179 lines
18 KiB
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179 lines
18 KiB
XML
<document ID-DOI="10.1111/zoj.12193" ID-ISSN="0024-4082" ID-Zenodo-Dep="5330460" approvalRequired="150" approvalRequired_for_taxonomicNames="66" approvalRequired_for_textStreams="58" approvalRequired_for_treatments="26" checkinTime="1630167834076" checkinUser="felipe" docAuthor="Norman, David B." docDate="2015" docId="03F9879B3208FFC8FC94FDCEFED67B25" docLanguage="en" docName="zoj12193.pdf" docOrigin="Zoological Journal of the Linnean Society 173 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12193" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.1:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="1" docTitle="Gilmoreosaurus Brett-Surmann 1979" docType="treatment" docVersion="3" lastPageNumber="162" masterDocId="FFC0FFE3324DFF8EFFBDFFADFFCD7D65" masterDocTitle="On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna)" masterLastPageNumber="189" masterPageNumber="92" pageNumber="161" updateTime="1630625843961" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna)</mods:title>
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</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Norman, David B.</mods:namePart>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-01-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>173</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>92</mods:start>
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<mods:location>
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<mods:url>https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12193</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1111/zoj.12193</mods:identifier>
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<mods:identifier type="ISSN">0024-4082</mods:identifier>
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<mods:identifier type="Zenodo-Dep">5330460</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03F9879B3208FFC8FC94FDCEFED67B25" httpUri="http://treatment.plazi.org/id/03F9879B3208FFC8FC94FDCEFED67B25" lastPageId="70" lastPageNumber="162" pageId="69" pageNumber="161">
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<subSubSection pageId="69" pageNumber="161" type="nomenclature">
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<paragraph blockId="69.[809,1418,611,665]" pageId="69" pageNumber="161">
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<heading centered="true" fontSize="8" level="2" pageId="69" pageNumber="161" reason="8">
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<emphasis box="[809,1195,611,632]" italics="true" pageId="69" pageNumber="161">
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<taxonomicName authorityName="Brett-Surmann" authorityYear="1979" box="[809,1018,611,632]" class="Reptilia" family="Hadrosauridae" genus="Gilmoreosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">GILMOREOSAURUS</taxonomicName>
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MONGOLIENSIS
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</emphasis>
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(
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<bibRefCitation author="Gilmore CW" box="[1211,1388,611,633]" pageId="69" pageNumber="161" pagination="23 - 78" refId="ref61143" refString="Gilmore CW. 1933. On the dinosaurian fauna of the Iren Dabasu Formation. Bulletin of the American Museum of Natural History 67: 23 - 78." type="journal article" year="1933">GILMORE, 1933</bibRefCitation>
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) – (PRIETO- MÁRQUEZ & NORELL, 2010)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="69" pageNumber="161" type="reference_group">
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<paragraph blockId="69.[806,1422,681,887]" pageId="69" pageNumber="161">
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<taxonomicName authorityName="Brett-Surmann" authorityYear="1979" box="[806,997,681,702]" class="Reptilia" family="Hadrosauridae" genus="Gilmoreosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[806,997,681,702]" italics="true" pageId="69" pageNumber="161">Gilmoreosaurus</emphasis>
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</taxonomicName>
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comprises the partial remains of four individuals of a hadrosauromorph collected from the Iren Dabasu Formation, Erenhot,
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<collectingCountry box="[1343,1418,743,765]" name="China" pageId="69" pageNumber="161">China</collectingCountry>
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: Turonian−Coniacian (
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<bibRefCitation author="Sues H-D & Averianov A" box="[1042,1303,773,795]" pageId="69" pageNumber="161" pagination="2549 - 2555" refId="ref64886" refString="Sues H-D, Averianov A. 2009. A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs. Proceedings of the Royal Society of London. Series B 276: 2549 - 2555." type="journal article" year="2009">Sues & Averianov, 2009</bibRefCitation>
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). However, it should (again) be noted that estimates of the age of these beds have ranged from Albian to Maastrichtian (
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<bibRefCitation author="Prieto-Marquez A" box="[814,1080,865,887]" pageId="69" pageNumber="161" pagination="773 - 792" refId="ref64053" refString="Prieto-Marquez A. 2011 a. Cranial and appendicular ontogeny of Bactrosaurus johnsoni, a hadrosauroid dinosaur from the late Cretaceous of northern China. Palaeontology 54: 773 - 792." type="journal article" year="2011">Prieto-Márquez, 2011a</bibRefCitation>
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).
