageId="30" pageNumber="247">
<quantity id="4CA89B36FF8FF742060914D549A2CAD3" box="[1148,1207,1168,1191]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="247" unit="km" value="8.0">8 km</quantity>
ENE of Cape Torrens
</locationDeviation>
,
<geoCoordinate id="EE645014FF8FF742013214E84ED0CAB0" box="[839,965,1197,1220]" degrees="14" direction="south" minutes="57" orientation="latitude" pageId="30" pageNumber="247" precision="15" seconds="49" value="-14.963611">14°57'49&quot;S</geoCoordinate>
125°09'30&quot;; KC-084 (leg.
<collectionCode id="ED41AE16FF8FF742068814E84804CAB0" box="[1277,1297,1197,1220]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="30" pageNumber="247" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FF8FF742076A14E8486DCAB0" box="[1311,1400,1197,1220]" pageId="30" pageNumber="247">Kessner</collectorName>
&amp;
<collectorName id="26A55305FF8FF74207D514E84EDBCA95" pageId="30" pageNumber="247">A. Longbottom</collectorName>
,
<date id="FFEE1013FF8FF74201AD148F4973CA95" box="[984,1126,1226,1249]" pageId="30" pageNumber="247" value="1988-07-22">
<collectingDate id="EFAAE9FBFF8FF74201AD148F4973CA95" box="[984,1126,1226,1249]" pageId="30" pageNumber="247" value="1988-07-22">22 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FF8FF74201321546497BCB32" blockId="30.[839,1467,1283,1350]" pageId="30" pageNumber="247">
<emphasis id="B924EAC1FF8FF742013215464EDECB62" bold="true" box="[839,971,1283,1302]" pageId="30" pageNumber="247">Type material</emphasis>
.
<materialsCitation id="3B383C8EFF8FF74201AF1546497BCB32" accessionNumber="S34613, S41492" collectionCode="WAM" pageId="30" pageNumber="247" specimenCode="FMNH 219330" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FF8FF74201AF1546493ECB62" box="[986,1067,1283,1302]" pageId="30" pageNumber="247" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FF8FF742064715464973CB62" box="[1074,1126,1283,1302]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="30" pageNumber="247" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF8FF742061A154649A4CB62" box="[1135,1201,1283,1302]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34613" pageId="30" pageNumber="247" type="EnaNcbi">S34613</accessionNumber>
(Pl. 1.19). AMS C463756 (5 preserved specimens),
<specimenCode id="DBF69EA8FF8FF742067A155E4981CB5A" box="[1039,1172,1307,1326]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="30" pageNumber="247" type="Museum">FMNH 219330</specimenCode>
(16 preserved specimens),
<collectionCode id="ED41AE16FF8FF74207F2155E48AECB5A" box="[1415,1467,1307,1326]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="30" pageNumber="247" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FF8FF742013215764E9DCB32" box="[839,904,1331,1350]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41492" pageId="30" pageNumber="247" type="EnaNcbi">S41492</accessionNumber>
(10 preserved specimens).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FF8FF77C0132152F4FD6C9AD" lastPageId="32" lastPageNumber="249" pageId="30" pageNumber="247" type="etymology">
<paragraph id="8BEF36D3FF8FF7420132152F4EFDCBEB" blockId="30.[839,1467,1386,1439]" pageId="30" pageNumber="247">
<emphasis id="B924EAC1FF8FF7420132152F4ED7CBF6" bold="true" box="[839,962,1386,1410]" pageId="30" pageNumber="247">Etymology</emphasis>
. In reference to Anderdon Islands, where this species occurs.
</paragraph>
<paragraph id="8BEF36D3FF8FF743013215874EC8C97E" blockId="30.[839,1153,1474,1497]" lastBlockId="31.[794,1421,1691,1802]" lastPageId="31" lastPageNumber="248" pageId="30" pageNumber="247">
Sealing strategy. Rock sealer.
