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<document id="CF90EE98EA9455896EC2684F6FE5364A" ID-CLB-Dataset="28135" ID-DOI="10.11646/zootaxa.4563.3.3" ID-GBIF-Dataset="6c280343-607d-4ff5-b2ba-1ed24db90c36" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601342" ID-ZooBank="86E75433-D91B-48E5-9807-9A0A460EFCB3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553172235077" checkinUser="plazi" docAuthor="Salgado, Alexandre &amp; Ruiz, Gustavo R. S." docDate="2019" docId="03F987C08A6501683AEE8C24FC3B4D36" docLanguage="en" docName="zootaxa.4563.3.3.pdf" docOrigin="Zootaxa 4563 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphidraus draconicaudatus Salgado &amp; Ruiz 2017" docType="treatment" docVersion="8" lastPageNumber="477" masterDocId="FFC0FFB88A7201723A798B0DFFC84C76" masterDocTitle="Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini)" masterLastPageNumber="481" masterPageNumber="451" pageNumber="474" updateTime="1698703444826" updateUser="ExternalLinkService">
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<mods:title id="016A03F7A4B2BBCA4912229E75CFA58D">Seven new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini)</mods:title>
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<mods:namePart id="FBE77062965E52470553142384E6ADAF">Salgado, Alexandre</mods:namePart>
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<mods:namePart id="CB6EB3E3A636390F5D6069F94A1183C5">Ruiz, Gustavo R. S.</mods:namePart>
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<mods:title id="90285AB4FD0F859ED6A749082B6FC819">Zootaxa</mods:title>
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<treatment id="03F987C08A6501683AEE8C24FC3B4D36" ID-DOI="http://doi.org/10.5281/zenodo.5931679" ID-GBIF-Taxon="154474856" ID-Zenodo-Dep="5931679" LSID="urn:lsid:plazi:treatment:03F987C08A6501683AEE8C24FC3B4D36" httpUri="http://treatment.plazi.org/id/03F987C08A6501683AEE8C24FC3B4D36" lastPageId="26" lastPageNumber="477" pageId="23" pageNumber="474">
<subSubSection id="C34A655D8A6501653AEE8C24FCDC4B32" box="[151,788,1833,1860]" pageId="23" pageNumber="474" type="nomenclature">
<paragraph id="8BEF36D68A6501653AEE8C24FCDC4B32" blockId="23.[151,788,1833,1893]" box="[151,788,1833,1860]" pageId="23" pageNumber="474">
<heading id="D0A781BA8A6501653AEE8C24FCDC4B32" bold="true" box="[151,788,1833,1860]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
<taxonomicName id="4C504D558A6501653AEE8C24FCDC4B32" ID-CoL="D2G8" authority="Salgado &amp; Ruiz, 2017" authorityName="Salgado &amp; Ruiz" authorityYear="2017" box="[151,788,1833,1860]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="draconicaudatus">
<emphasis id="B924EAC48A6501653AEE8C24FCDC4B32" bold="true" box="[151,788,1833,1860]" pageId="23" pageNumber="474">
<emphasis id="B924EAC48A6501653AEE8C24FE364B35" bold="true" box="[151,510,1833,1859]" italics="true" pageId="23" pageNumber="474">Amphidraus draconicaudatus</emphasis>
<bibRefCitation id="EFC14B278A650165387C8C27FCDC4B32" author="Salgado, A. &amp; Ruiz, G. R." box="[517,788,1834,1860]" pageId="23" pageNumber="474" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz, 2017</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A6501653AEE8C41FD484B13" box="[151,640,1868,1893]" pageId="23" pageNumber="474" type="description">
<paragraph id="8BEF36D68A6501653AEE8C41FD484B13" blockId="23.[151,788,1833,1893]" box="[151,640,1868,1893]" pageId="23" pageNumber="474">
<figureCitation id="136B2A538A6501653AEE8C41FF224B13" box="[151,234,1868,1893]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="23" pageNumber="474">Figs 26</figureCitation>
AD, 26G, 27AE, 28A, 29AB, 30
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A6501653AEE8C99FB9F4B9E" pageId="23" pageNumber="474" type="reference_group">
<paragraph id="8BEF36D68A6501653AEE8C99FB9F4B9E" blockId="23.[151,1436,1940,2024]" pageId="23" pageNumber="474">
<treatmentCitationGroup id="AB4011F88A6501653AEE8C99FB9F4B9E" infered="true" pageId="23" pageNumber="474">
<taxonomicName id="4C504D558A6501653AEE8C99FD224BDC" ID-CoL="D2G8" authority="Salgado &amp; Ruiz, 2017: 427" authorityName="Salgado &amp; Ruiz" authorityPageNumber="427" authorityYear="2017" box="[151,746,1940,1962]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="draconicaudatus">
<emphasis id="B924EAC48A6501653AEE8C99FE0D4BDF" box="[151,453,1940,1961]" italics="true" pageId="23" pageNumber="474">Amphidraus draconicaudatus</emphasis>
<treatmentCitation id="0AF110C78A6501653BB48C99FD224BDC" author="Salgado, A. &amp; Ruiz, G. R." box="[461,746,1940,1962]" page="427" pageId="23" pageNumber="474" year="2017">
<bibRefCitation id="EFC14B278A6501653BB48C99FD7F4BDC" author="Salgado, A. &amp; Ruiz, G. R." box="[461,695,1940,1962]" pageId="23" pageNumber="474" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz, 2017</bibRefCitation>
: 427
</treatmentCitation>
</taxonomicName>
, figs 2224 (