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</paragraph>
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<paragraph blockId="69.[806,1421,928,1348]" box="[806,979,928,950]" pageId="69" pageNumber="161">
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<emphasis box="[806,979,928,950]" italics="true" pageId="69" pageNumber="161">Teeth and jaws</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="70" lastPageNumber="162" pageId="69" pageNumber="161" type="description">
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<paragraph blockId="69.[806,1421,928,1348]" pageId="69" pageNumber="161">
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The maxillary crowns are narrower than the dentary crowns, but have a more prominent median primary ridge. The dentary crowns have a single median (or submedian) lower primary ridge. Neither dentary nor maxillary crowns appear to have accessory ridges. The marginal denticles are also ledge-like and bear mammillae [these were also reported to be present in
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<taxonomicName authorityName="Head" authorityYear="1998" box="[806,943,1173,1194]" class="Reptilia" family="Ornithischia (awaiting allocation)" genus="Protohadros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[806,943,1173,1194]" italics="true" pageId="69" pageNumber="161">Protohadros</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Langston" authorityYear="1960" box="[958,1115,1173,1194]" class="Reptilia" family="Hadrosauridae" genus="Lophorhothon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[958,1115,1173,1194]" italics="true" pageId="69" pageNumber="161">Lophorhothon</emphasis>
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</taxonomicName>
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, and some lambeosaurines (
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<bibRefCitation author="Prieto-Marquez A & Norell MA" box="[813,1187,1203,1226]" pageId="69" pageNumber="161" pagination="1 - 49" refId="ref64180" refString="Prieto-Marquez A, Norell MA. 2010. Anatomy and relationships of Gilmoreosaurus mongoliensis (Dinosauria: Hadrosauroidea) from the Late Cretaceous of Central Asia. American Museum Novitates 3694: 1 - 49." type="journal article" year="2010">Prieto-Márquez & Norell, 2010: 18</bibRefCitation>
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)]. The dentary ramus is imperfectly known, but differs very little from that described in
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<taxonomicName box="[962,1116,1265,1286]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[962,1116,1265,1286]" italics="true" pageId="69" pageNumber="161">Bactrosaurus</emphasis>
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</taxonomicName>
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, its sympatric contemporary. This dental morphology is distinct from that seen in
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<taxonomicName baseAuthorityName="NHMUK R" baseAuthorityYear="1831" box="[839,952,1326,1348]" class="Reptilia" family="Styracosterna incertae sedis" genus="Hypselospinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="species" species="fittoni">
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<emphasis box="[839,952,1326,1348]" italics="true" pageId="69" pageNumber="161">Hy. fittoni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="69.[806,1422,1389,1503]" box="[806,966,1389,1410]" pageId="69" pageNumber="161">
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<emphasis box="[806,966,1389,1410]" italics="true" pageId="69" pageNumber="161">Axial skeleton</emphasis>
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</paragraph>
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<paragraph blockId="69.[806,1422,1389,1503]" pageId="69" pageNumber="161">
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The axial skeleton is very similar to that described in
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<taxonomicName box="[806,958,1451,1472]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[806,958,1451,1472]" italics="true" pageId="69" pageNumber="161">Bactrosaurus</emphasis>
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</taxonomicName>
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, and displays no distinct characters of significance.