<emphasis id="B924EAC1FF8EF743020C16DE4DA4C8C7" bold="true" box="[121,177,1691,1715]" pageId="31" pageNumber="248">Shell</emphasis>
(
<figureCitation id="136B2A56FF8EF74302CA16DE4C55C8C6" box="[191,320,1691,1714]" captionStart="Figure 46" captionStartId="31.[121,184,1555,1576]" captionTargetBox="[121,1421,151,1540]" captionTargetId="figure-9@31.[121,1421,151,1540]" captionTargetPageId="31" captionText="Figure 46. SEM photographs of Amplirhagada anderdonensis n.sp. (AC) Shell, paratype AMS C463756: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second and third whorl viewed from above (scale 200 µm); (C) sculpture on upper part of last whorl (scale 100 µm). (D) Jaw (scale 100 µm). (EG) Radula, paratype FMNH 219330: (E) central and lateral teeth (scale 20 µm); (F) close-up of lateral teeth (scale 10 µm); (G) outer lateral and inner marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239181" httpUri="https://zenodo.org/record/5239181/files/figure.png" pageId="31" pageNumber="248">Fig. 46AC</figureCitation>
, Pl. 1.19). Broadly conical with medium high spire, solid. Periphery evenly rounded to slightly angulate; upper and basal sectors of whorls rounded. Umbilicus forming a chink, 90100 percent concealed by columellar reflection. Background colour greenish ochre to yellowish brown; sub-sutural and mid-whorl band absent or diffuse, darker than background, thin, most conspicuous on last whorl; ventral colour cream; outer lip colour same as shell; inner lip translucent, whitish. Protoconch c.
<quantity id="4CA89B36FF8EF74300E617C34FC5C9E9" box="[659,720,1926,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="248" unit="mm" value="2.0">2 mm</quantity>
in diameter, comprising 1.7 whorls, almost smooth. Teleoconch with inconspicuous axial growth lines, last whorl smooth. Angle of aperture 30°; outer lip rounded, moderate to thick, slightly expanded, slightly reflected; basal node of lip absent or weak; palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<caption id="DF2F665BFF8FF74202D317CE4CCAC9A2" ID-DOI="http://doi.org/10.5281/zenodo.5239177" ID-Zenodo-Dep="5239177" httpUri="https://zenodo.org/record/5239177/files/figure.png" pageId="30" pageNumber="247" startId="30.[166,228,1931,1952]" targetBox="[190,781,748,1873]" targetPageId="30">
<paragraph id="8BEF36D3FF8FF74202D317CE4CCAC9A2" blockId="30.[166,794,1931,2006]" pageId="30" pageNumber="247">
Figure 44. Interior of penial chamber of
<taxonomicName id="4C504D50FF8FF742006217C94E0FC9D5" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[535,794,1932,1953]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="lamarckiana" status="sp. nov.">
<emphasis id="B924EAC1FF8FF742006217C94E0FC9D5" box="[535,794,1932,1953]" italics="true" pageId="30" pageNumber="247">Amplirhagada lamarckiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8FF74202D317E34DC0C9CF" box="[166,213,1958,1979]" pageId="30" pageNumber="247" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219163 (14 July, scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFF8FF742013217CE4935C9A1" ID-DOI="http://doi.org/10.5281/zenodo.5239179" ID-Zenodo-Dep="5239179" httpUri="https://zenodo.org/record/5239179/files/figure.png" pageId="30" pageNumber="247" startId="30.[839,900,1931,1952]" targetBox="[844,1463,1516,1907]" targetPageId="30">