<typeStatus id="54EB88748A650165390A8C99FC1B4BDC" box="[883,979,1940,1962]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
<specimenCount id="9D56FD5F8A65016539A28C98FC264BDC" box="[987,1006,1941,1962]" pageId="23" pageNumber="474" type="male"></specimenCount>
from Pium,
<geoCoordinate id="EE6450118A6501653E0A8C99FB344BDC" box="[1139,1276,1940,1962]" direction="south" orientation="latitude" pageId="23" pageNumber="474" precision="1" value="-9.977416">09°58'38.7&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6450118A6501653F718C99FA504BDC" box="[1288,1432,1940,1962]" direction="west" orientation="longitude" pageId="23" pageNumber="474" precision="1" value="-50.03472">50°02'05.0&quot;W</geoCoordinate>
,
<collectingRegion id="4994F8348A6501653ABE8CBEFEE44BBF" box="[199,300,1971,1993]" country="Brazil" name="Tocantins" pageId="23" pageNumber="474">Tocantins</collectingRegion>
,
<collectingCountry id="F34776468A6501653B438CBEFEB24BBF" box="[314,378,1971,1993]" name="Brazil" pageId="23" pageNumber="474">Brazil</collectingCountry>
,
<date id="FFEE10168A6501653BF08CBEFDD04BBF" box="[393,536,1971,1993]" pageId="23" pageNumber="474" value="2016-05-11" valueMax="2016-05-26" valueMin="2016-05-11">1126.V.2016</date>
, Serrão et al. leg., deposided in MPEG 32689;
<specimenCount id="9D56FD5F8A6501653E658CBEFBF44BBF" box="[1052,1084,1971,1993]" pageId="23" pageNumber="474" type="male">1♂</specimenCount>
and
<specimenCount id="9D56FD5F8A6501653E0F8CBEFB5E4BBF" box="[1142,1174,1971,1993]" pageId="23" pageNumber="474" type="female">2♀</specimenCount>
<typeStatus id="54EB88748A6501653ED88CB9FACD4BBF" box="[1185,1285,1972,1993]" pageId="23" pageNumber="474" type="paratype">paratypes</typeStatus>
, same data as
<typeStatus id="54EB88748A6501653ABE8CDFFEEA4B9E" box="[199,290,2002,2024]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
, deposited in IBSP 211843211844, MPEG 32690);
<bibRefCitation id="EFC14B278A65016539438CDFFB9A4B9E" author="World Spider Catalog" box="[826,1106,2002,2024]" pageId="23" pageNumber="474" refId="ref9034" refString="World Spider Catalog (2018) World Spider Catalog. Uersion 18.0. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed 2 May 2018)" type="url" year="2018">World Spider Catalog 2018</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF2F665E8A6A016A3AEE8DE7FD7D4B0C" httpUri="https://zenodo.org/record/2601392/files/figure.png" pageId="24" pageNumber="475" startId="24.[151,250,1770,1792]" targetBox="[151,1432,204,1738]" targetPageId="24">
<paragraph id="8BEF36D68A6A016A3AEE8DE7FD7D4B0C" blockId="24.[151,1436,1770,1914]" pageId="24" pageNumber="475">
<emphasis id="B924EAC48A6A016A3AEE8DE7FEE94A89" bold="true" box="[151,289,1770,1792]" pageId="24" pageNumber="475">FIGURE 23.</emphasis>
<taxonomicName id="4C504D558A6A016A3B508DE7FDFE4A89" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[297,566,1770,1791]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="draconitupan" status="sp. nov.">
<emphasis id="B924EAC48A6A016A3B508DE7FDFE4A89" box="[297,566,1770,1791]" italics="true" pageId="24" pageNumber="475">Amphidraus draconitupan</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6A016A38448DE7FD474A89" bold="true" box="[573,655,1770,1791]" pageId="24" pageNumber="475">
<taxonomicNameLabel id="A21757BF8A6A016A38448DE7FD474A89" box="[573,655,1770,1791]" pageId="24" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AC male (A frontal, B dorsal, C ventral); DG left male palp (D embolic disc, ventral; E ventral; F prolateral; G retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc. Black arrows in D show reduced spikes on elongate projection of embolic disc. White arrow in E shows rounded projection on the elongate projection of embolic disc.
</paragraph>
</caption>
<caption id="DF2F665E8A6B016B3AEE8F80FE3148B4" httpUri="https://zenodo.org/record/2601394/files/figure.png" pageId="25" pageNumber="476" startId="25.[151,250,1165,1187]" targetBox="[151,1436,185,1152]" targetPageId="25">
<paragraph id="8BEF36D68A6B016B3AEE8F80FE3148B4" blockId="25.[151,1437,1165,1218]" pageId="25" pageNumber="476">
<emphasis id="B924EAC48A6B016B3AEE8F80FED748D5" bold="true" box="[151,287,1165,1187]" pageId="25" pageNumber="476">FIGURE 24.</emphasis>
<taxonomicName id="4C504D558A6B016B3B5C8F83FDE748D5" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[293,559,1166,1187]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconitupan" status="sp. nov.">
<emphasis id="B924EAC48A6B016B3B5C8F83FDE748D5" box="[293,559,1166,1187]" italics="true" pageId="25" pageNumber="476">Amphidraus draconitupan</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B384C8F83FD4D48D5" bold="true" box="[565,645,1166,1187]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B384C8F83FD4D48D5" box="[565,645,1166,1187]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AC female (A dorsal, B ventral, C live specimen); D epigyne/vulva, ventral. Abbreviation: po, coupling pocket.
