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<document id="8D3FAB7C17ADFA140D23A8F03A59AECC" ID-CLB-Dataset="24425" ID-DOI="10.11646/zootaxa.4718.3.1" ID-GBIF-Dataset="0539e3ec-e2d6-4ed3-9f5e-22677babf264" ID-ISSN="1175-5326" ID-Zenodo-Dep="3602430" ID-ZooBank="09F9270F-589A-471D-A912-2F2788544399" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1578564776944" checkinUser="plazi" docAuthor="Mound, Laurence A., Wang, Jun &amp; Tree, Desley J." docDate="2020" docId="03F987BEFF982722FF2BFDEBFB29D5CF" docLanguage="en" docName="zootaxa.4718.3.1.pdf" docOrigin="Zootaxa 4718 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplothrips nelsoni Mound &amp; Wang &amp; Tree 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="310" masterDocId="FFC0FFC6FF91272BFFBCFFCEFFBFD258" masterDocTitle="The genus Hoplothrips in Australia (Thysanoptera, Phlaeothripinae), with eleven new species" masterLastPageNumber="323" masterPageNumber="301" pageNumber="310" updateTime="1698776662637" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="338EB7D43EE1709AA8929684BE07160B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0DA3658264562EA189B39FE86614FE6D">
<mods:title id="085859CE7CCBA175447C61A89B7BE52C">The genus Hoplothrips in Australia (Thysanoptera, Phlaeothripinae), with eleven new species</mods:title>
</mods:titleInfo>
<mods:name id="C7236BFC1D499D35734076BE93DFDADE" type="personal">
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<mods:namePart id="D5466A9EC1FD66AEC68EF46E71125B2B">Mound, Laurence A.</mods:namePart>
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<mods:namePart id="A65B2A5DA2A9DA426FCC4172B77EBF04">Wang, Jun</mods:namePart>
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<mods:roleTerm id="034DB0CEFDDF21D3EB92E361639D369E">Author</mods:roleTerm>
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<mods:namePart id="63BFF8A0B44BDEB3745AA1926209D8F9">Tree, Desley J.</mods:namePart>
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<mods:title id="01E0C7DDA30DFE319EA37C4D4B4F33AF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="027DAAF204B18CC17EB45AB478424438">
<mods:date id="9DBE333E65F7C55D262BE5122869F3CC">2020</mods:date>
<mods:detail id="AC60BB00CE4F03EB33168FC394D3DF60" type="pubDate">
<mods:number id="C5C313C2BEAEA6383F53113566D02FF3">2020-01-07</mods:number>
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<mods:detail id="FBB480E1AD2AE8F8C163DB3098DED7F8" type="volume">
<mods:number id="1232392316FC6D386D7CEA82603A520D">4718</mods:number>
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<mods:number id="BE6768854474DC3BA250E2627F712179">3</mods:number>
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<mods:classification id="F317FB36E60BEB3D88B63FDF030BF49D">journal article</mods:classification>
<mods:identifier id="E03F5C95D71E1CF1A2DA17D0E83E5BB0" type="CLB-Dataset">24425</mods:identifier>
<mods:identifier id="DF2E1DAA5BE7C4CC7ACB9923216EB8A9" type="DOI">10.11646/zootaxa.4718.3.1</mods:identifier>
<mods:identifier id="39D70B06281A12F5B721D2E5B506CE52" type="GBIF-Dataset">0539e3ec-e2d6-4ed3-9f5e-22677babf264</mods:identifier>
<mods:identifier id="C6C9D3100D788A7827CF74205AF5DD2A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F987BEFF982722FF2BFDEBFB29D5CF" ID-DOI="http://doi.org/10.5281/zenodo.5920708" ID-GBIF-Taxon="161670903" ID-Zenodo-Dep="5920708" LSID="urn:lsid:plazi:treatment:03F987BEFF982722FF2BFDEBFB29D5CF" httpUri="http://treatment.plazi.org/id/03F987BEFF982722FF2BFDEBFB29D5CF" lastPageNumber="310" pageId="9" pageNumber="310">
<subSubSection id="C34A6523FF982722FF2BFDEBFE6ED067" box="[151,465,549,576]" pageId="9" pageNumber="310" type="nomenclature">
<paragraph id="8BEF36A8FF982722FF2BFDEBFE6ED067" blockId="9.[151,465,549,612]" box="[151,465,549,576]" pageId="9" pageNumber="310">
