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<document id="C2A33A84FA470E43DCAE0BD67198FF48" ID-DOI="10.5281/zenodo.4504495" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504495" ID-ZooBank="70097B8F-3EF2-4740-97D4-AD2357234011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612466532816" checkinUser="carolina" docAuthor="Kottelat, Maurice" docDate="2017" docId="03F987ACFFBDFF8BFED3CFC18DF5FE99" docLanguage="en" docName="RafflesBZool.65.80-99.pdf" docOrigin="Raffles Bulletin of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Malihkaia aligera Kottelat 2017, new species" docType="treatment" docUuid="9DE7EBCC-00EB-49CE-9F47-551D629190AC" docUuidSource="ZooBank" docVersion="5" lastPageNumber="87" masterDocId="FFC0FFD4FFB8FF83FFCACA0F8F10FFCD" masterDocTitle="A new genus and three new species of nemacheilid loaches from northern Irrawaddy drainage, Myanmar (Teleostei: Cypriniformes)" masterLastPageNumber="99" masterPageNumber="80" pageNumber="85" updateTime="1698866246585" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9C3CE24023C51F6EAB51E6692A554DEC">A new genus and three new species of nemacheilid loaches from northern Irrawaddy drainage, Myanmar (Teleostei: Cypriniformes)</mods:title>
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<mods:namePart id="C334C3CEE98B40839B1BAE101DA0C23D">Kottelat, Maurice</mods:namePart>
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<mods:title id="A254109320392A422BA4A2A872FD476B">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="0FFB45CEBEE9572AC0891EECC044E9FD">2017</mods:date>
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<subSubSection id="C34A6531FFBDFF86FED3CFC18D66FA2B" box="[281,630,1486,1510]" pageId="5" pageNumber="85" type="nomenclature">
<paragraph id="8BEF36BAFFBDFF86FED3CFC18D66FA2B" blockId="5.[281,630,1486,1542]" box="[281,630,1486,1510]" pageId="5" pageNumber="85">
<heading id="D0A781D6FFBDFF86FED3CFC18D66FA2B" bold="true" box="[281,630,1486,1510]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="85" reason="2">
<emphasis id="B924EAA8FFBDFF86FED3CFC18D66FA2B" bold="true" box="[281,630,1486,1510]" pageId="5" pageNumber="85">
<taxonomicName id="4C504D39FFBDFF86FED3CFC18EF1FA2B" ID-CoL="3XMXP" authority="Kottelat, 2017" authorityName="Kottelat" authorityYear="2017" box="[281,481,1486,1510]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="85" phylum="Chordata" rank="species" species="aligera" status="sp. nov.">
<emphasis id="B924EAA8FFBDFF86FED3CFC18EF1FA2B" bold="true" box="[281,481,1486,1510]" italics="true" pageId="5" pageNumber="85">Malihkaia aligera</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21757D3FFBDFF86FE3ACFC18D66FA2B" box="[496,630,1486,1510]" pageId="5" pageNumber="85" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34A6531FFBDFF86FE41CFE18D14F9CB" box="[395,516,1518,1542]" pageId="5" pageNumber="85" type="description">
<paragraph id="8BEF36BAFFBDFF86FE41CFE18D14F9CB" blockId="5.[281,630,1486,1542]" box="[395,516,1518,1542]" pageId="5" pageNumber="85">
(
<figureCitation id="136B2A3FFFBDFF86FE59CFE18ECEF9CB" box="[403,478,1518,1542]" captionStart="Fig" captionStartId="2.[113,148,1528,1550]" captionTargetBox="[114,1452,189,1512]" captionTargetId="figure-9@2.[114,1452,189,1512]" captionTargetPageId="2" captionText="Fig. 1. Malihkaia aligera, MHNG 2766.051, holotype, male, 65.7 mm SL; Myanmar: Irrawaddy drainage; Mali Hka River near Putao." figureDoi="http://doi.org/10.5281/zenodo.4504497" httpUri="https://zenodo.org/record/4504497/files/figure.png" pageId="5" pageNumber="85">Figs. 1</figureCitation>
,
<figureCitation id="136B2A3FFFBDFF86FE24CFE18EECF9CB" box="[494,508,1518,1542]" captionStart="Fig" captionStartId="3.[135,170,1975,1997]" captionTargetBox="[135,1473,278,1960]" captionTargetId="figure-33@3.[135,1473,278,1960]" captionTargetPageId="3" captionText="Fig. 2. Malihkaia aligera, CMK 25508, paratypes; a, male, 73.0 mm SL; b, female, 77.2 mm SL; Myanmar: Irrawaddy drainage: Mali Hka River near Putao." figureDoi="http://doi.org/10.5281/zenodo.4504499" httpUri="https://zenodo.org/record/4504499/files/figure.png" pageId="5" pageNumber="85">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A6531FFBDFF86FF42CC218E52F8EB" pageId="5" pageNumber="85" type="materials_examined">
<paragraph id="8BEF36BAFFBDFF86FF42CC218D21F96B" blockId="5.[135,776,1582,1702]" pageId="5" pageNumber="85">