<paragraph id="8BEF36D3FF8FF742013217CE4935C9A1" blockId="30.[839,1467,1931,2005]" pageId="30" pageNumber="247">
Figure 45. Genitalia of
<taxonomicName id="4C504D50FF8FF742066917CE483AC9D4" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[1052,1327,1931,1952]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="247" phylum="Mollusca" rank="species" species="anderdonensis" status="sp. nov.">
<emphasis id="B924EAC1FF8FF742066917CE483AC9D4" box="[1052,1327,1931,1952]" italics="true" pageId="30" pageNumber="247">Amplirhagada anderdonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8FF742074617CE4877C9D4" box="[1331,1378,1931,1952]" pageId="30" pageNumber="247" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219330 (22 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFF8EF743020C1656491FC80C" ID-DOI="http://doi.org/10.5281/zenodo.5239181" ID-Zenodo-Dep="5239181" httpUri="https://zenodo.org/record/5239181/files/figure.png" pageId="31" pageNumber="248" startId="31.[121,184,1555,1576]" targetBox="[121,1421,151,1540]" targetPageId="31">
<paragraph id="8BEF36D3FF8EF743020C1656491FC80C" blockId="31.[121,1422,1555,1656]" pageId="31" pageNumber="248">
Figure 46. SEM photographs of
<taxonomicName id="4C504D50FF8EF74303DB16564FDCC85C" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[430,713,1555,1576]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="248" phylum="Mollusca" rank="species" species="anderdonensis" status="sp. nov.">
<emphasis id="B924EAC1FF8EF74303DB16564FDCC85C" box="[430,713,1555,1576]" italics="true" pageId="31" pageNumber="248">Amplirhagada anderdonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFF8EF74300BB16564FE9C85C" box="[718,764,1555,1576]" pageId="31" pageNumber="248" rank="species">n.sp.</taxonomicNameLabel>
(
<emphasis id="B924EAC1FF8EF743017C16564E21C85C" box="[777,820,1555,1576]" italics="true" pageId="31" pageNumber="248">AC</emphasis>
) Shell, paratype AMS C463756: (
<emphasis id="B924EAC1FF8EF743060E1656499FC85C" box="[1147,1162,1555,1576]" italics="true" pageId="31" pageNumber="248">A</emphasis>
) apical whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF8EF743032A16684C7BC836" box="[351,366,1581,1602]" italics="true" pageId="31" pageNumber="248">B</emphasis>
) sculpture on second and third whorl viewed from above (scale 200 µm); (
<emphasis id="B924EAC1FF8EF743063C1668494CC836" box="[1097,1113,1581,1602]" italics="true" pageId="31" pageNumber="248">C</emphasis>
) sculpture on upper part of last whorl (scale 100 µm). (
<emphasis id="B924EAC1FF8EF7430329160D4C78C829" box="[348,365,1608,1629]" italics="true" pageId="31" pageNumber="248">D</emphasis>
) Jaw (scale 100 µm). (
<emphasis id="B924EAC1FF8EF743003E160D4F62C829" box="[587,631,1608,1629]" italics="true" pageId="31" pageNumber="248">EG</emphasis>
) Radula, paratype FMNH 219330: (
<emphasis id="B924EAC1FF8EF74301AD160D4EF2C829" box="[984,999,1608,1629]" italics="true" pageId="31" pageNumber="248">E</emphasis>
) central and lateral teeth (scale 20 µm); (
<emphasis id="B924EAC1FF8EF7430702160D4893C829" box="[1399,1414,1608,1629]" italics="true" pageId="31" pageNumber="248">F</emphasis>
) close-up of lateral teeth (scale 10 µm); (
<emphasis id="B924EAC1FF8EF743038D16264F1CC80C" box="[504,521,1635,1656]" italics="true" pageId="31" pageNumber="248">G</emphasis>
) outer lateral and inner marginal teeth (scale 10 µm).