<heading id="D0A781C4FF982722FF2BFDEBFE6ED067" bold="true" box="[151,465,549,576]" fontSize="11" level="1" pageId="9" pageNumber="310" reason="1">
<emphasis id="B924EABAFF982722FF2BFDEBFE6ED067" bold="true" box="[151,465,549,576]" pageId="9" pageNumber="310">
<taxonomicName id="4C504D2BFF982722FF2BFDEBFE37D018" ID-CoL="3MNCD" authority="Mound &amp; Wang &amp; Tree, 2020" authorityName="Mound &amp; Wang &amp; Tree" authorityYear="2020" box="[151,392,549,576]" class="Insecta" family="Phlaeothripidae" genus="Hoplothrips" kingdom="Animalia" order="Thysanoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="nelsoni" status="sp. nov.">
<emphasis id="B924EABAFF982722FF2BFDEBFE37D018" bold="true" box="[151,392,549,576]" italics="true" pageId="9" pageNumber="310">Hoplothrips nelsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757C1FF982722FE33FDEBFE6ED067" box="[399,465,549,575]" pageId="9" pageNumber="310" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6523FF982722FF2BFD87FCC2D55F" pageId="9" pageNumber="310" type="description">
<paragraph id="8BEF36A8FF982722FF2BFD87FEEED03C" blockId="9.[151,465,549,612]" box="[151,337,585,612]" pageId="9" pageNumber="310">
(
<figureCitation id="136B2A2DFF982722FF23FD87FE9FD03B" box="[159,288,585,611]" captionStart="FIGURES 10–20" captionStartId="12.[151,264,1944,1969]" captionTargetBox="[151,1435,337,1925]" captionTargetId="figure@12.[151,1435,337,1925]" captionTargetPageId="12" captionText="FIGURES 10–20. Hoplothrips species—head and pronotum. (10) lowdeni male; (11) lowdeni female; (12) nelsoni male; (13) nelsoni female; (14) oakeyi female; (15) oakeyi male; (16) reedi male; (17) reedi pronotum; (18) melanurus pronotum; (19) tarsus; (20) semicaecus female macroptera." figureDoi="http://doi.org/10.5281/zenodo.3602434" httpUri="https://zenodo.org/record/3602434/files/figure.png" pageId="9" pageNumber="310">Figs 12, 13</figureCitation>
,
<figureCitation id="136B2A2DFF982722FE90FD87FEF6D03C" box="[300,329,585,612]" captionStart="FIGURES 31–45" captionStartId="17.[151,264,1924,1949]" captionTargetBox="[151,1435,196,1891]" captionTargetId="figure@17.[151,1435,196,1891]" captionTargetPageId="17" captionText="FIGURES 31–45. Hoplothrips species antennae. (31) bellingeni; (32) corticis (III–V); (33) giganteus; (34) lihongae; (35) lowdeni; (36) nelsoni; (37) oakeyi; (38) wrightae; (39) orientalis (III–V); (40) poultoni; (41) reedi; (42) semicaecus female macroptera; (43) semicaecus female microptera; (44) semicaecus male microptera; (45) woodsi." figureDoi="http://doi.org/10.5281/zenodo.3602438" httpUri="https://zenodo.org/record/3602438/files/figure.png" pageId="9" pageNumber="310">36</figureCitation>
)
</paragraph>
<paragraph id="8BEF36A8FF982722FF2BFD5FFAE8D627" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF2BFD5FFECAD0F3" box="[151,373,657,683]" italics="true" pageId="9" pageNumber="310">Female macroptera</emphasis>
. Body, femora and antennae light brown with tube darker; tibiae shaded with brown, tarsi almost yellow; antennal segment III with basal third pale (
<figureCitation id="136B2A2DFF982722FCAFFD7BFCD7D088" box="[787,872,693,720]" captionStart="FIGURES 31–45" captionStartId="17.[151,264,1924,1949]" captionTargetBox="[151,1435,196,1891]" captionTargetId="figure@17.[151,1435,196,1891]" captionTargetPageId="17" captionText="FIGURES 31–45. Hoplothrips species antennae. (31) bellingeni; (32) corticis (III–V); (33) giganteus; (34) lihongae; (35) lowdeni; (36) nelsoni; (37) oakeyi; (38) wrightae; (39) orientalis (III–V); (40) poultoni; (41) reedi; (42) semicaecus female macroptera; (43) semicaecus female microptera; (44) semicaecus male microptera; (45) woodsi." figureDoi="http://doi.org/10.5281/zenodo.3602438" httpUri="https://zenodo.org/record/3602438/files/figure.png" pageId="9" pageNumber="310">Fig. 36</figureCitation>