<materialsCitation id="3B383CE7FFBDFF86FF42CC218D21F96B" collectingDate="2014-11-26" collectionCode="MHNG" collectorName="M. Kottelat &amp; Nyein Chan" country="Myanmar" elevation="402" latitude="27.431665" location="Kang Mu Lon" longLatPrecision="20" longitude="97.46555" municipality="Hka River" pageId="5" pageNumber="85" specimenCode="MHNG 2766.051" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B924EAA8FFBDFF86FF42CC218FE7F98B" bold="true" box="[136,247,1582,1606]" pageId="5" pageNumber="85">
<typeStatus id="54EB8818FFBDFF86FF42CC218FE2F98B" box="[136,242,1582,1606]" pageId="5" pageNumber="85" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBF69EC1FFBDFF86FF37CC218EABF98B" box="[253,443,1582,1606]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="85" type="Museum">MHNG 2766.051</specimenCode>
, male,
<quantity id="4CA89B5FFFBDFF86FDCCCC218D74F98B" box="[518,612,1582,1606]" metricMagnitude="-2" metricUnit="m" metricValue="6.57" pageId="5" pageNumber="85" unit="mm" value="65.7">65.7 mm</quantity>
SL;
<collectingCountry id="F347762AFFBDFF86FD52CC208C14F98A" box="[664,772,1583,1607]" name="Myanmar" pageId="5" pageNumber="85">Myanmar</collectingCountry>
:
<collectingRegion id="4994F858FFBDFF86FF42CC418E03F9AB" box="[136,275,1614,1638]" country="Myanmar" name="Kachin" pageId="5" pageNumber="85">Kachin State</collectingRegion>
: Mali
<collectingMunicipality id="6B8BACC0FFBDFF86FE9ECC418ED4F9AB" box="[340,452,1614,1638]" pageId="5" pageNumber="85">Hka River</collectingMunicipality>
, about
<quantity id="4CA89B5FFFBDFF86FDC4CC418D53F9AB" box="[526,579,1614,1638]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="85" unit="km" value="9.0">9 km</quantity>
upstream of
<location id="8E8F6061FFBDFF86FD06CC408FF4F94B" LSID="urn:lsid:plazi:treatment:03F987ACFFBDFF8BFED3CFC18DF5FE99:8E8F6061FFBDFF86FD06CC408FF4F94B" country="Myanmar" latitude="27.431665" longLatPrecision="20" longitude="97.46555" municipality="Hka River" name="Kang Mu Lon" pageId="5" pageNumber="85" stateProvince="Kachin">Kang Mu Lon</location>
;
<quantity id="4CA89B5FFFBDFF86FF39CC618E4BF94B" box="[243,347,1646,1670]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="5" pageNumber="85" unit="m" value="402.0">
<elevation id="007DD189FFBDFF86FF39CC618E4BF94B" box="[243,347,1646,1670]" metricMagnitude="2" metricUnit="m" metricValue="4.02" pageId="5" pageNumber="85" unit="m" value="402.0">402 masl</elevation>
</quantity>
;
<geoCoordinate id="EE64507DFFBDFF86FEA3CC628EF8F94B" box="[361,488,1645,1670]" degrees="27" direction="north" minutes="25" orientation="latitude" pageId="5" pageNumber="85" precision="15" seconds="54" value="27.431665">27°2554″N</geoCoordinate>
<geoCoordinate id="EE64507DFFBDFF86FE39CC628D63F94B" box="[499,627,1645,1670]" degrees="97" direction="east" minutes="27" orientation="longitude" pageId="5" pageNumber="85" precision="15" seconds="56" value="97.46555">97°2756″E</geoCoordinate>
;
<collectorName id="26A5536CFFBDFF86FD4ACC608C12F94B" box="[640,770,1646,1670]" pageId="5" pageNumber="85">M. Kottelat</collectorName>
,
<collectorName id="26A5536CFFBDFF86FF42CC818E1AF96B" box="[136,266,1678,1702]" pageId="5" pageNumber="85">Nyein Chan</collectorName>
et al.,
<date id="FFEE107AFFBDFF86FE92CC818D3EF96B" box="[344,558,1678,1702]" pageId="5" pageNumber="85" value="2014-11-26">
<collectingDate id="EFAAE992FFBDFF86FE92CC818D3EF96B" box="[344,558,1678,1702]" pageId="5" pageNumber="85" value="2014-11-26">26 November 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEF36BAFFBDFF86FF42CCC18E52F8EB" blockId="5.[136,775,1742,1830]" pageId="5" pageNumber="85">
<materialsCitation id="3B383CE7FFBDFF86FF42CCC18E5DF8CB" accessionNumber="CMK 25508" collectionCode="CMK" pageId="5" pageNumber="85" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<emphasis id="B924EAA8FFBDFF86FF42CCC18E14F92B" bold="true" box="[136,260,1742,1766]" pageId="5" pageNumber="85">
<typeStatus id="54EB8818FFBDFF86FF42CCC18FEFF92B" box="[136,255,1742,1766]" pageId="5" pageNumber="85" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED41AE7FFFBDFF86FEC4CCC18E5BF92B" box="[270,331,1742,1766]" pageId="5" pageNumber="85">CMK</collectionCode>
25508,
<specimenCount id="9D56FD33FFBDFF86FE61CCC18D15F92B" box="[427,517,1742,1766]" pageId="5" pageNumber="85" type="male">2 males</specimenCount>
, 64.973.0,
<specimenCount id="9D56FD33FFBDFF86FD59CCC18C12F92B" box="[659,770,1742,1766]" pageId="5" pageNumber="85" type="female">2 females</specimenCount>
,
<quantity id="4CA89B5FFFBDFF86FF42CCE18E33F8CB" box="[136,291,1774,1798]" metricMagnitude="-2" metricUnit="m" metricValue="7.515" metricValueMax="7.72" metricValueMin="7.31" pageId="5" pageNumber="85" unit="mm" value="75.15" valueMax="77.2" valueMin="73.1">73.177.2 mm</quantity>
SL
</materialsCitation>
;