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</paragraph>
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<paragraph blockId="69.[806,1422,1544,1903]" box="[806,1059,1544,1565]" pageId="69" pageNumber="161">
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<emphasis box="[806,1059,1544,1565]" italics="true" pageId="69" pageNumber="161">Appendicular skeleton</emphasis>
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</paragraph>
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<paragraph blockId="69.[806,1422,1544,1903]" lastBlockId="70.[164,779,198,342]" lastPageId="70" lastPageNumber="162" pageId="69" pageNumber="161">
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The pectoral girdle and forelimb are very similar in morphology to those described above for
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<taxonomicName box="[1265,1414,1606,1627]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[1265,1414,1606,1627]" italics="true" pageId="69" pageNumber="161">Bactrosaurus</emphasis>
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</taxonomicName>
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. The ilium differs in the more posterior positioning of the transverse expansion of the dorsal iliac blade when compared to that of
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<taxonomicName box="[1049,1201,1698,1719]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[1049,1201,1698,1719]" italics="true" pageId="69" pageNumber="161">Bactrosaurus</emphasis>
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</taxonomicName>
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, and in the development of a bar-like postacetabular process. The pubis has a prepubic process that is less expanded proximally, the distal expansion is less extreme, and the process overall appears to be longer than that seen in
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<taxonomicName box="[837,986,1851,1872]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="69" pageNumber="161" phylum="Chordata" rank="genus">
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<emphasis box="[837,986,1851,1872]" italics="true" pageId="69" pageNumber="161">Bactrosaurus</emphasis>
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</taxonomicName>
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. The remaining elements of the pelvis and hindlimb seem indistinguishable in these two taxa; however, the unguals of the pes are notable narrower and taper to a bluntly truncated tip, rather than been broad and rounded as in the case of those described for
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<taxonomicName box="[310,455,290,311]" class="Reptilia" family="Lambeosauridae" genus="Bactrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="70" pageNumber="162" phylum="Chordata" rank="genus">
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<emphasis box="[310,455,290,311]" italics="true" pageId="70" pageNumber="162">Bactrosaurus</emphasis>
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</taxonomicName>
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(cf.
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<bibRefCitation author="Prieto-Marquez A & Norell MA" pageId="70" pageNumber="162" pagination="1 - 49" refId="ref64180" refString="Prieto-Marquez A, Norell MA. 2010. Anatomy and relationships of Gilmoreosaurus mongoliensis (Dinosauria: Hadrosauroidea) from the Late Cretaceous of Central Asia. American Museum Novitates 3694: 1 - 49." type="journal article" year="2010">Prieto-Márquez & Norell, 2010</bibRefCitation>
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: fig. 18 and
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<bibRefCitation author="Godefroit P & Dong Z & Bultynk P & Li H & Feng L" box="[367,614,320,342]" pageId="70" pageNumber="162" pagination="3 - 70" refId="ref61214" refString="Godefroit P, Dong Z, Bultynk P, Li H, Feng L. 1998. New Bactrosaurus (Dinosauria: Hadrosauroidea) material from Iren Dabasu (Inner Mongolia, P. R. China). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique. Sciences de la Terre 68: 3 - 70." type="journal article" year="1998">
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Godefroit
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<emphasis box="[483,532,320,342]" italics="true" pageId="70" pageNumber="162">et al</emphasis>
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., 1998
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</bibRefCitation>
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: pl. 14).
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</paragraph>
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<paragraph blockId="70.[260,682,399,452]" pageId="70" pageNumber="162">
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<emphasis box="[260,618,399,420]" italics="true" pageId="70" pageNumber="162">
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<taxonomicName box="[260,500,399,420]" class="Reptilia" family="Iguanodontia incertae sedis" genus="Shuangmiaosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="70" pageNumber="162" phylum="Chordata" rank="genus">SHUANGMIAOSAURUS</taxonomicName>
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GILMOREI
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</emphasis>
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<bibRefCitation author="You H-L & Ji Q & Li J & Li Y" pageId="70" pageNumber="162" pagination="148 - 154" refId="ref65786" refString="You H-L, Ji Q, Li J, Li Y. 2003 a. A new hadrosauroid dinosaur from the mid-Cretaceous of Liaoning Province, China. Acta Geologica Sinica 77: 148 - 154." type="journal article" year="2003">YOU, JI, LI & LI, 2003</bibRefCitation>
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A
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</paragraph>
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<paragraph blockId="70.[164,779,469,827]" pageId="70" pageNumber="162">
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This taxon is represented by a few cranial elements collected from the Sunjiawan Formation (‘middle’ Cretaceous), Beipiao,