). Head without sculpture apart from a few lines at posterior margin; genae subparallel; ocelli present; dorsal length of compound eyes greater than one third of head length; postocular setae capitate, as longas dorsal eye length; maxillary stylets retracted to postocular setae and about one quarter of head width apart medially, weak maxillary bridge present (
<figureCitation id="136B2A2DFF982722FB8FFCEFFB36D163" box="[1075,1161,801,827]" captionStart="FIGURES 10–20" captionStartId="12.[151,264,1944,1969]" captionTargetBox="[151,1435,337,1925]" captionTargetId="figure@12.[151,1435,337,1925]" captionTargetPageId="12" captionText="FIGURES 10–20. Hoplothrips species—head and pronotum. (10) lowdeni male; (11) lowdeni female; (12) nelsoni male; (13) nelsoni female; (14) oakeyi female; (15) oakeyi male; (16) reedi male; (17) reedi pronotum; (18) melanurus pronotum; (19) tarsus; (20) semicaecus female macroptera." figureDoi="http://doi.org/10.5281/zenodo.3602434" httpUri="https://zenodo.org/record/3602434/files/figure.png" pageId="9" pageNumber="310">Fig. 13</figureCitation>
). Antennal segments III and IV each with 2 sense cones, IV–VII each with pedicel; VIII broadly connected with VII (
<figureCitation id="136B2A2DFF982722FB7FFC8BFAA6D138" box="[1219,1305,837,864]" captionStart="FIGURES 31–45" captionStartId="17.[151,264,1924,1949]" captionTargetBox="[151,1435,196,1891]" captionTargetId="figure@17.[151,1435,196,1891]" captionTargetPageId="17" captionText="FIGURES 31–45. Hoplothrips species antennae. (31) bellingeni; (32) corticis (III–V); (33) giganteus; (34) lihongae; (35) lowdeni; (36) nelsoni; (37) oakeyi; (38) wrightae; (39) orientalis (III–V); (40) poultoni; (41) reedi; (42) semicaecus female macroptera; (43) semicaecus female microptera; (44) semicaecus male microptera; (45) woodsi." figureDoi="http://doi.org/10.5281/zenodo.3602438" httpUri="https://zenodo.org/record/3602438/files/figure.png" pageId="9" pageNumber="310">Fig. 36</figureCitation>
). Pronotum without sculpture; 4 pairs of capitate major setae, am setae small and pointed. Mesonotum with with weak transverse sculpture, lateral setal pair capitate; metanotum without sculpture medially, median setal pair long and finely pointed. Prosternal basantra absent, ferna acute medially but not meeting; mesopresternum reduced to 2 triangles; mesoeusternal anterior margin entire. Fore tarsal tooth pointed, length equals half of tarsal width (
<figureCitation id="136B2A2DFF982722FB41FC1BFAEDD1B7" box="[1277,1362,981,1007]" captionStart="FIGURES 10–20" captionStartId="12.[151,264,1944,1969]" captionTargetBox="[151,1435,337,1925]" captionTargetId="figure@12.[151,1435,337,1925]" captionTargetPageId="12" captionText="FIGURES 10–20. Hoplothrips species—head and pronotum. (10) lowdeni male; (11) lowdeni female; (12) nelsoni male; (13) nelsoni female; (14) oakeyi female; (15) oakeyi male; (16) reedi male; (17) reedi pronotum; (18) melanurus pronotum; (19) tarsus; (20) semicaecus female macroptera." figureDoi="http://doi.org/10.5281/zenodo.3602434" httpUri="https://zenodo.org/record/3602434/files/figure.png" pageId="9" pageNumber="310">Fig. 13</figureCitation>
). Fore wing with three small, weakly capitate, sub-basal setae; about 6 duplicated cilia present. Pelta broadly triangular with small lateral lobes; tergites III–VII each with two pairs of sigmoid wing-retaining setae, these are weak on tergite II; median tergal setae acute but not reaching posterior margins of tergites; tergite VIII median setae capitate, tergite IX setae S1 shorter than tube and capitate, S2 shorter than tube and blunt; anal setae longer than tube.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BFB47FCE6D70F" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BFB47FEC9D6FB" bold="true" box="[199,374,1161,1187]" pageId="9" pageNumber="310">Measurements</emphasis>
(
<typeStatus id="54EB880AFF982722FE38FB47FE57D6FB" box="[388,488,1161,1187]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