<materialsCitation id="3B383CE7FFBDFF86FE93CCE18E2EF8EB" accessionNumber="ZRC 55630" collectingDate="2014-11-26" collectionCode="ZRC" collectorName="M. Kottelat &amp; Nyein Chan" country="Myanmar" elevation="402" latitude="27.431665" location="Kang Mu Lon" longLatPrecision="20" longitude="97.46555" municipality="Hka River" pageId="5" pageNumber="85" specimenCount="1" specimenCount-male="1" stateProvince="Kachin" typeStatus="paratype">
<collectionCode id="ED41AE7FFFBDFF86FE93CCE18E9DF8CB" box="[345,397,1774,1798]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="85">ZRC</collectionCode>
55630,
<specimenCount id="9D56FD33FFBDFF86FE2FCCE18D3FF8CB" box="[485,559,1774,1798]" pageId="5" pageNumber="85" type="male">1 male</specimenCount>
,
<quantity id="4CA89B5FFFBDFF86FDF1CCE18D8AF8CB" box="[571,666,1774,1798]" metricMagnitude="-2" metricUnit="m" metricValue="6.59" pageId="5" pageNumber="85" unit="mm" value="65.9">65.9 mm</quantity>
SL; same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6531FFBDFF86FF42CD418D29F84B" pageId="5" pageNumber="85" type="diagnosis">
<paragraph id="8BEF36BAFFBDFF86FF42CD418D29F84B" blockId="5.[134,775,1870,1926]" pageId="5" pageNumber="85">
<emphasis id="B924EAA8FFBDFF86FF42CD418FE9F8AB" bold="true" box="[136,249,1870,1894]" pageId="5" pageNumber="85">Diagnosis.</emphasis>
See generic diagnosis for characters distinguishing the species from all other nemacheilids.
</paragraph>
</subSubSection>
<subSubSection id="C34A6531FFBDFF84FF42CDA18DD8FF38" lastPageId="7" lastPageNumber="87" pageId="5" pageNumber="85" type="description">
<paragraph id="8BEF36BAFFBDFF86FF42CDA18C8CF8AA" blockId="5.[136,776,1966,2086]" lastBlockId="5.[834,1476,1486,1895]" pageId="5" pageNumber="85">
<emphasis id="B924EAA8FFBDFF86FF42CDA18E04F80B" bold="true" box="[136,276,1966,1990]" pageId="5" pageNumber="85">Description.</emphasis>
See
<figureCitation id="136B2A3FFFBDFF86FE84CDA18E8AF80B" box="[334,410,1966,1990]" captionStart="Fig" captionStartId="2.[113,148,1528,1550]" captionTargetBox="[114,1452,189,1512]" captionTargetId="figure-9@2.[114,1452,189,1512]" captionTargetPageId="2" captionText="Fig. 1. Malihkaia aligera, MHNG 2766.051, holotype, male, 65.7 mm SL; Myanmar: Irrawaddy drainage; Mali Hka River near Putao." figureDoi="http://doi.org/10.5281/zenodo.4504497" httpUri="https://zenodo.org/record/4504497/files/figure.png" pageId="5" pageNumber="85">Figs. 1</figureCitation>
&amp;
<figureCitation id="136B2A3FFFBDFF86FE0ECDA18EC1F80B" box="[452,465,1966,1990]" captionStart="Fig" captionStartId="3.[135,170,1975,1997]" captionTargetBox="[135,1473,278,1960]" captionTargetId="figure-33@3.[135,1473,278,1960]" captionTargetPageId="3" captionText="Fig. 2. Malihkaia aligera, CMK 25508, paratypes; a, male, 73.0 mm SL; b, female, 77.2 mm SL; Myanmar: Irrawaddy drainage: Mali Hka River near Putao." figureDoi="http://doi.org/10.5281/zenodo.4504499" httpUri="https://zenodo.org/record/4504499/files/figure.png" pageId="5" pageNumber="85">2</figureCitation>
for general appearance and
<tableCitation id="C6D20301FFBDFF86FF42CDC18FCBF82B" box="[136,219,1998,2022]" captionStart="Table 1" captionStartId="5.[136,191,189,211]" captionText="Table 1. Morphometric data of type material of Malihkaia aligera (n=6). Range and mean include holotype data." pageId="5" pageNumber="85">Table 1</tableCitation>
for morphometric data of
<typeStatus id="54EB8818FFBDFF86FDC9CDC18D70F82B" box="[515,608,1998,2022]" pageId="5" pageNumber="85" type="holotype">holotype</typeStatus>
and
<typeStatus id="54EB8818FFBDFF86FD50CDC08C14F82A" box="[666,772,1999,2023]" pageId="5" pageNumber="85" type="paratype">paratypes</typeStatus>
. An elongate nemacheilid with body depth increasing until somewhat in front of dorsal-fin origin, then decreasing until posterior extremity of dorsal-fin base. Behind dorsal fin, body depth almost uniform until caudal-fin base. Dorsal profile with a marked hump behind head. Head strongly arched in lateral view, slightly depressed; body slightly compressed anteriorly to very compressed on caudal peduncle. Interorbital area arched. Eye below dorsal profile of head. Head deep, with high cheek. Cheeks not swollen (in both sexes). Snout rounded in dorsal and lateral view. Caudal peduncle 1.31.5 times longer than deep, of uniform depth. Low dorsal keel on posterior half of post-dorsal area. Low ventral keel on posterior half of caudal peduncle. Dorsal keel continuous with upper margin of caudal fin. Largest recorded size
<quantity id="4CA89B5FFFBDFF86FA59CD208C7CF8AB" metricMagnitude="-2" metricUnit="m" metricValue="7.72" pageId="5" pageNumber="85" unit="mm" value="77.2">77.2 mm</quantity>
SL.
