251 lines
29 KiB
XML
251 lines
29 KiB
XML
<document id="D91756D54013351CDCC4535422BE6136" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF9AE8160D068D2227A8FB8C" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia lapidini Liew & Schilthuizen & Vermeulen 2009, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="536" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="536" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="15A7BDD3122E4D52E6987CB61815C158">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:namePart id="ED81B1AB958680715B1997F45338D017">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="0D63BC97FF29765804C7D64BBBBC8052">Schilthuizen, Menno</mods:namePart>
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<mods:roleTerm id="0EE6A213EE8782762A31E1BBDA630D78">Author</mods:roleTerm>
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<mods:namePart id="0220286283071AD8698F3272E5E4F6ED">Vermeulen, Jaap Jan</mods:namePart>
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<mods:typeOfResource id="697861C88CEE9789DFF7FF9A69757C88">text</mods:typeOfResource>
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<mods:title id="61B4522A49C1F601F55BA57723668EAC">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="D1FB22085972A0F35ED832B43DC91212">
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<mods:date id="FB8D3DB1B3199AE8E85A108A7F2B03E9">2009</mods:date>
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<mods:number id="58CDB6B68EFFC10A9DB75EBAF4D791C6">2009-10-26</mods:number>
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<mods:number id="6DA8A59C1B448ACD77F870910577E0A5">157</mods:number>
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<mods:number id="214633D450C1B83CFE6E80E41C238068">3</mods:number>
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<mods:identifier id="1346E25402B31D73E2A3F3ADCA0C58A5" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
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<mods:identifier id="75BDCF37207AC21492315BE69046B11A" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F9878AFF9AE8160D068D2227A8FB8C" ID-DOI="http://doi.org/10.5281/zenodo.5492191" ID-Zenodo-Dep="5492191" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8160D068D2227A8FB8C" httpUri="http://treatment.plazi.org/id/03F9878AFF9AE8160D068D2227A8FB8C" lastPageNumber="536" pageId="21" pageNumber="536">
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<subSubSection id="C34A6517FF9AE8160D068D2221ECFE8C" box="[293,649,305,326]" pageId="21" pageNumber="536" type="nomenclature">
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<paragraph id="8BEF369CFF9AE8160D068D2221ECFE8C" blockId="21.[293,649,305,326]" box="[293,649,305,326]" pageId="21" pageNumber="536">
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<heading id="D0A781F0FF9AE8160D068D2221ECFE8C" box="[293,649,305,326]" centered="true" fontSize="9" level="2" pageId="21" pageNumber="536" reason="2">
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<taxonomicName id="4C504D1FFF9AE8160D068D22217EFE8F" ID-CoL="6HLW3" authority="Liew & Schilthuizen & Vermeulen, 2009" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[293,539,305,326]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="lapidini" status="sp. nov.">EVERETTIA LAPIDINI</taxonomicName>
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<emphasis id="B924EA8EFF9AE8160E078D2721ECFE8C" bold="true" box="[548,649,305,326]" pageId="21" pageNumber="536">
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<taxonomicNameLabel id="A21757F5FF9AE8160E078D2721ECFE8C" box="[548,649,305,326]" pageId="21" pageNumber="536" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6517FF9AE8160C878D4B2107FD8F" pageId="21" pageNumber="536" type="reference_group">
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<paragraph id="8BEF369CFF9AE8160C878D4B211DFEA6" blockId="21.[164,632,343,365]" box="[164,632,343,365]" pageId="21" pageNumber="536">
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<treatmentCitationGroup id="AB4011B2FF9AE8160C878D4B211DFEA6" box="[164,632,343,365]" pageId="21" pageNumber="536">
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<treatmentCitation id="0AF1108DFF9AE8160C878D4B2111FEA6" author="Laidlaw FF" box="[164,628,343,365]" page="180" pageId="21" pageNumber="536" year="1937">