female in microns). Body length 1950. Head, length 193; width 207; postocular setae 57, distance between their bases 163; postocellar setae 10, distance between their bases 53. Pronotum, length 125; median width 250; major setae am 15, aa 45, ml 40, epim 51, pa 33. Mesonotum lateral setae 30; Metanotum median setae 29. Tergite IV median marginal setae 65; tergite VIII median setae 60, tergite IX setae S1 78, S2 78. Tube length 118. Anal setae 175. Antennal segments III–VIII length (width) 60 (30), 60 (30), 55 (26), 45 (23), 43 (21), 30 (13). Fore wing length 75, sub-basal setae
<geoCoordinate id="EE64506FFF982722FDC8FAF3FD09D70F" box="[628,694,1341,1367]" degrees="1" direction="south" minutes="28" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="-1.4666667">S1 28</geoCoordinate>
,
<geoCoordinate id="EE64506FFF982722FD7EFAF3FCBAD70F" box="[706,773,1341,1367]" degrees="2" direction="south" minutes="28" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="-2.4666667">S2 28</geoCoordinate>
,
<geoCoordinate id="EE64506FFF982722FCADFAF3FCEAD70F" box="[785,853,1340,1367]" degrees="3" direction="south" minutes="25" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="-3.4166667">S3 25</geoCoordinate>
.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BFAAFFD2DD7C7" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BFAAFFED1D723" box="[199,366,1377,1403]" italics="true" pageId="9" pageNumber="310">Female aptera</emphasis>
. Similar to macropterous female, but compound eyes smaller, ocelli absent; pelta D-shaped; tergal wing-retaining setae short and pointed.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BFA67FAA7D79B" blockId="9.[151,1437,657,1943]" box="[199,1304,1448,1476]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BFA67FEC9D79B" bold="true" box="[199,374,1449,1475]" pageId="9" pageNumber="310">Measurements</emphasis>
(in microns). Body length 1680. Head, length 185; width 170; postocular setae 51.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BFA03FE17D477" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BFA03FEEFD7BF" box="[199,336,1485,1511]" italics="true" pageId="9" pageNumber="310">Male aptera</emphasis>
. Similar to apterous female but smaller, with fore leg and tarsal tooth larger (
<figureCitation id="136B2A2DFF982722FB1BFA03FB46D7BF" box="[1191,1273,1485,1511]" captionStart="FIGURES 10–20" captionStartId="12.[151,264,1944,1969]" captionTargetBox="[151,1435,337,1925]" captionTargetId="figure@12.[151,1435,337,1925]" captionTargetPageId="12" captionText="FIGURES 10–20. Hoplothrips species—head and pronotum. (10) lowdeni male; (11) lowdeni female; (12) nelsoni male; (13) nelsoni female; (14) oakeyi female; (15) oakeyi male; (16) reedi male; (17) reedi pronotum; (18) melanurus pronotum; (19) tarsus; (20) semicaecus female macroptera." figureDoi="http://doi.org/10.5281/zenodo.3602434" httpUri="https://zenodo.org/record/3602434/files/figure.png" pageId="9" pageNumber="310">Fig. 12</figureCitation>
); tergite IX S2 setae shorter; sternite VIII pore plate transverse, about a third of sternite width; sternites VI–VII without specialised reticulate areas laterally.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BF9F7FAC6D4C3" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BF9F7FEC9D40B" bold="true" box="[199,374,1593,1619]" pageId="9" pageNumber="310">Measurements</emphasis>
(in microns). Body length 1390. Head, length 175; width 158; postocular setae 70. Pronotum, length 128; median width 225; major setae am 10, aa 51, ml 25, epim 55, pa 40. Tergite VIII median setae 41, tergite IX setae S1 73,
<geoCoordinate id="EE64506FFF982722FEF5F94FFE32D4C3" box="[329,397,1665,1691]" degrees="2" direction="south" minutes="36" orientation="latitude" pageId="9" pageNumber="310" precision="925" value="-2.6">S2 36</geoCoordinate>
. Tube length 88. anal setae 133. Antennal segments III–VIII length 53, 50, 50, 49, 42, 23.