</paragraph>
<paragraph id="8BEF36BAFFBDFF85FC89CD818D41FD9B" blockId="5.[835,1474,1933,2087]" lastBlockId="6.[113,753,189,598]" lastPageId="6" lastPageNumber="86" pageId="5" pageNumber="85">
Dorsal fin with 4 unbranched and 8½ branched rays; distal margin straight to slightly concave. Second branched ray longest. In first branched ray, anterior branch not branched a second time, shorter than posterior branch. Pectoral fin with 1 unbranched and 10 branched rays, falcate in males, reaching beyond pelvic-fin base, rounded in females, reaching about two thirds of distance to pelvic-fin base (see diagnosis of genus for more detailed description of pectoral fin;
<figureCitation id="136B2A3FFFBEFF85FF6ACB128FF6FEF8" box="[160,230,285,309]" captionStart="Fig" captionStartId="4.[813,848,863,885]" captionTargetBox="[813,1452,189,848]" captionTargetId="figure-50@4.[813,1452,189,848]" captionTargetPageId="4" captionText="Fig. 4. Malihkaia aligera, CMK 25508, 73.0 mm SL; left pectoral fin of male; dorsal (above) and ventral (below) view. Extremity of fourth branched ray damaged." figureDoi="http://doi.org/10.5281/zenodo.4504505" httpUri="https://zenodo.org/record/4504505/files/figure.png" pageId="6" pageNumber="86">Fig. 4</figureCitation>
). Pelvic fin with 1 unbranched and 6 (1) or 7 (5*) branched rays, reaching about three-fifths of distance to anal-fin origin (not reaching anus); rounded; posterior margin straight; origin below base of branched dorsal-fin rays 1 to 2. Axillary pelvic lobe present, entirely free. Anus situated about 1.52 eye diameter in front of anal fin. Anal fin with 3 unbranched and 5½ branched rays; distal margin straight. Caudal fin with 9+8 branched rays; forked, lobes rounded, lower lobe slightly longer than upper one; upper lobe 1.31.5 times longer than median rays.
</paragraph>
<paragraph id="8BEF36BAFFBEFF85FFBBC8728E44FCD8" blockId="6.[112,753,637,789]" pageId="6" pageNumber="86">Body entirely scaled except on anterior half of predorsal area and belly in front of pelvic fins. Lateral line complete, with 8999 pores. Cephalic lateral line system with 6 supraorbital, 4+1011 infraorbital, 89 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8BEF36BAFFBEFF85FFBBC9328D50FB18" blockId="6.[112,755,829,1237]" pageId="6" pageNumber="86">
Anterior nostril pierced in front side of a pointed flap-like tube. Posterior nostril adjacent to anterior one. Mouth U-shaped, gape about 1.5 times wider than long (
<figureCitation id="136B2A3FFFBEFF85FDA5C9728DBEFC58" box="[623,686,893,917]" captionStart="Fig" captionStartId="4.[113,149,651,673]" captionTargetBox="[113,752,189,636]" captionTargetId="figure-49@4.[113,752,189,636]" captionTargetPageId="4" captionText="Fig. 3. Malihkaia aligera, CMK 25508, 73.0 mm SL; ventral view of head." figureDoi="http://doi.org/10.5281/zenodo.4504501" httpUri="https://zenodo.org/record/4504501/files/figure.png" pageId="6" pageNumber="86">Fig. 3</figureCitation>
). Lips thick, with numerous, closely set, deep furrows. Upper lip with a small median notch, with furrows on its whole length, edge crenulated. Processus dentiformis present. Lower lip with a median interruption; with furrows on its whole length, edge crenulated. Tip of lower jaw not exposed. No median concavity in lower jaw. Inner rostral barbel reaching beyond corner of mouth; outer one reaching vertical of middle of eye. Maxillary barbel reaching beyond posterior margin of eye. Intestine with a loop posterior to stomach (
<figureCitation id="136B2A3FFFBEFF85FDBDCE928DA9FB78" box="[631,697,1181,1205]" captionStart="Fig" captionStartId="6.[813,853,1036,1058]" captionTargetBox="[1001,1270,191,1021]" captionTargetId="figure-722@6.[951,1312,176,1024]" captionTargetPageId="6" captionText="Fig. 5. Malihkaia aligera, MHNG 2766.051, holotype, 65.7 mm SL; digestive tract. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4504507" httpUri="https://zenodo.org/record/4504507/files/figure.png" pageId="6" pageNumber="86">Fig. 5</figureCitation>
). Air bladder without visible posterior chamber.