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<taxonomicName id="4C504D1FFF9AE8160C878D4B2111FEA6" ID-CoL="3DHB4" authority=", Laidlaw, 1937: 180" authorityName=", Laidlaw" authorityPageNumber="180" authorityYear="1937" box="[164,628,343,365]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="subconsul">
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<emphasis id="B924EA8EFF9AE8160C878D4B22E0FEA6" box="[164,389,343,365]" italics="true" pageId="21" pageNumber="536">Everettia subconsul</emphasis>
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,
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<bibRefCitation id="EFC14B6DFF9AE8160DB68D442111FEA6" author="Laidlaw FF" box="[405,628,343,365]" pageId="21" pageNumber="536" pagination="177 - 189" refId="ref25632" refString="Laidlaw FF. 1937. Notes on land Mollusca from Mount Kinabalu, British North Borneo. The Bulletin of the Raffles Museum 13: 177 - 189." type="journal article" year="1937">Laidlaw, 1937: 180</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BEF369CFF9AE8160C878D852107FD8F" blockId="21.[164,779,406,581]" pageId="21" pageNumber="536">
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<emphasis id="B924EA8EFF9AE8160C878D85238FFE61" box="[164,234,406,427]" italics="true" pageId="21" pageNumber="536">Types:</emphasis>
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<materialsCitation id="3B383CC1FF9AE8160CD58D85213BFD8F" collectingDate="2007-05-11" collectionCode="BMNH" country="Malaysia" county="State" elevation="1900" latitude="6.0333333" location="Mesilau Resort" longLatPrecision="1305" longitude="116.583336" municipality="Ranau District" pageId="21" pageNumber="536" specimenCode="BMNH 20080636" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<collectingCountry id="F347760CFF9AE8160CD58D85221AFE61" box="[246,383,406,427]" name="Malaysia" pageId="21" pageNumber="536">MALAYSIA</collectingCountry>
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:
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<collectingCounty id="628E4E10FF9AE8160DAC8D8522A9FE61" box="[399,460,406,427]" pageId="21" pageNumber="536">State</collectingCounty>
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of
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<collectingRegion id="4994F87EFF9AE8160DD98D852120FE66" box="[506,581,406,428]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">Sabah</collectingRegion>
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:
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<collectingMunicipality id="6B8BACE6FF9AE8160E768D85206DFE66" box="[597,776,406,428]" pageId="21" pageNumber="536">Ranau District</collectingMunicipality>
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. Southern part of Kinabalu Park. At
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<quantity id="4CA89B79FF9AE8160E5C8DA621B6FE00" box="[639,723,437,458]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="536" unit="m" value="1900.0">
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||
<elevation id="007DD1AFFF9AE8160E5C8DA621B6FE00" box="[639,723,437,458]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="536" unit="m" value="1900.0">1900 m</elevation>
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</quantity>
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alt.
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<location id="8E8F6047FF9AE8160C878DC02231FE22" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8160D068D2227A8FB8C:8E8F6047FF9AE8160C878DC02231FE22" box="[164,340,467,489]" country="Malaysia" county="State" latitude="6.0333333" longLatPrecision="1305" longitude="116.583336" municipality="Ranau District" name="Mesilau Resort" pageId="21" pageNumber="536" stateProvince="Sabah">Mesilau Resort</location>
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(
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<geoCoordinate id="EE64505BFF9AE8160D4B8DC022CCFE22" box="[360,425,466,489]" degrees="6" direction="north" minutes="2" orientation="latitude" pageId="21" pageNumber="536" precision="925" value="6.0333333">
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6°2
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<emphasis id="B924EA8EFF9AE8160DAE8DC122F6FE23" box="[397,403,466,489]" italics="true" pageId="21" pageNumber="536">′</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE64505BFF9AE8160D998DC02144FE22" box="[442,545,466,489]" degrees="116" direction="east" minutes="35" orientation="longitude" pageId="21" pageNumber="536" precision="925" value="116.583336">