</paragraph>
</caption>
<paragraph id="8BEF36D3FF8EF743016F1768489EC9F4" blockId="31.[794,1421,1837,1920]" pageId="31" pageNumber="248">
<emphasis id="B924EAC1FF8EF743016F17684EE9C931" bold="true" box="[794,1020,1837,1861]" pageId="31" pageNumber="248">Pallial morphology</emphasis>
. Pallial cavity deep, extending one whorl. Pigmentation on mantle consists of sparse greyish to brown spots. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FF8EF77C016F17E64CF8CB92" blockId="31.[794,1421,1955,2008]" lastBlockId="32.[166,795,989,1510]" lastPageId="32" lastPageNumber="249" pageId="31" pageNumber="248">
<emphasis id="B924EAC1FF8EF743016F17E6491BC9CF" bold="true" box="[794,1038,1955,1979]" pageId="31" pageNumber="248">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FF8EF743065617E64993C9CE" box="[1059,1158,1955,1978]" captionStart="Figure 45" captionStartId="30.[839,900,1931,1952]" captionTargetBox="[844,1463,1516,1907]" captionTargetId="figure-347@30.[839,1466,1510,1920]" captionTargetPageId="30" captionText="Figure 45.Genitalia of Amplirhagadaanderdonensis n.sp., paratype FMNH 219330 (22 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." figureDoi="http://doi.org/10.5281/zenodo.5239179" httpUri="https://zenodo.org/record/5239179/files/figure.png" pageId="31" pageNumber="248">Figs. 45</figureCitation>
,
<figureCitation id="136B2A56FF8EF74306E317E649A2C9CE" box="[1174,1207,1955,1978]" captionStart="Figure 47" captionStartId="32.[426,489,350,371]" captionTargetBox="[181,794,161,958]" captionTargetId="figure-8@32.[167,794,151,964]" captionTargetPageId="32" captionText="Figure 47. Interior of penial chamber of Amplirhagada anderdonensis n.sp., paratype FMNH 219330 (22 July, scale 5 mm). Compare with Fig. 4 for labelling of structures." figureDoi="http://doi.org/10.5281/zenodo.5239183" httpUri="https://zenodo.org/record/5239183/files/figure.png" pageId="31" pageNumber="248">47</figureCitation>
). Penis straight to slightly curved, more or less of same length as anterior part of oviduct. Vas deferens coils once before entering penis. Penial retractor muscle shorter than penis. Penial verge moderately long (
<emphasis id="B924EAC1FFB1F77C032814524C70CA51" bold="true" box="[349,357,1047,1061]" pageId="32" pageNumber="249">
<superScript id="7C259B9BFFB1F77C032814524C70CA51" attach="none" box="[349,357,1047,1061]" fontSize="6" pageId="32" pageNumber="249">1</superScript>
</emphasis>
<emphasis id="B924EAC1FFB1F77C031F145B4C92CA51" bold="true" box="[362,391,1047,1070]" pageId="32" pageNumber="249">
<subScript id="17D43496FFB1F77C031F145B4C67CA58" attach="none" box="[362,370,1054,1068]" fontSize="6" pageId="32" pageNumber="249">8</subScript>
<superScript id="7C259B9BFFB1F77C030A14524C92CA51" attach="none" box="[383,391,1047,1061]" fontSize="6" pageId="32" pageNumber="249">1</superScript>
</emphasis>
<emphasis id="B924EAC1FFB1F77C03F9145B4C81CA58" bold="true" box="[396,404,1054,1068]" pageId="32" pageNumber="249">
<subScript id="17D43496FFB1F77C03F9145B4C81CA58" attach="left" box="[396,404,1054,1068]" fontSize="6" pageId="32" pageNumber="249">4</subScript>
</emphasis>
penial chamber), slender to spatulate, with pointed tip. Penial wall pustules small, arranged in rows. Main stimulatory pilaster differentiated, relatively thin, cone-shaped, with horizontal ridged that support little hooks, comprising apical to median portion of penial chamber. Vas deferens entering penial sheath in upper third. Vagina moderately long, tubular, posteriorly inflated; inner vaginal wall with smooth longitudinal pilasters. Spermatheca very short, barely reaching base of spermoviduct. Spermathecal duct normal, internally with smooth longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with delicate wall. Free oviduct comprising more than half of anterior part of oviduct, more or less straight. Spermoviduct longer than anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<caption id="DF2F665BFFB1F77C03DF111B4F97CFA9" ID-DOI="http://doi.org/10.5281/zenodo.5239183" ID-Zenodo-Dep="5239183" httpUri="https://zenodo.org/record/5239183/files/figure.png" pageId="32" pageNumber="249" startId="32.[426,489,350,371]" targetBox="[181,794,161,958]" targetPageId="32">