</paragraph>
</caption>
<subSubSection id="C34A655D8A6B016B3AEE8FE5FE84493F" pageId="25" pageNumber="476" type="materials_examined">
<paragraph id="8BEF36D68A6B016B3AEE8FE5FE84493F" blockId="25.[151,1437,1256,2037]" pageId="25" pageNumber="476">
<emphasis id="B924EAC48A6B016B3AEE8FE5FCCB4977" bold="true" box="[151,771,1256,1281]" pageId="25" pageNumber="476">
Additional material examined.
<materialsCitation id="3B383C8B8A6B016B38748FE5FCCB4977" ID-GBIF-Occurrence="2013952931" box="[525,771,1256,1281]" collectingDate="2016-05-11" collectionCode="MPEG" collectorName="A. B. Bonaldo" country="Brazil" latitude="-9.304334" location="Caseara" longLatPrecision="1" longitude="-49.95789" pageId="25" pageNumber="476" specimenCode="MPEG 35018" specimenCount="2" specimenCount-male="2" stateProvince="Tocantins">
<collectingCountry id="F34776468A6B016B38748FE5FDB54977" box="[525,637,1256,1281]" name="Brazil" pageId="25" pageNumber="476">BRAZIL</collectingCountry>
:
<emphasis id="B924EAC48A6B016B38F08FE5FD324977" bold="true" box="[649,762,1256,1281]" italics="true" pageId="25" pageNumber="476">
<collectingRegion id="4994F8348A6B016B38F08FE5FD324977" box="[649,762,1256,1281]" country="Brazil" name="Tocantins" pageId="25" pageNumber="476">Tocantins</collectingRegion>
</emphasis>
:
</materialsCitation>
</emphasis>
<location id="8E8F600D8A6B016B39728FE4FCAF4977" LSID="urn:lsid:plazi:treatment:03F987C08A6501683AEE8C24FC3B4D36:8E8F600D8A6B016B39728FE4FCAF4977" box="[779,871,1257,1281]" country="Brazil" latitude="-9.304334" longLatPrecision="1" longitude="-49.95789" name="Caseara" pageId="25" pageNumber="476" stateProvince="Tocantins">Caseara</location>
,
<geoCoordinate id="EE6450118A6B016B390B8FE5FC334977" box="[882,1019,1256,1281]" direction="south" orientation="latitude" pageId="25" pageNumber="476" precision="1" value="-9.304334">9°18'15.6&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6450118A6B016B3E7E8FE5FB614977" box="[1031,1193,1256,1281]" direction="west" orientation="longitude" pageId="25" pageNumber="476" precision="1" value="-49.95789">49°57'28.4&quot;W</geoCoordinate>
:
<specimenCount id="9D56FD5F8A6B016B3ECC8FE4FB124977" box="[1205,1242,1257,1281]" pageId="25" pageNumber="476" type="male">2♂</specimenCount>
,
<date id="FFEE10168A6B016B3E9E8FE5FA934977" box="[1255,1371,1256,1281]" pageId="25" pageNumber="476" value="2016-05-11">
<collectingDate id="EFAAE9FE8A6B016B3E9E8FE5FA934977" box="[1255,1371,1256,1281]" pageId="25" pageNumber="476" value="2016-05-11">11.V.2016</collectingDate>
</date>
,
<collectorName id="26A553008A6B016B3F1E8FE4FF3E4953" pageId="25" pageNumber="476">A.B. Bonaldo</collectorName>
<emphasis id="B924EAC48A6B016B3A868E00FEF14952" box="[255,313,1292,1316]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
leg. (
<specimenCode id="DBF69EAD8A6B016B3B058E00FDD34953" box="[380,539,1292,1317]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="25" pageNumber="476">MPEG 35018</specimenCode>
);
<materialsCitation id="3B383C8B8A6B016B384C8E01FE80493F" ID-GBIF-Occurrence="2013952928" collectingDate="2016-05-19" collectionCode="MPEG" collectorName="A. B. Bonaldo" country="Brazil" latitude="-9.977416" location="Pium" longLatPrecision="1" longitude="-50.03472" pageId="25" pageNumber="476" specimenCode="MPEG 35019" specimenCount="1" specimenCount-male="1" stateProvince="Tocantins">
<location id="8E8F600D8A6B016B384C8E01FDBB4952" LSID="urn:lsid:plazi:treatment:03F987C08A6501683AEE8C24FC3B4D36:8E8F600D8A6B016B384C8E01FDBB4952" box="[565,627,1292,1316]" country="Brazil" latitude="-9.977416" longLatPrecision="1" longitude="-50.03472" name="Pium" pageId="25" pageNumber="476" stateProvince="Tocantins">Pium</location>
,
<geoCoordinate id="EE6450118A6B016B38F98E01FCDF4952" box="[640,791,1292,1316]" direction="south" orientation="latitude" pageId="25" pageNumber="476" precision="1" value="-9.977416">09°58'38.7&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6450118A6B016B395D8E01FC0F4952" box="[804,967,1292,1316]" direction="west" orientation="longitude" pageId="25" pageNumber="476" precision="1" value="-50.03472">50°02'05.0&quot;W</geoCoordinate>
:
<specimenCount id="9D56FD5F8A6B016B39AD8E01FC304952" box="[980,1016,1292,1316]" pageId="25" pageNumber="476" type="male">1♂</specimenCount>
,
<date id="FFEE10168A6B016B3E7E8E01FBB44952" box="[1031,1148,1292,1316]" pageId="25" pageNumber="476" value="2016-05-19">
<collectingDate id="EFAAE9FE8A6B016B3E7E8E01FBB44952" box="[1031,1148,1292,1316]" pageId="25" pageNumber="476" value="2016-05-19">19.V.2016</collectingDate>
</date>
,
<collectorName id="26A553008A6B016B3EF08E01FAEE4952" box="[1161,1318,1292,1316]" pageId="25" pageNumber="476">A.B. Bonaldo</collectorName>
<emphasis id="B924EAC48A6B016B3F568E00FAA24952" box="[1327,1386,1292,1316]" italics="true" pageId="25" pageNumber="476">et al.</emphasis>
leg. (
<specimenCode id="DBF69EAD8A6B016B3AE78E3CFE88493F" box="[158,320,1328,1353]" collectionCode="MPEG" country="Brazil" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="25" pageNumber="476">MPEG 35019</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A6B016B3ABE8E59FB6C4B6B" pageId="25" pageNumber="476" type="diagnosis">