</paragraph>
<caption id="DF2F6632FFBEFF85FCE7CE038A15FBF3" ID-DOI="http://doi.org/10.5281/zenodo.4504507" ID-Zenodo-Dep="4504507" httpUri="https://zenodo.org/record/4504507/files/figure.png" pageId="6" pageNumber="86" startId="6.[813,853,1036,1058]" targetBox="[1001,1270,191,1021]" targetPageId="6">
<paragraph id="8BEF36BAFFBEFF85FCE7CE038A15FBF3" blockId="6.[813,1449,1036,1087]" pageId="6" pageNumber="86">
Fig. 5.
<taxonomicName id="4C504D39FFBEFF85FC4CCE038BA9FBEC" authority=", MHNG" authorityName="MHNG" box="[902,1209,1036,1057]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="86" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBEFF85FC4CCE038B46FBEC" box="[902,1110,1036,1057]" italics="true" pageId="6" pageNumber="86">Malihkaia aligera</emphasis>
, MHNG
</taxonomicName>
2766.051, holotype, 65.7 mm SL; digestive tract. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BEF36BAFFBEFF85FFBBCEF28E3FF998" blockId="6.[112,753,1277,1621]" pageId="6" pageNumber="86">
<emphasis id="B924EAA8FFBEFF85FFBBCEF28E4CFAD8" bold="true" box="[113,348,1277,1301]" pageId="6" pageNumber="86">Sexual dimorphism.</emphasis>
Males with suborbital flap; females with neither suborbital flap nor suborbital slit. Pectoral fin rounded in females, strongly falcate in males. In males, unbranched and first branched rays rigid, arched and curled upwards. First branched ray about 5 times wider than other branched rays, flattened dorso-ventrally, reaching beyond pelvic-fin base; without membrane between the branches and all sub-branches. Second branched ray slender, branched only at tip, membrane between branches very narrow. See diagnosis of genus for more detailed description of pectoral fin of both sexes.
</paragraph>
<paragraph id="8BEF36BAFFBEFF85FFBBCC728DA8F838" blockId="6.[112,753,1661,2037]" pageId="6" pageNumber="86">
<emphasis id="B924EAA8FFBEFF85FFBBCC728FEEF958" bold="true" box="[113,254,1661,1685]" pageId="6" pageNumber="86">Coloration.</emphasis>
In formalin, shortly after fixation. Body background colour pale yellowish brown, throat and belly whitish; except otherwise stated, all markings blackish brown to black. Head black, cheek marmorated, lower side whitish. Irregularly set and shaped patches of black pigments on lips, but always a patch on posterior edge of lower lip and on adjacent part of throat. Body with 912 bars (34 predorsal, 3 subdorsal, 35 postdorsal), extending from dorsal midline to level of pectoral fin (or ventral midline on caudal peduncle), mostly continuous over back with contralaterals; bars of quite regular width and shape in front of dorsal fin, less regular posteriorly; much wider than interspaces.
</paragraph>
<paragraph id="8BEF36BAFFBEFF85FCE7CE798B62FA23" blockId="6.[812,1453,1142,1518]" pageId="6" pageNumber="86">A conspicuous more or less squarish black blotch at middle of caudal-fin base, depth about ⅓–¼ of depth of base of fin. A more or less triangular black blotch over dorsal procurrent rays, including dorsal midline. Fainter pigments over base of uppermost and lowermost 46 principal rays of caudal fin, making a slightly elongated, arched mark, variably contrasted, and variably connected with other two blotches. Space between upper triangular blotch and last bar on body somewhat reddish. Axial stripe faint, or not distinct except on caudal peduncle where darker and wider, combining with blotch at middle of caudal-fin base to make a larger and more conspicuous blotch.</paragraph>
<paragraph id="8BEF36BAFFBEFF85FCE7CC198A90F8C3" blockId="6.[812,1452,1558,1806]" pageId="6" pageNumber="86">
Dorsal fin hyaline, no pigments on membranes, with a small black spot at base of unbranched rays and middle and distal parts of last unbranched ray black; pigments on rays at branching points and on middle of unbranched part of first branched rays (and branched rays
<quantity id="4CA89B5FFFBEFF85FB5DCC998BCAF963" box="[1175,1242,1686,1710]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="6" pageNumber="86" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">24 in</quantity>
largest specimens). Caudal fin hyaline, with pigments between segments of branched rays and near branching points. Anal fin hyaline, with pigments near branching points of branched rays.