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<collectingRegion box="[375,479,530,552]" country="China" name="Liaoning" pageId="70" pageNumber="162">Liaoning</collectingRegion>
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,
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<collectingCountry box="[493,566,530,552]" name="China" pageId="70" pageNumber="162">China</collectingCountry>
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. The specimens, a maxilla plus articulated lacrimal and an edentulous dentary, were not associated and show evidence of post-mortem distortion, which may have contributed to the way in which its anatomy has been described and interpreted. In systematic analyses
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<bibRefCitation author="You H-L & Ji Q & Li J & Li Y" box="[594,778,683,705]" pageId="70" pageNumber="162" pagination="148 - 154" refId="ref65786" refString="You H-L, Ji Q, Li J, Li Y. 2003 a. A new hadrosauroid dinosaur from the mid-Cretaceous of Liaoning Province, China. Acta Geologica Sinica 77: 148 - 154." type="journal article" year="2003">
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You
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<emphasis box="[641,686,683,705]" italics="true" pageId="70" pageNumber="162">et al</emphasis>
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. (2003a)
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</bibRefCitation>
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placed this taxon as the sister taxon to the
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<taxonomicName box="[164,336,744,766]" class="Reptilia" family="Hadrosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="70" pageNumber="162" phylum="Chordata" rank="family">Hadrosauridae</taxonomicName>
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(= Euhadrosauria
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<emphasis box="[563,626,745,766]" italics="true" pageId="70" pageNumber="162">sensu</emphasis>
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<bibRefCitation author="Weishampel DB & Norman DB & Grigorescu D" pageId="70" pageNumber="162" pagination="361 - 385" refId="ref65483" refString="Weishampel DB, Norman DB, Grigorescu D. 1993. Telmatosaurus transsylvanicus from the Late Cretaceous of Romania: the most basal hadrosaurid dinosaur. Palaeontology 36: 361 - 385." type="journal article" year="1993">
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Weishampel
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<emphasis box="[164,209,775,796]" italics="true" pageId="70" pageNumber="162">et al</emphasis>
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., 1993
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</bibRefCitation>
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;
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<bibRefCitation author="Norman DB" box="[292,447,775,797]" pageId="70" pageNumber="162" pagination="10 - 43" refId="ref63328" refString="Norman DB. 2014. Iguanodonts from the Wealden of England: do they contribute to the discussion concerning hadrosaur origins? In: Evans D, Eberth D, eds. Hadrosaurs. Bloomington: Indiana University Press, 10 - 43." type="book chapter" year="2014">Norman, 2014</bibRefCitation>
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) and
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<collectingCountry box="[507,617,775,797]" name="Heard Island and McDonald Islands" pageId="70" pageNumber="162">McDonald</collectingCountry>
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(2012b) placed it at just one further step removed.
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</paragraph>
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<paragraph blockId="70.[164,780,870,1504]" box="[164,337,870,892]" pageId="70" pageNumber="162">
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<emphasis box="[164,337,870,892]" italics="true" pageId="70" pageNumber="162">Teeth and jaws</emphasis>
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</paragraph>
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<paragraph blockId="70.[164,780,870,1504]" pageId="70" pageNumber="162">Only maxillary crowns are known and exhibit a lanceolate shape, have a single median primary ridge and no accessory ridges, and the mesial and distal margins of the crowns bear denticles. The dentary is very elongate and slightly arched along its length; it also appears to have had a relatively short diastema. The medial surface of the dentary ramus shows a deep and elongate alveolar trough that is backed by sets of alveolar grooves that appear to show the outlines of replacement crowns, rather than forming consistent parallel troughs (this is a nonhadrosauromorph characteristic). The alveolar trough extends back toward the posterior margin of the base of the coronoid process. The coronoid process is elongate but appears to form an obtuse angle to the long axis of the dentary ramus. The extent of post-mortem distortion in this specimen (which is clearly evident in the maxilla that is described) makes it difficult to discern genuine and unique anatomy from structures that may simply reflect postburial distortion.</paragraph>
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<paragraph blockId="70.[164,546,1547,1600]" box="[164,546,1547,1568]" pageId="70" pageNumber="162">
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<emphasis box="[164,546,1547,1568]" italics="true" pageId="70" pageNumber="162">Axial and appendicular skeletons</emphasis>
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</paragraph>
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<paragraph blockId="70.[164,546,1547,1600]" box="[164,283,1578,1600]" pageId="70" pageNumber="162">Unknown.</paragraph>
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</subSubSection>
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</treatment>
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</document> |