<paragraph id="8BEF36D3FFB1F77C03DF111B4F97CFA9" blockId="32.[426,794,350,477]" pageId="32" pageNumber="249">
Figure 47. Interior of penial chamber of
<taxonomicName id="4C504D50FFB1F77C03B1113D4FF5CFF9" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[452,736,376,397]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="anderdonensis" status="sp. nov.">
<emphasis id="B924EAC1FFB1F77C03B1113D4FF5CFF9" box="[452,736,376,397]" italics="true" pageId="32" pageNumber="249">Amplirhagada anderdonensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFB1F77C0093113D4E03CFF9" box="[742,790,376,397]" pageId="32" pageNumber="249" rank="species">n.sp.</taxonomicNameLabel>
, paratype FMNH 219330 (22 July, scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<paragraph id="8BEF36D3FFB1F77C02D3164F4F5DC978" blockId="32.[166,794,1546,1804]" pageId="32" pageNumber="249">
<emphasis id="B924EAC1FFB1F77C02D3164F4C8EC856" bold="true" box="[166,411,1546,1570]" pageId="32" pageNumber="249">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFB1F77C03DB164F4F20C855" box="[430,565,1546,1569]" captionStart="Figure 46" captionStartId="31.[121,184,1555,1576]" captionTargetBox="[121,1421,151,1540]" captionTargetId="figure-9@31.[121,1421,151,1540]" captionTargetPageId="31" captionText="Figure 46. SEM photographs of Amplirhagada anderdonensis n.sp. (AC) Shell, paratype AMS C463756: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on second and third whorl viewed from above (scale 200 µm); (C) sculpture on upper part of last whorl (scale 100 µm). (D) Jaw (scale 100 µm). (EG) Radula, paratype FMNH 219330: (E) central and lateral teeth (scale 20 µm); (F) close-up of lateral teeth (scale 10 µm); (G) outer lateral and inner marginal teeth (scale 10 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239181" httpUri="https://zenodo.org/record/5239181/files/figure.png" pageId="32" pageNumber="249">Fig. 46EG</figureCitation>
). Rectangular. Tooth formula C+1415+2+2023. In average with 145±12.0 rows of teeth, 31.6±4.2 rows per mm (n = 2). Central teeth with bluntly pointed, ovate mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with bluntly pointed, ovate mesocones, length equal to base of tooth; ectocones small, endocones vestigial. Marginal teeth with triangular to rounded mesocones; ectocones shorter and narrower than mesocones; endocones reduced in size.
</paragraph>
<paragraph id="8BEF36D3FFB1F77C02D3176A4FD6C9AD" blockId="32.[166,794,1839,2009]" pageId="32" pageNumber="249">
<emphasis id="B924EAC1FFB1F77C02D3176A4C8EC933" bold="true" box="[166,411,1839,1863]" pageId="32" pageNumber="249">Comparative remarks</emphasis>
. Shell colour very distinctive, similar only to
<taxonomicName id="4C504D50FFB1F77C037717084C81C910" authorityName="Köhler" authorityYear="2010" box="[258,404,1869,1892]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="tricenaria">
<emphasis id="B924EAC1FFB1F77C037717084C81C910" box="[258,404,1869,1892]" italics="true" pageId="32" pageNumber="249">A. tricenaria</emphasis>
</taxonomicName>
(see below). Penis proportionally smaller than in other species. In addition, it differs from
<taxonomicName id="4C504D50FFB1F77C0176172F4C3EC9EB" authorityName="Köhler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="lamarckiana">
<emphasis id="B924EAC1FFB1F77C0176172F4C3EC9EB" italics="true" pageId="32" pageNumber="249">A. lamarckiana</emphasis>
</taxonomicName>
by finer pustulation of inner penial wall and smaller main pilaster. Material of the present species has been labelled as “
<taxonomicName id="4C504D50FFB1F77C031617874F37C9AD" box="[355,546,1986,2009]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="249" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FFB1F77C031617874CE8C9AD" box="[355,509,1986,2009]" italics="true" pageId="32" pageNumber="249">Amplirhagada</emphasis>
sp.
</taxonomicName>
75” by Solem.
</paragraph>
</subSubSection>
</treatment>
</document>