</paragraph>
<paragraph id="8BEF36BAFFBEFF85FCE7CD398B57F7C0" blockId="6.[811,1452,1846,2062]" pageId="6" pageNumber="86">Pelvic fin hyaline, with pigments along rays and near branching points of branched rays. Pectoral fin with membranes hyalines, pigments on rays, darker along posterior edge of branched rays 210. In males, unbranched and first branched rays more or less completely covered by pigments, on dorsal side, on whole length; identical in females, but pigments less densely set.</paragraph>
<paragraph id="8BEF36BAFFBFFF84FF4DCAB28DD8FF38" blockId="7.[134,774,189,245]" pageId="7" pageNumber="87">
<emphasis id="B924EAA8FFBFFF84FF4DCAB28E56FF18" bold="true" box="[135,326,189,213]" pageId="7" pageNumber="87">Notes on biology.</emphasis>
The two females did not have ripe ovaries. The stomach contained unidentifiable insect remains.
</paragraph>
</subSubSection>
<subSubSection id="C34A6531FFBFFF84FF4DCB128D5BFD78" pageId="7" pageNumber="87" type="distribution">
<paragraph id="8BEF36BAFFBFFF84FF4DCB128D5BFD78" blockId="7.[135,775,285,693]" pageId="7" pageNumber="87">
<emphasis id="B924EAA8FFBFFF84FF4DCB128EAAFEF8" bold="true" box="[135,442,285,309]" pageId="7" pageNumber="87">Distribution and habitat.</emphasis>
<taxonomicName id="4C504D39FFBFFF84FE0CCB128D85FEF8" box="[454,661,285,309]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBFFF84FE0CCB128D85FEF8" box="[454,661,285,309]" italics="true" pageId="7" pageNumber="87">Malihkaia aligera</emphasis>
</taxonomicName>
has been observed only once, in the
<collectingCountry id="F347762AFFBFFF84FE0CCB328EEAFE98" box="[454,506,317,341]" name="Serbia" pageId="7" pageNumber="87">Mali</collectingCountry>
Hka River near Putao. It was collected over gravel and cobble, in fast riffles, in about
<quantity id="4CA89B5FFFBFFF84FF06CB728E01FE59" box="[204,273,381,405]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="87" unit="cm" value="60.0">60 cm</quantity>
depth (
<figureCitation id="136B2A3FFFBFFF84FEACCB728EBBFE58" box="[358,427,381,405]" captionStart="Fig" captionStartId="7.[835,870,580,602]" captionTargetBox="[835,1474,189,565]" captionTargetId="figure-148@7.[835,1474,189,565]" captionTargetPageId="7" captionText="Fig. 6. Type locality of Malihkaia aligera and Schistura nubigena; Myanmar: Kachin state: Mali Hka River near Kang Mu Lon; 402 masl; 26 November 2014." figureDoi="http://doi.org/10.5281/zenodo.4504509" httpUri="https://zenodo.org/record/4504509/files/figure.png" pageId="7" pageNumber="87">Fig. 6</figureCitation>
). Other species collected in the same microhabitat at this locality include
<taxonomicName id="4C504D39FFBFFF84FDBBCB918DC5FE78" authorityName="Hamilton" authorityYear="1822" box="[625,725,414,437]" class="Actinopterygii" family="Cyprinidae" genus="Bangana" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="B924EAA8FFBFFF84FDBBCB918DC5FE78" box="[625,725,414,437]" italics="true" pageId="7" pageNumber="87">Bangana</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C504D39FFBFFF84FF4DCBB28FDCFE19" box="[135,204,445,468]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="B924EAA8FFBFFF84FF4DCBB28FDCFE19" box="[135,204,445,468]" italics="true" pageId="7" pageNumber="87">Garra</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C504D39FFBFFF84FEC1CBB28E9EFE18" box="[267,398,445,469]" class="Actinopterygii" family="Cyprinidae" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="family">Cyprinidae</taxonomicName>
),
<taxonomicName id="4C504D39FFBFFF84FE6FCBB28E57FE38" authorityName="Conway &amp; Britz" authorityYear="2010" class="Actinopterygii" family="Psilorhynchidae" genus="Psilorhynchus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="brachyrhynchus">
<emphasis id="B924EAA8FFBFFF84FE6FCBB28C15FE18" box="[421,773,445,469]" italics="true" pageId="7" pageNumber="87">Psilorhynchus brachyrhynchus</emphasis>
(Psilorhynchidae)
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FE99CBD28C12FE38" baseAuthorityName="Prashad &amp; Mukerji" baseAuthorityYear="1929" box="[339,770,477,501]" class="Actinopterygii" family="Balitoridae" genus="Homalopteroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="rupicola">
<emphasis id="B924EAA8FFBFFF84FE99CBD28D7FFE38" box="[339,623,477,501]" italics="true" pageId="7" pageNumber="87">Homalopteroides rupicola</emphasis>
(Balitoridae)
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FF43CBF28E22FDD8" authorityName="Peters" authorityYear="1861" box="[137,306,509,533]" class="Actinopterygii" family="Nemacheilidae" genus="Acanthocobitis" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="genus">