<paragraph id="8BEF36D68A6B016B3ABE8E59FB6C4B6B" blockId="25.[151,1437,1256,2037]" pageId="25" pageNumber="476">
<emphasis id="B924EAC48A6B016B3ABE8E59FE6C491B" bold="true" box="[199,420,1364,1389]" pageId="25" pageNumber="476">Revised diagnosis.</emphasis>
<taxonomicName id="4C504D558A6B016B3BD78E58FD45491A" authorityName="Salgado &amp; Ruiz" authorityYear="2017" box="[430,653,1364,1388]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconicaudatus">
<emphasis id="B924EAC48A6B016B3BD78E58FD45491A" box="[430,653,1364,1388]" italics="true" pageId="25" pageNumber="476">A. draconicaudatus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C504D558A6B016B39548E58FC7D491A" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[813,949,1364,1388]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="shenlong" status="sp. nov.">
<emphasis id="B924EAC48A6B016B39548E58FC7D491A" box="[813,949,1364,1388]" italics="true" pageId="25" pageNumber="476">A. shenlong</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B39B98E59FBD5491B" bold="true" box="[960,1053,1364,1389]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B39B98E59FBD5491B" box="[960,1053,1364,1389]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having an elongate and stout projection on the retrolateral border of the embolic disc that extends towards the retrolateral tegulum border and bears several branches (
<figureCitation id="136B2A538A6B016B3BDC8E91FDD949C3" box="[421,529,1436,1461]" captionStart="FIGURE 21" captionStartId="22.[151,250,1923,1945]" captionTargetBox="[159,1422,795,1903]" captionTargetId="figure@22.[159,1423,795,1903]" captionTargetPageId="22" captionText="FIGURE 21. Amphidraus shenlong sp. nov. AB male (A dorsal, B ventral); CF left male palp (C ventral; D retroventral; E bulb, ventral; F femur + patella + tibia, retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. White arrow in D shows branchy projection of embolic disc." httpUri="https://zenodo.org/record/2601388/files/figure.png" pageId="25" pageNumber="476">Figs 21D</figureCitation>
,
<figureCitation id="136B2A538A6B016B38678E91FD4649C3" box="[542,654,1436,1461]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">26A, 26C</figureCitation>
,
<figureCitation id="136B2A538A6B016B38E28E90FD0749C3" box="[667,719,1437,1461]" captionStart="FIGURE 27" captionStartId="28.[151,250,1131,1153]" captionTargetBox="[151,1436,228,1080]" captionTargetId="figure@28.[151,1436,228,1080]" captionTargetPageId="28" captionText="FIGURE 27. Copulatory structures of Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AD left male palp (A embolic disc, ventral; patella + tibia, retrolateral; C detail of RvTA projection; D detail of reduced RTA). E epigyne/vulva, dorsal. Abbreviations: dPED, distal processes on embolic disc; e, embolus shaft; ef, embolic filament; po, coupling pocket; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc. White arrows in B and C show projection on RvTA, and black arrow in C shows one spike of the reduced RTA." httpUri="https://zenodo.org/record/2601400/files/figure.png" pageId="25" pageNumber="476">27A</figureCitation>
); also, by having the RvTA pointing ventrally, with projections on its border (
<figureCitation id="136B2A538A6B016B3B4E8ECDFE5549AF" box="[311,413,1472,1497]" captionStart="FIGURE 21" captionStartId="22.[151,250,1923,1945]" captionTargetBox="[159,1422,795,1903]" captionTargetId="figure@22.[159,1423,795,1903]" captionTargetPageId="22" captionText="FIGURE 21. Amphidraus shenlong sp. nov. AB male (A dorsal, B ventral); CF left male palp (C ventral; D retroventral; E bulb, ventral; F femur + patella + tibia, retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. White arrow in D shows branchy projection of embolic disc." httpUri="https://zenodo.org/record/2601388/files/figure.png" pageId="25" pageNumber="476">Figs 21F</figureCitation>
,
<figureCitation id="136B2A538A6B016B3BD08ECCFE1349AF" box="[425,475,1473,1497]" captionStart="FIGURE 22" captionStartId="23.[151,250,1670,1692]" captionTargetBox="[177,1383,890,1615]" captionTargetId="figure@23.[151,1436,879,1648]" captionTargetPageId="23" captionText="FIGURE 22. Amphidraus shenlong sp. nov., left male palp (A ventral; B retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal processes on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601390/files/figure.png" pageId="25" pageNumber="476">22B</figureCitation>
,
<figureCitation id="136B2A538A6B016B3B9E8ECDFDD449AF" box="[487,540,1472,1497]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">26G</figureCitation>
). However, the male of
<taxonomicName id="4C504D558A6B016B39548ECCFBC149AE" authorityName="Salgado &amp; Ruiz" authorityYear="2017" box="[813,1033,1472,1496]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconicaudatus">
<emphasis id="B924EAC48A6B016B39548ECCFBC149AE" box="[813,1033,1472,1496]" italics="true" pageId="25" pageNumber="476">A. draconicaudatus</emphasis>
</taxonomicName>
can be distinguished by having the RvTA with a single serrated projection, and a RTA reduced to spikes (