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116°35
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<emphasis id="B924EA8EFF9AE8160E258DC12169FE23" box="[518,524,466,489]" italics="true" pageId="21" pageNumber="536">′</emphasis>
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E
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</geoCoordinate>
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||
). Collected by
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<location id="8E8F6047FF9AE8160EF08DC02386FDC2" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8160D068D2227A8FB8C:8E8F6047FF9AE8160EF08DC02386FDC2" country="Malaysia" county="State" latitude="6.0333333" longLatPrecision="1305" longitude="116.583336" municipality="Ranau District" name="Martinah" pageId="21" pageNumber="536" stateProvince="Sabah">Martinah</location>
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,
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<location id="8E8F6047FF9AE8160CD68DE12259FDC2" LSID="urn:lsid:plazi:treatment:03F9878AFF9AE8160D068D2227A8FB8C:8E8F6047FF9AE8160CD68DE12259FDC2" box="[245,316,498,520]" country="Malaysia" county="State" latitude="6.0333333" longLatPrecision="1305" longitude="116.583336" municipality="Ranau District" name="David" pageId="21" pageNumber="536" stateProvince="Sabah">David</location>
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, and Safrie. Date:
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<date id="FFEE105CFF9AE8160E058DE121F7FDCD" box="[550,658,498,519]" pageId="21" pageNumber="536" value="2007-05-11">
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<collectingDate id="EFAAE9B4FF9AE8160E058DE121F7FDCD" box="[550,658,498,519]" pageId="21" pageNumber="536" value="2007-05-11">11.v.2007</collectingDate>
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</date>
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,
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<typeStatus id="54EB883EFF9AE8160E808DE12063FDC2" box="[675,774,498,520]" pageId="21" pageNumber="536" type="holotype">holotype</typeStatus>
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, SP 12914,
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<figureCitation id="136B2A19FF9AE8160D058E0322FEFDEC" box="[294,411,528,550]" captionStart="Figure 6" captionStartId="12.[144,223,1616,1635]" captionTargetBox="[306,1260,198,1584]" captionTargetId="figure-74@12.[303,1263,195,1586]" captionTargetPageId="12" captionText="Figure 6. Shells. A, Everettia lapidini sp. nov.; B, Everettia klemmantanica; C, Everettia subconsul; D, Everettia paulbasintali sp. nov.; E, Everettia themis; F, Everettia layanglayang sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443035" httpUri="https://zenodo.org/record/5443035/files/figure.png" pageId="21" pageNumber="536">Figure 6A</figureCitation>
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;
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<specimenCount id="9D56FD15FF9AE8160D888E022136FDEC" box="[427,595,529,550]" pageId="21" pageNumber="536" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
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, one in
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<specimenCode id="DBF69EE7FF9AE8160E9A8E022274FD8F" collectionCode="BMNH" pageId="21" pageNumber="536">BMNH 20080636</specimenCode>
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and one in BOR/MOL 5500
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6517FF9AE8160C878E7D27A8FB8C" pageId="21" pageNumber="536" type="etymology">
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<paragraph id="8BEF369CFF9AE8160C878E7D218FFD15" blockId="21.[164,779,622,735]" pageId="21" pageNumber="536">
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<emphasis id="B924EA8EFF9AE8160C878E7D2247FD49" box="[164,290,622,643]" italics="true" pageId="21" pageNumber="536">Etymology:</emphasis>
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This species is named after Johny Lapidin, a staff member at Kinabalu Park, who dedicatedly assisted the first author during the intensive malacological survey on Mount Kinabalu in 2005.