<emphasis id="B924EAA8FFBFFF84FF43CBF28E22FDD8" box="[137,306,509,533]" italics="true" pageId="7" pageNumber="87">Acanthocobitis</emphasis>
</taxonomicName>
sp.,
<taxonomicName id="4C504D39FFBFFF84FEA4CBF28D53FDD8" authorityName="Kottelat" authorityYear="1990" box="[366,579,509,533]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="malaisei">
<emphasis id="B924EAA8FFBFFF84FEA4CBF28D53FDD8" box="[366,579,509,533]" italics="true" pageId="7" pageNumber="87">Schistura malaisei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FD9CCBF28DCEFDD8" authorityName="Kottelat" authorityYear="2017" box="[598,734,509,533]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="nubigena">
<emphasis id="B924EAA8FFBFFF84FD9CCBF28DCEFDD8" box="[598,734,509,533]" italics="true" pageId="7" pageNumber="87">S. nubigena</emphasis>
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FD3BCBF28EAFFDF8" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="sikmaiensis">
<emphasis id="B924EAA8FFBFFF84FD3BCBF28E12FDF8" italics="true" pageId="7" pageNumber="87">S. sikmaiensis</emphasis>
(Nemacheilidae)
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FE04C8118C11FDF8" authorityName="Ng" authorityYear="2008" box="[462,769,541,565]" class="Actinopterygii" family="Bagridae" genus="Batasio" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="procerus">
<emphasis id="B924EAA8FFBFFF84FE04C8118D97FDF8" box="[462,647,542,565]" italics="true" pageId="7" pageNumber="87">Batasio procerus</emphasis>
(Bagridae)
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FF43C8328D4CFD98" authorityName="Chaudhuri" authorityYear="1919" box="[137,604,573,597]" class="Actinopterygii" family="Amblycipitidae" genus="Amblyceps" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="murraystuarti">
<emphasis id="B924EAA8FFBFFF84FF43C8328E8AFD98" box="[137,410,573,597]" italics="true" pageId="7" pageNumber="87">Amblyceps murraystuarti</emphasis>
(Amblycipitidae)
</taxonomicName>
,
<taxonomicName id="4C504D39FFBFFF84FDA3C8328E8EFDB8" authorityName="L.i.Zhou &amp; Yang" authorityYear="2008" class="Actinopterygii" family="Sisoridae" genus="Pseudecheneis" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="brachyurus">
<emphasis id="B924EAA8FFBFFF84FDA3C8328E1EFDB8" italics="true" pageId="7" pageNumber="87">Pseudecheneis brachyurus</emphasis>
(Sisoridae)
</taxonomicName>
and
<taxonomicName id="4C504D39FFBFFF84FE28C8528E44FD58" class="Actinopterygii" family="Mastacembelidae" genus="Mastacembelus" kingdom="Animalia" order="Synbranchiformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="armatus">
<emphasis id="B924EAA8FFBFFF84FE28C8528C16FDB8" box="[482,774,605,629]" italics="true" pageId="7" pageNumber="87">Mastacembelus armatus</emphasis>
(Mastacembelidae)
</taxonomicName>
.
<taxonomicName id="4C504D39FFBFFF84FE97C8728D0EFD58" authorityName="Kottelat" authorityYear="1990" box="[349,542,637,661]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="malaisei">
<emphasis id="B924EAA8FFBFFF84FE97C8728D0EFD58" box="[349,542,637,661]" italics="true" pageId="7" pageNumber="87">Schistura malaisei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D39FFBFFF84FD87C8728FF1FD78" baseAuthorityName="Prashad &amp; Mukerji" baseAuthorityYear="1929" class="Actinopterygii" family="Balitoridae" genus="Homalopteroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="rupicola">
<emphasis id="B924EAA8FFBFFF84FD87C8728FF1FD78" italics="true" pageId="7" pageNumber="87">Homalopteroides rupicola</emphasis>
</taxonomicName>
were the most abundant species.
</paragraph>
</subSubSection>
<caption id="DF2F6632FFBFFF84FC89C84B8B54FD5E" ID-DOI="http://doi.org/10.5281/zenodo.4504509" ID-Zenodo-Dep="4504509" httpUri="https://zenodo.org/record/4504509/files/figure.png" pageId="7" pageNumber="87" startId="7.[835,870,580,602]" targetBox="[835,1474,189,565]" targetPageId="7">
<paragraph id="8BEF36BAFFBFFF84FC89C84B8B54FD5E" blockId="7.[835,1474,580,660]" pageId="7" pageNumber="87">
Fig. 6. Type locality of
<taxonomicName id="4C504D39FFBFFF84FBEFC84B8BC2FD94" box="[1061,1234,580,601]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBFFF84FBEFC84B8BC2FD94" box="[1061,1234,580,601]" italics="true" pageId="7" pageNumber="87">Malihkaia aligera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C504D39FFBFFF84FACBC84B8AAAFD94" authorityName="Kottelat" authorityYear="2017" box="[1281,1466,580,601]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="nubigena">
<emphasis id="B924EAA8FFBFFF84FACBC84B8AAAFD94" box="[1281,1466,580,601]" italics="true" pageId="7" pageNumber="87">Schistura nubigena</emphasis>
</taxonomicName>
; Myanmar: Kachin state: Mali Hka River near Kang Mu Lon; 402 masl; 26 November 2014.