<figureCitation id="136B2A538A6B016B39CC8EE9FBD7498B" box="[949,1055,1508,1533]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">Figs 26G</figureCitation>
,
<figureCitation id="136B2A538A6B016B3E528EE8FB80498B" box="[1067,1096,1509,1533]" captionStart="FIGURE 27" captionStartId="28.[151,250,1131,1153]" captionTargetBox="[151,1436,228,1080]" captionTargetId="figure@28.[151,1436,228,1080]" captionTargetPageId="28" captionText="FIGURE 27. Copulatory structures of Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AD left male palp (A embolic disc, ventral; patella + tibia, retrolateral; C detail of RvTA projection; D detail of reduced RTA). E epigyne/vulva, dorsal. Abbreviations: dPED, distal processes on embolic disc; e, embolus shaft; ef, embolic filament; po, coupling pocket; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc. White arrows in B and C show projection on RvTA, and black arrow in C shows one spike of the reduced RTA." httpUri="https://zenodo.org/record/2601400/files/figure.png" pageId="25" pageNumber="476">27</figureCitation>
BD), whereas in
<taxonomicName id="4C504D558A6B016B3F6F8EE8FA54498A" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1302,1436,1508,1532]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="shenlong" status="sp. nov.">
<emphasis id="B924EAC48A6B016B3F6F8EE8FA54498A" box="[1302,1436,1508,1532]" italics="true" pageId="25" pageNumber="476">A. shenlong</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3AEE8D05FF3A4A57" bold="true" box="[151,242,1544,1569]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3AEE8D05FF3A4A57" box="[151,242,1544,1569]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the RvTA has four projections (three with acute tip and one with serrated tip), and the RTA serrated tip reaches the same height of the RvTA tip along palp axis (
<figureCitation id="136B2A538A6B016B39658D21FC494A33" box="[796,897,1580,1605]" captionStart="FIGURE 21" captionStartId="22.[151,250,1923,1945]" captionTargetBox="[159,1422,795,1903]" captionTargetId="figure@22.[159,1423,795,1903]" captionTargetPageId="22" captionText="FIGURE 21. Amphidraus shenlong sp. nov. AB male (A dorsal, B ventral); CF left male palp (C ventral; D retroventral; E bulb, ventral; F femur + patella + tibia, retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc. White arrow in D shows branchy projection of embolic disc." httpUri="https://zenodo.org/record/2601388/files/figure.png" pageId="25" pageNumber="476">Figs 21F</figureCitation>
,
<figureCitation id="136B2A538A6B016B39F58D20FC764A33" box="[908,958,1581,1605]" captionStart="FIGURE 22" captionStartId="23.[151,250,1670,1692]" captionTargetBox="[177,1383,890,1615]" captionTargetId="figure@23.[151,1436,879,1648]" captionTargetPageId="23" captionText="FIGURE 22. Amphidraus shenlong sp. nov., left male palp (A ventral; B retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal processes on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601390/files/figure.png" pageId="25" pageNumber="476">22B</figureCitation>
,
<figureCitation id="136B2A538A6B016B39B08D21FC364A33" box="[969,1022,1580,1605]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">26H</figureCitation>
). The female of
<taxonomicName id="4C504D558A6B016B3ECF8D20FAF34A32" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[1206,1339,1580,1604]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="shenlong" status="sp. nov.">
<emphasis id="B924EAC48A6B016B3ECF8D20FAF34A32" box="[1206,1339,1580,1604]" italics="true" pageId="25" pageNumber="476">A. shenlong</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3F3B8D21FA544A33" bold="true" box="[1346,1436,1580,1605]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3F3B8D21FA544A33" box="[1346,1436,1580,1605]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unknown, but among the currently known females of
<taxonomicName id="4C504D558A6B016B397C8D5DFC464A1E" authorityName="Simon" authorityYear="1900" box="[773,910,1616,1640]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="B924EAC48A6B016B397C8D5DFC464A1E" box="[773,910,1616,1640]" italics="true" pageId="25" pageNumber="476">Amphidraus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D558A6B016B39E28D5CFBBD4A1E" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[923,1141,1616,1640]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconicaudatus" status="sp. nov.">
<emphasis id="B924EAC48A6B016B39E28D5CFBBD4A1E" box="[923,1141,1616,1640]" italics="true" pageId="25" pageNumber="476">A. draconicaudatus</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3E058D5DFB1D4A1F" bold="true" box="[1148,1237,1616,1641]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3E058D5DFB1D4A1F" box="[1148,1237,1616,1641]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to those of
<taxonomicName id="4C504D558A6B016B3A828D78FE7B4AFA" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[251,435,1652,1676]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconitupan" status="sp. nov.">