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</paragraph>
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<paragraph id="8BEF369CFF9AE8160C878F1B22B2FB23" blockId="21.[164,779,776,1257]" pageId="21" pageNumber="536">
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<emphasis id="B924EA8EFF9AE8160C878F1B22E7FCD7" box="[164,386,776,797]" italics="true" pageId="21" pageNumber="536">Material examined:</emphasis>
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<collectingRegion id="4994F87EFF9AE8160DA98F1B2286FCD7" box="[394,483,776,797]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">SABAH</collectingRegion>
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– Mount Kinabalu: BOR/ MOL 973, southern slope,
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<quantity id="4CA89B79FF9AE8160DC38F342151FCF6" box="[480,564,807,828]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="536" unit="m" value="1800.0">1800 m</quantity>
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||
alt.,
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<date id="FFEE105CFF9AE8160E558F3421D8FCF7" box="[630,701,807,829]" pageId="21" pageNumber="536" value="2001-01">i.2001</date>
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; BOR/ MOL 5320, southern slope,
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<quantity id="4CA89B79FF9AE8160E2D8F562104FC91" box="[526,609,837,859]" metricMagnitude="3" metricUnit="m" metricValue="2.244" pageId="21" pageNumber="536" unit="m" value="2244.0">2244 m</quantity>
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||
alt.,
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<date id="FFEE105CFF9AE8160E978F562063FC91" box="[692,774,837,859]" pageId="21" pageNumber="536" value="2005-04">iv.2005</date>
|
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; BOR/MOL 5321, southern slope,
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<quantity id="4CA89B79FF9AE8160E018F772113FCB3" box="[546,630,868,889]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="21" pageNumber="536" unit="m" value="2040.0">2040 m</quantity>
|
||
alt.,
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<date id="FFEE105CFF9AE8160E978F772063FCB0" box="[692,774,868,890]" pageId="21" pageNumber="536" value="2005-04">iv.2005</date>
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||
; BOR/MOL 5265, southern slope,
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<quantity id="4CA89B79FF9AE8160E038F912116FC52" box="[544,627,898,920]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="21" pageNumber="536" unit="m" value="1950.0">1950 m</quantity>
|
||
alt.,
|
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<date id="FFEE105CFF9AE8160E938F912063FC52" box="[688,774,898,920]" pageId="21" pageNumber="536" value="2003-09">ix 2003</date>
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; SP 12121, southern slope, Kiau view trail,
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<date id="FFEE105CFF9AE8160E948FB22063FC7D" box="[695,774,929,951]" pageId="21" pageNumber="536" value="2002-02">ii.2002</date>
|
||
; SP12460, southern slope,
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<quantity id="4CA89B79FF9AE8160DC18FD32153FC1F" box="[482,566,960,981]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="536" unit="m" value="1900.0">1900 m</quantity>
|
||
alt.,
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<date id="FFEE105CFF9AE8160EA38FD321B2FC1C" box="[640,727,960,982]" pageId="21" pageNumber="536" value="2005-03">iii.2005</date>
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||
; SP 12500, eastern slope, Mansaranam, Kg. Pahu,
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<date id="FFEE105CFF9AE8160E9F8FCD2062FC3E" box="[700,775,990,1012]" pageId="21" pageNumber="536" value="2005-05">v.2005</date>
|
||
; SP12916, southern slope,
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||
<quantity id="4CA89B79FF9AE8160E258FEE213CFBD8" box="[518,601,1021,1042]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="536" unit="m" value="1900.0">1900 m</quantity>
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alt.,
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<date id="FFEE105CFF9AE8160E9F8FEE2062FBD8" box="[700,775,1021,1042]" pageId="21" pageNumber="536" value="2007-05">v.2007</date>
|
||
; SP12924, southern slope, Marai-parai,
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<date id="FFEE105CFF9AE8160EA8880F21B3FBFB" box="[651,726,1052,1073]" pageId="21" pageNumber="536" value="2007-05">v.2007</date>