</paragraph>
</caption>
<subSubSection id="C34A6531FFBFFF84FF4DC8D28E66FCD8" pageId="7" pageNumber="87" type="etymology">
<paragraph id="8BEF36BAFFBFFF84FF4DC8D28E66FCD8" blockId="7.[135,774,733,789]" pageId="7" pageNumber="87">
<emphasis id="B924EAA8FFBFFF84FF4DC8D28E1FFD38" bold="true" box="[135,271,733,757]" pageId="7" pageNumber="87">Etymology.</emphasis>
From the Latin
<emphasis id="B924EAA8FFBFFF84FE1FC8D28D08FD38" box="[469,536,733,757]" italics="true" pageId="7" pageNumber="87">aliger</emphasis>
(
<taxonomicName id="4C504D39FFBFFF84FDE7C8D28D6EFD38" box="[557,638,733,757]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBFFF84FDE7C8D28D6EFD38" box="[557,638,733,757]" italics="true" pageId="7" pageNumber="87">aligera</emphasis>
</taxonomicName>
,
<emphasis id="B924EAA8FFBFFF84FD5AC8D28DE5FD38" box="[656,757,733,757]" italics="true" pageId="7" pageNumber="87">aligerum</emphasis>
), winged. An adjective.
</paragraph>
</subSubSection>
<subSubSection id="C34A6531FFBFFF8BFF4DC9328DF5FE99" lastPageId="8" lastPageNumber="88" pageId="7" pageNumber="87" type="discussion">
<paragraph id="8BEF36BAFFBFFF8BFF4DC9328DF5FE99" blockId="7.[135,774,829,885]" lastBlockId="8.[113,755,188,341]" lastPageId="8" lastPageNumber="88" pageId="7" pageNumber="87">
<emphasis id="B924EAA8FFBFFF84FF4DC9328FE6FC98" bold="true" box="[135,246,829,853]" pageId="7" pageNumber="87">Remarks.</emphasis>
The colour pattern of the six available specimens of
<taxonomicName id="4C504D39FFBFFF84FF6DC9518E0BFCB8" box="[167,283,861,885]" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBFFF84FF6DC9518E0BFCB8" box="[167,283,861,885]" italics="true" pageId="7" pageNumber="87">M. aligera</emphasis>
</taxonomicName>
is made of very regular bars on body, with very well marked edges, on a pale background. This general pattern is rare among Southeast Asian nemacheilids. Interestingly, one of the species collected together with
<taxonomicName id="4C504D39FFBFFF84FA6DC9038C80FC8E" class="Actinopterygii" family="Nemacheilidae" genus="Malihkaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="aligera">
<emphasis id="B924EAA8FFBFFF84FA6DC9038C80FC8E" italics="true" pageId="7" pageNumber="87">M. aligera</emphasis>
</taxonomicName>
is
<taxonomicName id="4C504D39FFBFFF84FC61C9248B9EFC8E" box="[939,1166,811,835]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="sikmaiensis">
<emphasis id="B924EAA8FFBFFF84FC61C9248B9EFC8E" box="[939,1166,811,835]" italics="true" pageId="7" pageNumber="87">Schistura sikmaiensis</emphasis>
</taxonomicName>
, which also has a regular and contrasted colour pattern (
<figureCitation id="136B2A3FFFBFFF84FB94C9448BB1FCAE" box="[1118,1185,843,867]" captionStart="Fig" captionStartId="7.[136,171,2093,2115]" captionTargetBox="[274,1336,951,2077]" captionTargetId="figure-200@7.[274,1336,951,2077]" captionTargetPageId="7" captionText="Fig. 7. Schistura sikmaiensis, CMK 25560; Myanmar: Irrawaddy drainage: Mula River at Lika; a, female, 61.6 mm SL; b, male, 72.7 mm SL." figureDoi="http://doi.org/10.5281/zenodo.4504513" httpUri="https://zenodo.org/record/4504513/files/figure.png" pageId="7" pageNumber="87">Fig. 7</figureCitation>
). A third species collected at the same locality is
<taxonomicName id="4C504D39FFBFFF84FBFAC9638BBDFC4E" authorityName="Kottelat" authorityYear="2017" box="[1072,1197,875,899]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="87" phylum="Chordata" rank="species" species="nubigena">
<emphasis id="B924EAA8FFBFFF84FBFAC9638BBDFC4E" box="[1072,1197,875,899]" italics="true" pageId="7" pageNumber="87">S. nubigena</emphasis>
</taxonomicName>
(described below), which has a sharply contrasted colour pattern of 4 bold black bars on a whitish background. Finally,
<taxonomicName id="4C504D39FFB0FF8BFDCBCAD28DA1FF39" authorityName="Kottelat" authorityYear="2017" box="[513,689,220,244]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="88" phylum="Chordata" rank="species" species="wanlainensis">
<emphasis id="B924EAA8FFB0FF8BFDCBCAD28DA1FF39" box="[513,689,220,244]" italics="true" pageId="8" pageNumber="88">S. wanlainensis</emphasis>
</taxonomicName>
, also described below, from the same area but at higher altitude and habitat with higher gradient, also has a very contrasted colour pattern of very regular bars on a pale background.
</paragraph>
</subSubSection>
</treatment>
</document>