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BF96AFE38D4BB" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BF96AFE13D4E7" bold="true" box="[199,428,1700,1727]" pageId="9" pageNumber="310">Specimens studied.</emphasis>
<materialsCitation id="3B383CF5FF982722FE09F96BFE38D4BB" ID-GBIF-Occurrence="2545450435" collectingDate="2013-10-05" collectionCode="ANIC" location="Victoria, Nelson" pageId="9" pageNumber="310" specimenCode="DJT 1687" specimenCount="1" stateProvince="Victoria, Nelson" typeStatus="holotype">
<typeStatus id="54EB880AFF982722FE09F96BFDA2D4E7" box="[437,541,1701,1727]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
female macroptera.
<collectingRegion id="4994F84AFF982722FCB4F96AFC74D4E7" box="[776,971,1700,1727]" isEnumeration="true" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FCB4F96AFCD6D4E7" bold="true" box="[776,873,1700,1727]" pageId="9" pageNumber="310">Victoria</emphasis>
, Nelson
</collectingRegion>
, from bark spray of living tree,
<date id="FFEE1068FF982722FA86F96AFA23D4E7" box="[1338,1436,1700,1727]" pageId="9" pageNumber="310" value="2013-10-05">
<collectingDate id="EFAAE980FF982722FA86F96AFA23D4E7" box="[1338,1436,1700,1727]" pageId="9" pageNumber="310" value="2013-10-05">5.x.2013</collectingDate>
</date>
(
<specimenCode id="DBF69ED3FF982722FF23F907FEAFD4BC" box="[159,272,1737,1764]" collectionCode="DJT" pageId="9" pageNumber="310">DJT 1687</specimenCode>
) in
<collectionCode id="ED41AE6DFF982722FE87F907FE3CD4BB" box="[315,387,1737,1763]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="310">ANIC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEF36A8FF982722FF7BF923FCC2D55F" blockId="9.[151,1437,657,1943]" box="[199,893,1773,1799]" pageId="9" pageNumber="310">
<typeStatus id="54EB880AFF982722FF7BF923FE87D55F" box="[199,312,1773,1799]" pageId="9" pageNumber="310" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D56FD21FF982722FEFFF923FE1BD55F" box="[323,420,1773,1799]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
aptera and
<materialsCitation id="3B383CF5FF982722FD9AF923FCC5D55F" ID-GBIF-Occurrence="2545450414" box="[550,890,1773,1799]" collectingDate="2013-10-05" collectionCode="ANIC" location="Victoria, Nelson" pageId="9" pageNumber="310" specimenCount="1" specimenCount-male="1" stateProvince="Victoria, Nelson" typeStatus="paratype">
<specimenCount id="9D56FD21FF982722FD9AF923FDCBD55F" box="[550,628,1773,1799]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6523FF982722FF7BF8DEFB29D5CF" pageId="9" pageNumber="310" type="discussion">
<paragraph id="8BEF36A8FF982722FF7BF8DEFB29D5CF" blockId="9.[151,1437,657,1943]" pageId="9" pageNumber="310">
<emphasis id="B924EABAFF982722FF7BF8DEFEF0D573" bold="true" box="[199,335,1808,1835]" pageId="9" pageNumber="310">Comments.</emphasis>
Within the genus
<taxonomicName id="4C504D2BFF982722FDA3F8DFFD19D573" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[543,678,1809,1835]" class="Insecta" family="Phlaeothripidae" genus="Hoplothrips" kingdom="Animalia" order="Thysanoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">
<emphasis id="B924EABAFF982722FDA3F8DFFD19D573" box="[543,678,1809,1835]" italics="true" pageId="9" pageNumber="310">Hoplothrips</emphasis>
</taxonomicName>
this species is unusual in having a maxillary bridge. It is generally similar to
<taxonomicName id="4C504D2BFF982722FEB6F8F8FE2AD517" authorityName="Hood" authorityYear="1919" box="[266,405,1846,1871]" class="Insecta" family="Phlaeothripidae" genus="Trichothrips" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="connexus">
<emphasis id="B924EABAFF982722FEB6F8F8FE2AD517" box="[266,405,1846,1871]" italics="true" pageId="9" pageNumber="310">H. connexus</emphasis>
</taxonomicName>
but with shorter median setae on tergites II–VII, and the macroptera has two pairs of welldeveloped sigmoid wing-retaining setae. The pore plate on sternite VIII of the male is transverse but present only on the median third of the sternite, and sternite VII has no specialised reticulate areas laterally.
</paragraph>
</subSubSection>
</treatment>
</document>
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