|
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; SP 13019, southern slope,
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<quantity id="4CA89B79FF9AE8160D8F88292165FB9A" box="[428,512,1082,1104]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="536" unit="m" value="1800.0">1800 m</quantity>
|
||
alt.,
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<date id="FFEE105CFF9AE8160E1F882921F6FB9A" box="[572,659,1082,1104]" pageId="21" pageNumber="536" value="2008-03">iii.2008</date>
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; SP12924, southern slope, Marai-parai,
|
||
<date id="FFEE105CFF9AE8160E65884A21F4FBA4" box="[582,657,1113,1134]" pageId="21" pageNumber="536" value="2007-05">v.2007</date>
|
||
; BMNH 20080202, southern slope,
|
||
<quantity id="4CA89B79FF9AE8160DD988642128FB47" box="[506,589,1143,1165]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="536" unit="m" value="1600.0">1600 m</quantity>
|
||
alt.,
|
||
<date id="FFEE105CFF9AE8160E8188642063FB47" box="[674,774,1143,1165]" pageId="21" pageNumber="536" value="1967-08">viii.1967</date>
|
||
; BMNH 20080203, southern slope,
|
||
<quantity id="4CA89B79FF9AE8160E51888521A0FB61" box="[626,709,1174,1195]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="21" pageNumber="536" unit="m" value="2000.0">2000 m</quantity>
|
||
alt.,
|
||
<date id="FFEE105CFF9AE8160C8788A7226CFB00" box="[164,265,1204,1226]" pageId="21" pageNumber="536" value="1967-08">viii.1967</date>
|
||
. Mount Trusmadi: BOR/MOL 1293, Waras cave,
|
||
<quantity id="4CA89B79FF9AE8160CC488C0225FFB23" box="[231,314,1235,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="536" unit="m" value="1600.0">1600 m</quantity>
|
||
alt.,
|
||
<date id="FFEE105CFF9AE8160D5988C022B7FB23" box="[378,466,1235,1257]" pageId="21" pageNumber="536" value="2002-03">iii.2002</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF9AE8160C878901221BFA68" blockId="21.[164,779,1298,1442]" pageId="21" pageNumber="536">
|
||
<emphasis id="B924EA8EFF9AE8160C87890122B2FAED" box="[164,471,1298,1319]" italics="true" pageId="21" pageNumber="536">Diagnostic characteristics:</emphasis>
|
||
The shell shape of this species is similar to
|
||
<taxonomicName id="4C504D1FFF9AE8160DA8892221CFFA8F" authorityName="COI sensu Gude" authorityYear="1918" box="[395,682,1328,1350]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="klemmantanica">
|
||
<emphasis id="B924EA8EFF9AE8160DA8892221CFFA8F" box="[395,682,1328,1350]" italics="true" pageId="21" pageNumber="536">Everettia klemmantanica</emphasis>
|
||
</taxonomicName>
|
||
, but the spiral striation above the shell characterizes this species, compared with the granule-like sculpture in
|
||
<taxonomicName id="4C504D1FFF9AE8160C87899E2212FA6B" authorityName="COI sensu Gude" authorityYear="1918" box="[164,375,1420,1442]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="klemmantanica">
|
||
<emphasis id="B924EA8EFF9AE8160C87899E2212FA6B" box="[164,375,1420,1442]" italics="true" pageId="21" pageNumber="536">E. klemmantanica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF9AE8160C8789DF26C7FDC4" blockId="21.[164,779,1483,1903]" lastBlockId="21.[826,1442,198,526]" pageId="21" pageNumber="536">
|
||
<emphasis id="B924EA8EFF9AE8160C8789DF2249FA2B" box="[164,300,1484,1505]" italics="true" pageId="21" pageNumber="536">Description:</emphasis>
|
||
Shell (
|
||
<figureCitation id="136B2A19FF9AE8160DA689D822BAFA2B" box="[389,479,1483,1505]" captionStart="Figure 6" captionStartId="12.[144,223,1616,1635]" captionTargetBox="[306,1260,198,1584]" captionTargetId="figure-74@12.[303,1263,195,1586]" captionTargetPageId="12" captionText="Figure 6. Shells. A, Everettia lapidini sp. nov.; B, Everettia klemmantanica; C, Everettia subconsul; D, Everettia paulbasintali sp. nov.; E, Everettia themis; F, Everettia layanglayang sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443035" httpUri="https://zenodo.org/record/5443035/files/figure.png" pageId="21" pageNumber="536">Figs 6A</figureCitation>
|
||
,
|
||
<figureCitation id="136B2A19FF9AE8160DCD89D82178FA2B" box="[494,541,1483,1505]" captionStart="Figure 11" captionStartId="18.[144,223,1209,1228]" captionTargetBox="[305,1263,196,1180]" captionTargetId="figure-241@18.[303,1263,196,1180]" captionTargetPageId="18" captionText="Figure 11. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia corrugata williamsi ssp. nov.; B, underside of E. corrugata williamsi ssp. nov.; C, underside of Everettia lapidini; D, upper surface of Everettia occidentalis sp. nov.; E, corrugation at the suture of Everettia corrugata corrugata; F, upper surface of Everettia jucundior sp. nov. Scale bar = 500 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443045" httpUri="https://zenodo.org/record/5443045/files/figure.png" pageId="21" pageNumber="536">11C</figureCitation>