<emphasis id="B924EAC48A6B016B3A828D78FE7B4AFA" box="[251,435,1652,1676]" italics="true" pageId="25" pageNumber="476">A. draconitupan</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3BC58D79FDDF4AFB" bold="true" box="[444,535,1652,1677]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3BC58D79FDDF4AFB" box="[444,535,1652,1677]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, both with spermathecae kidney-shaped, with glands near copulatory openings and with a narrow dorsal pocket placed posteriorly on epigynal plate (
<figureCitation id="136B2A538A6B016B39E68D95FBC24AC7" box="[927,1034,1688,1713]" captionStart="FIGURE 24" captionStartId="25.[151,250,1165,1187]" captionTargetBox="[151,1436,185,1152]" captionTargetId="figure@25.[151,1436,185,1152]" captionTargetPageId="25" captionText="FIGURE 24. Amphidraus draconitupan sp. nov. AC female (A dorsal, B ventral, C live specimen); D epigyne/vulva, ventral. Abbreviation: po, coupling pocket." httpUri="https://zenodo.org/record/2601394/files/figure.png" pageId="25" pageNumber="476">Figs 24D</figureCitation>
,
<figureCitation id="136B2A538A6B016B3E6C8D95FB8F4AC7" box="[1045,1095,1688,1713]" captionStart="FIGURE 25" captionStartId="26.[151,250,1787,1809]" captionTargetBox="[167,1392,393,1736]" captionTargetId="figure@26.[151,1436,370,1755]" captionTargetPageId="26" captionText="FIGURE 25. Amphidraus draconitupan sp. nov., AC left male palp (A ventral, B prolateral, C retrolateral); D epigyne/vulva, cleared, ventral. Abbreviations: cc, cymbial conductor; co, copulatory opening; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; po, coupling pocket; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601396/files/figure.png" pageId="25" pageNumber="476">25D</figureCitation>
,
<figureCitation id="136B2A538A6B016B3E2A8D94FB4C4AC7" box="[1107,1156,1689,1713]" captionStart="FIGURE 27" captionStartId="28.[151,250,1131,1153]" captionTargetBox="[151,1436,228,1080]" captionTargetId="figure@28.[151,1436,228,1080]" captionTargetPageId="28" captionText="FIGURE 27. Copulatory structures of Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AD left male palp (A embolic disc, ventral; patella + tibia, retrolateral; C detail of RvTA projection; D detail of reduced RTA). E epigyne/vulva, dorsal. Abbreviations: dPED, distal processes on embolic disc; e, embolus shaft; ef, embolic filament; po, coupling pocket; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc. White arrows in B and C show projection on RvTA, and black arrow in C shows one spike of the reduced RTA." httpUri="https://zenodo.org/record/2601400/files/figure.png" pageId="25" pageNumber="476">27E</figureCitation>
). However, the female of
<taxonomicName id="4C504D558A6B016B3AEE8DB0FEB14AA2" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[151,377,1724,1748]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconicaudatus" status="sp. nov.">
<emphasis id="B924EAC48A6B016B3AEE8DB0FEB14AA2" box="[151,377,1724,1748]" italics="true" pageId="25" pageNumber="476">A. draconicaudatus</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3BFE8DB1FE2F4AA3" bold="true" box="[391,487,1724,1749]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3BFE8DB1FE2F4AA3" box="[391,487,1724,1749]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from that of
<taxonomicName id="4C504D558A6B016B39EE8DB0FB9C4AA2" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[919,1108,1724,1748]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconitupan" status="sp. nov.">
<emphasis id="B924EAC48A6B016B39EE8DB0FB9C4AA2" box="[919,1108,1724,1748]" italics="true" pageId="25" pageNumber="476">A. draconitupan</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6B016B3E1B8DB1FB0B4AA3" bold="true" box="[1122,1219,1724,1749]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21757BF8A6B016B3E1B8DB1FB0B4AA3" box="[1122,1219,1724,1749]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having longer copulatory ducts connected to the inner posterior border of spermathecae (see
<bibRefCitation id="EFC14B278A6B016B3E5D8DEDFAEB4A8F" author="Salgado, A. &amp; Ruiz, G. R." box="[1060,1315,1760,1785]" pageId="25" pageNumber="476" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: fig. 24F), whereas they connect to the outer border of the spermathecae in
<taxonomicName id="4C504D558A6B016B39108C08FBD74B6A" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[873,1055,1796,1820]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="draconitupan">
<emphasis id="B924EAC48A6B016B39108C08FBD74B6A" box="[873,1055,1796,1820]" italics="true" pageId="25" pageNumber="476">A. draconitupan</emphasis>
</taxonomicName>
(
<figureCitation id="136B2A538A6B016B3E578C09FB5F4B6B" box="[1070,1175,1796,1821]" captionStart="FIGURE 25" captionStartId="26.[151,250,1787,1809]" captionTargetBox="[167,1392,393,1736]" captionTargetId="figure@26.[151,1436,370,1755]" captionTargetPageId="26" captionText="FIGURE 25. Amphidraus draconitupan sp. nov., AC left male palp (A ventral, B prolateral, C retrolateral); D epigyne/vulva, cleared, ventral. Abbreviations: cc, cymbial conductor; co, copulatory opening; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; po, coupling pocket; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601396/files/figure.png" pageId="25" pageNumber="476">Fig. 25D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A6B01683ABE8C25FAA34D6A" lastPageId="26" lastPageNumber="477" pageId="25" pageNumber="476" type="description">
<paragraph id="8BEF36D68A6B016B3ABE8C25FD534B37" blockId="25.[151,1437,1256,2037]" box="[199,667,1832,1857]" pageId="25" pageNumber="476">
<emphasis id="B924EAC48A6B016B3ABE8C25FE914B37" bold="true" box="[199,345,1832,1857]" pageId="25" pageNumber="476">Description.</emphasis>
See
<bibRefCitation id="EFC14B278A6B016B3BF68C25FD5E4B37" author="Salgado, A. &amp; Ruiz, G. R." box="[399,662,1832,1857]" pageId="25" pageNumber="476" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz (2017)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEF36D68A6B016B3ABE8C41FC424B83" blockId="25.[151,1437,1256,2037]" pageId="25" pageNumber="476">
<emphasis id="B924EAC48A6B016B3ABE8C41FEF74B13" bold="true" box="[199,319,1868,1893]" pageId="25" pageNumber="476">Variation.</emphasis>
Among the males examined, we observed the following variation in palpal structures (
<figureCitation id="136B2A538A6B016B3F738C41FA974B13" box="[1290,1375,1868,1893]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">Figs 26</figureCitation>
AD, 26G): the superior border of the branched projection of embolic disc can be smooth (
<figureCitation id="136B2A538A6B016B3E208C7DFB664BFF" box="[1113,1198,1904,1929]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">Figs 26</figureCitation>
AB) or occupied by large serial spikes (similar to a comb) (
<figureCitation id="136B2A538A6B016B38358C99FD6B4BDB" box="[588,675,1940,1965]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">Figs 26</figureCitation>
CD). We decided to refrain from describing this variation as a new species and all these forms are considered conspecific. Also, the number and shape of branches on the RvTA is variable (
<figureCitation id="136B2A538A6B016B3B798CD1FEA04B83" box="[256,360,2012,2037]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="25" pageNumber="476">Fig. 26G</figureCitation>
; see
<bibRefCitation id="EFC14B278A6B016B3BE68CD1FD5C4B83" author="Salgado, A. &amp; Ruiz, G. R." box="[415,660,2012,2037]" pageId="25" pageNumber="476" pagination="401 - 437" refId="ref8937" refString="Salgado, A. &amp; Ruiz, G. R. S (2017) Ten new species of Amphidraus Simon, 1900 (Araneae: Salticidae: Euophryini) and three new combinations. Zootaxa, 4312 (3), 401 - 437. https: // doi. org / 10.11646 / zootaxa. 4312.3.1" type="journal article" year="2017">Salgado &amp; Ruiz 2017</bibRefCitation>
: figs 23B, 24B, 24D).
</paragraph>
<paragraph id="8BEF36D68A6801683ABE8B9AFAA34D6A" blockId="26.[151,1436,151,320]" pageId="26" pageNumber="477">
<emphasis id="B924EAC48A6801683ABE8B9AFECE4CC6" bold="true" box="[199,262,151,176]" pageId="26" pageNumber="477">Note.</emphasis>
<emphasis id="B924EAC48A6801683B698B95FD314CC6" box="[272,761,152,176]" italics="true" pageId="26" pageNumber="477">
<taxonomicName id="4C504D558A6801683B698B95FDAB4CC6" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[272,611,152,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="draconicaudatus">Amphidraus draconicaudatus</taxonomicName>
,
<taxonomicName id="4C504D558A680168380B8B95FD314CC6" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[626,761,152,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="shenlong" status="sp. nov.">A. shenlong</taxonomicName>
</emphasis>
<emphasis id="B924EAC48A680168397D8B9AFCA94CC6" bold="true" box="[772,865,151,176]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="A21757BF8A680168397D8B9AFCA94CC6" box="[772,865,151,176]" pageId="26" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C504D558A68016839E68B95FB904CC6" authority="Salgado &amp; Ruiz, 2019" authorityName="Salgado &amp; Ruiz" authorityYear="2019" box="[927,1112,152,176]" class="Arachnida" family="Salticidae" genus="Amphidraus" kingdom="Animalia" order="Araneae" pageId="26" pageNumber="477" phylum="Arthropoda" rank="species" species="draconitupan" status="sp. nov.">
<emphasis id="B924EAC48A68016839E68B95FB904CC6" box="[927,1112,152,176]" italics="true" pageId="26" pageNumber="477">A. draconitupan</emphasis>
</taxonomicName>
<emphasis id="B924EAC48A6801683E1A8B9AFB084CC6" bold="true" box="[1123,1216,151,176]" pageId="26" pageNumber="477">
<taxonomicNameLabel id="A21757BF8A6801683E1A8B9AFB084CC6" box="[1123,1216,151,176]" pageId="26" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
clearly compose a particular group within the genus. The males in this group have one stout projection that emerges from the retrolateral border of the embolic disc and extends towards to retrolateral border of tegulum (
<figureCitation id="136B2A538A6801683ED78BD2FAD34C8E" box="[1198,1307,223,248]" captionStart="FIGURE 22" captionStartId="23.[151,250,1670,1692]" captionTargetBox="[177,1383,890,1615]" captionTargetId="figure@23.[151,1436,879,1648]" captionTargetPageId="23" captionText="FIGURE 22. Amphidraus shenlong sp. nov., left male palp (A ventral; B retrolateral). Abbreviations: cc, cymbial conductor; dPED, distal processes on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601390/files/figure.png" pageId="26" pageNumber="477">Figs 22A</figureCitation>
,
<figureCitation id="136B2A538A6801683F5F8BD2FA914C8E" box="[1318,1369,223,248]" captionStart="FIGURE 25" captionStartId="26.[151,250,1787,1809]" captionTargetBox="[167,1392,393,1736]" captionTargetId="figure@26.[151,1436,370,1755]" captionTargetPageId="26" captionText="FIGURE 25. Amphidraus draconitupan sp. nov., AC left male palp (A ventral, B prolateral, C retrolateral); D epigyne/vulva, cleared, ventral. Abbreviations: cc, cymbial conductor; co, copulatory opening; dPED, distal process on embolic disc; e, embolus shaft; ed, embolic disc; ef, embolic filament; fd, fertilization duct; gl, gland; po, coupling pocket; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; S, spermatheca; TL, tegular lobe; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601396/files/figure.png" pageId="26" pageNumber="477">25A</figureCitation>
,
<figureCitation id="136B2A538A6801683F1C8BD2FA504C8E" box="[1381,1432,223,248]" captionStart="FIGURE 26" captionStartId="27.[151,250,1594,1616]" captionTargetBox="[170,1396,218,1541]" captionTargetId="figure@27.[151,1436,203,1563]" captionTargetPageId="27" captionText="FIGURE 26. Left male palp in Amphidraus spp. AD, G Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AB, G new specimen from Pium (A bulb, ventral; B retroventral; G tibia, retrolateral). CD specimen from Caseara (C bulb, ventral; D retroventral). EF, H Amphidraus shenlong sp. nov. (E bulb, ventral; F retroventral; H tibia, retrolateral). Abbreviations: dPED, distal process on embolic disc; e, embolus shaft; ef, embolic filament; RTA, retrolateral tibial apophysis; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc." httpUri="https://zenodo.org/record/2601398/files/figure.png" pageId="26" pageNumber="477">26A</figureCitation>
,
<figureCitation id="136B2A538A6801683AEE8A09FF044D6A" box="[151,204,260,284]" captionStart="FIGURE 27" captionStartId="28.[151,250,1131,1153]" captionTargetBox="[151,1436,228,1080]" captionTargetId="figure@28.[151,1436,228,1080]" captionTargetPageId="28" captionText="FIGURE 27. Copulatory structures of Amphidraus draconicaudatus Salgado &amp; Ruiz, 2017. AD left male palp (A embolic disc, ventral; patella + tibia, retrolateral; C detail of RvTA projection; D detail of reduced RTA). E epigyne/vulva, dorsal. Abbreviations: dPED, distal processes on embolic disc; e, embolus shaft; ef, embolic filament; po, coupling pocket; RvTA, retroventral tibial apophysis; vPED, ventral process on embolic disc. White arrows in B and C show projection on RvTA, and black arrow in C shows one spike of the reduced RTA." httpUri="https://zenodo.org/record/2601400/files/figure.png" pageId="26" pageNumber="477">27A</figureCitation>
). Moreover, males also have pockets on outer faces of chelicerae (
<figureCitation id="136B2A538A68016839C88A09FBCE4D6A" box="[945,1030,260,285]" captionStart="FIGURE 28" captionStartId="28.[151,250,1966,1988]" captionTargetBox="[151,1436,1345,1922]" captionTargetId="figure@28.[151,1436,1345,1922]" captionTargetPageId="28" captionText="FIGURE 28. Right chelicera, retrolateral (A Amphidraus draconicaudatus, B A. shenlong sp. nov. C A. draconitupan sp. nov.). Black arrow shows pocket on outer face." httpUri="https://zenodo.org/record/2601402/files/figure.png" pageId="26" pageNumber="477">Figs 28</figureCitation>
AC, 29AB), of unknown use.
</paragraph>
</subSubSection>
<subSubSection id="C34A655D8A6801683ABE8A2AFC3B4D36" box="[199,1011,295,320]" pageId="26" pageNumber="477" type="distribution">
<paragraph id="8BEF36D68A6801683ABE8A2AFC3B4D36" blockId="26.[151,1436,151,320]" box="[199,1011,295,320]" pageId="26" pageNumber="477">
<emphasis id="B924EAC48A6801683ABE8A2AFEA84D36" bold="true" box="[199,352,295,320]" pageId="26" pageNumber="477">Distribution.</emphasis>
Known only from the state of
<collectingRegion id="4994F8348A68016838C18A2AFCEC4D36" box="[696,804,295,320]" country="Brazil" name="Tocantins" pageId="26" pageNumber="477">Tocantins</collectingRegion>
(
<collectingCountry id="F34776468A680168394D8A2AFCB34D36" box="[820,891,295,320]" name="Brazil" pageId="26" pageNumber="477">Brazil</collectingCountry>
) (
<figureCitation id="136B2A538A68016839EB8A2AFC2F4D36" box="[914,999,295,320]" captionStart="FIGURE 30" captionStartId="29.[151,250,1729,1751]" captionTargetBox="[151,1436,929,1708]" captionTargetId="figure@29.[151,1436,929,1708]" captionTargetPageId="29" captionText="FIGURE 30. Map of northern Brazil showing records of Amphidraus draconicaudatus, A. shenlong sp. nov. and A. draconitupan sp. nov. White lines represent the borders of Brazilian states. Acronyms of states: MG, Minas Gerais; MS, Mato Grosso do Sul; PA, Pará; SP, São Paulo; TO, Tocantins. Map modified from Google Earth ®." httpUri="https://zenodo.org/record/2601406/files/figure.png" pageId="26" pageNumber="477">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>