|
||
): large, rather thin, yellowish brown. Spire moderately elevated, outer whorls shouldered below the suture. Periphery slightly shouldered to almost rounded, more distinctly angular in juveniles. Above the periphery, shell shiny with very weak (but distinct) oblique wrinkling, slightly stronger towards the suture and with densely, regularly placed spiral grooves,
|
||
<emphasis id="B924EA8EFF9AE8160E308AB12140F97D" box="[531,549,1698,1719]" italics="true" pageId="21" pageNumber="536">c.</emphasis>
|
||
7–8 per mm. Below the periphery, shell has fine, densely placed spiral grooves. Height up to
|
||
<quantity id="4CA89B79FF9AE8160D818ACD2162F93E" box="[418,519,1758,1780]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="21" pageNumber="536" unit="mm" value="16.6">16.6 mm</quantity>
|
||
; width up to 29.0 mm; diameter of the first three whorls 1.4–1.5, 1.0–1.1, and 2.0–
|
||
<quantity id="4CA89B79FF9AE8160D2B8B0F2238F8FB" box="[264,349,1820,1841]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="21" pageNumber="536" unit="mm" value="2.1">2.1 mm</quantity>
|
||
, respectively; number of whorls up to five and a half; height aperture up to
|
||
<quantity id="4CA89B79FF9AE8160E7B8B2921D9F89A" box="[600,700,1850,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="21" pageNumber="536" unit="mm" value="12.1">12.1 mm</quantity>
|
||
; width aperture up to 16.0 mm. Genitalia (
|
||
<figureCitation id="136B2A19FF9AE8160E8A8B4A219AF8A4" box="[681,767,1881,1903]" captionStart="Figure 7" captionStartId="13.[164,243,1755,1774]" captionTargetBox="[167,1442,198,1722]" captionTargetId="figure-0@13.[164,1444,194,1725]" captionTargetPageId="13" captionText="Figure 7. Genitalia. A, Everettia klemmantanica; B, Everettia lapidini sp. nov.; C, Everettia paulbasintali sp. nov.; D, Everettia subconsul; E, Everettia themis; F, Everettia layanglayang sp. nov.; G, Everettia jucundior sp. nov.; H, Everettia interior sp. nov.; I, Everettia corrugata corrugata." figureDoi="http://doi.org/10.5281/zenodo.5443037" httpUri="https://zenodo.org/record/5443037/files/figure.png" pageId="21" pageNumber="536">Fig. 7B</figureCitation>
|
||
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
|
||
<quantity id="4CA89B79FF9AE8160F198D1020EEFED2" box="[826,907,259,280]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="21" pageNumber="536" unit="mm" value="19.0">19 mm</quantity>
|
||
. The penis, dart-sac, and vagina with thick muscular walls. P, V arranged near the GO then followed by BC and DS where there is much space in the atrium between the openings of P + V and BC + DS. BC about two-thirds of total DS length. Animal (
|
||
<figureCitation id="136B2A19FF9AE8160FBC8D8F208AFE78" box="[927,1007,412,434]" captionStart="Figure 8" captionStartId="14.[144,223,1760,1779]" captionTargetBox="[147,1419,200,1728]" captionTargetId="figure-0@14.[144,1424,195,1730]" captionTargetPageId="14" captionText="Figure 8. Animal and mantle pigmentation. A, Everettia subconsul west coast lowland; B, Everettia layanglayang sp. nov.; C, Everettia monticola sp. nov.; D, Everettia safriei sp. nov.; E, Everettia interior sp. nov.; F, Everettia subconsul east coast lowland; G, Everettia corrugata corrugata; H, Everettia jucundior sp. nov.; I, Everettia klemmantanica; J, Everettia lapidini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443039" httpUri="https://zenodo.org/record/5443039/files/figure.png" pageId="21" pageNumber="536">Fig. 8J</figureCitation>
|
||
): the whole animal head is uniformly black. The mantle is black and covered by sparsely distributed tiny bright spots and followed by regularly spaced radial black bands at the last half whorl.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF9AE8160F198E3C2708FD48" blockId="21.[826,1441,559,642]" pageId="21" pageNumber="536">
|
||
<emphasis id="B924EA8EFF9AE8160F198E3C270CFD8E" box="[826,1129,559,580]" italics="true" pageId="21" pageNumber="536">Distribution and habitat:</emphasis>
|
||
Montane primary forest, from
|
||
<quantity id="4CA89B79FF9AE8160F5E8E5D275DFDA9" box="[893,1080,590,611]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="21" pageNumber="536" unit="m" value="1700.0" valueMax="2000.0" valueMin="1400.0">1400 to 2000 m</quantity>
|
||
alt.
|
||
<collectingRegion id="4994F87EFF9AE816085B8E5E27A6FDA9" box="[1144,1219,589,611]" country="Malaysia" name="Sabah" pageId="21" pageNumber="536">Sabah</collectingRegion>
|
||
: Mount Trusmadi, Mount Kinabalu (
|
||
<figureCitation id="136B2A19FF9AE81608288E7F2705FD48" box="[1035,1120,620,642]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="21" pageNumber="536">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF9AE8160F198EB027A8FB8C" blockId="21.[826,1441,675,1095]" pageId="21" pageNumber="536">
|
||
<emphasis id="B924EA8EFF9AE8160F198EB020C3FD72" box="[826,934,675,696]" italics="true" pageId="21" pageNumber="536">Remarks:</emphasis>
|
||
Spiral striation on the shell is the key character for this species. The animals usually have a uniform black head, but there is a rare case where a white stripe is present between the tentacles, which is similar to the pattern in
|
||
<taxonomicName id="4C504D1FFF9AE816085C8F0D2636FCF8" authorityName="COI sensu Gude" authorityYear="1918" box="[1151,1363,797,819]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="klemmantanica">
|
||
<emphasis id="B924EA8EFF9AE816085C8F0D2636FCF8" box="[1151,1363,797,819]" italics="true" pageId="21" pageNumber="536">E. klemmantanica</emphasis>
|
||
</taxonomicName>
|
||
. Some immature specimens have only a few spiral striations above the shell near the periphery, and are distinctly shouldered at the periphery. We believe that this species was first collected in 1929 on Mount Kinabalu (
|
||
<quantity id="4CA89B79FF9AE8160F618FA520FDFC06" box="[834,920,950,972]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="21" pageNumber="536" unit="m" value="1600.0">1600 m</quantity>
|
||
;
|
||
<bibRefCitation id="EFC14B6DFF9AE8160F8A8FA5279CFC07" author="Pendlebury HM & Chasen FN" box="[937,1273,950,973]" pageId="21" pageNumber="536" pagination="1 - 38" refId="ref25826" refString="Pendlebury HM, Chasen FN. 1932. A zoological expedition to Mt. Kinabalu, British North Borneo (1929). Journal of the Federated Malay States Museums 17: 1 - 38." type="journal article" year="1932">Pendlebury & Chasen, 1932</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="EFC14B6DFF9AE81609678FA520E7FC21" author="Laidlaw FF" pageId="21" pageNumber="536" pagination="177 - 189" refId="ref25632" refString="Laidlaw FF. 1937. Notes on land Mollusca from Mount Kinabalu, British North Borneo. The Bulletin of the Raffles Museum 13: 177 - 189." type="journal article" year="1937">Laidlaw (1937)</bibRefCitation>
|
||
mentioned that the shells were ‘definitely shouldered at the periphery, and like other Kinabalu specimens seem to me to differ from the
|
||
<typeStatus id="54EB883EFF9AE816096888002619FBE2" box="[1355,1404,1043,1064]" pageId="21" pageNumber="536">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C504D1FFF9AE8160F19882220CFFB8C" authorityName=", Laidlaw" authorityYear="1937" box="[826,938,1073,1094]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="536" phylum="Mollusca" rank="species" species="subconsul">
|
||
<emphasis id="B924EA8EFF9AE8160F19882220CFFB8C" box="[826,938,1073,1094]" italics="true" pageId="21" pageNumber="536">subconsul</emphasis>
|
||
</taxonomicName>
|
||
in the British Museum’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |