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<document id="67AC29407BC90CC9CCE33573C4C0E357" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno &amp; Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF95E8180FB48D2121F7FA53" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia monticola Liew &amp; Schilthuizen &amp; Vermeulen 2009, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="542" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="541" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="25EB579475E5112E35A6F72012BDF099">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:namePart id="F295171A168FEB0FB77665F14665415C">Liew, Thor-Seng</mods:namePart>
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<mods:namePart id="0E21C508E47D3FE18CE91033AA86993B">Schilthuizen, Menno</mods:namePart>
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<mods:namePart id="3443C480C3E7CD36F96AA8C113CB2481">Vermeulen, Jaap Jan</mods:namePart>
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<mods:title id="60FC515C64A204257FF9234DBE779B77">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="27FD0F8310021456E14180A6ED3E19F8">
<mods:date id="29D3C2109268661FBD27A62CEE67A926">2009</mods:date>
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<mods:number id="E3D8FF4CBC352EF1A88BCB1E178FD8B8">2009-10-26</mods:number>
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<mods:number id="1BCC97AB2CD116E17286A3A8DDF83688">157</mods:number>
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<mods:classification id="ADAE498D3F4DDCE128F90F7B4D4F3D6F">journal article</mods:classification>
<mods:identifier id="06EF549A8023355351AEDB95865619E8" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
<mods:identifier id="026D7311182849C77589CDB2F45102B4" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03F9878AFF95E8180FB48D2121F7FA53" ID-DOI="http://doi.org/10.5281/zenodo.5492205" ID-Zenodo-Dep="5492205" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8180FB48D2121F7FA53" httpUri="http://treatment.plazi.org/id/03F9878AFF95E8180FB48D2121F7FA53" lastPageId="27" lastPageNumber="542" pageId="26" pageNumber="541">
<subSubSection id="C34A6517FF95E8190FB48D212747FDEF" pageId="26" pageNumber="541" type="nomenclature">
<paragraph id="8BEF369CFF95E8190FB48D212679FE8C" blockId="26.[919,1308,305,327]" box="[919,1308,305,327]" pageId="26" pageNumber="541">
<heading id="D0A781F0FF95E8190FB48D212679FE8C" box="[919,1308,305,327]" centered="true" fontSize="9" level="2" pageId="26" pageNumber="541" reason="2">
<taxonomicName id="4C504D1FFF95E8190FB48D2127C8FE8C" ID-CoL="6HM83" authority="Liew &amp; Schilthuizen &amp; Vermeulen, 2009" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[919,1197,306,327]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="541" phylum="Mollusca" rank="species" species="monticola" status="sp. nov.">EVERETTIA MONTICOLA</taxonomicName>
<emphasis id="B924EA8EFF95E81908948D262679FE8C" bold="true" box="[1207,1308,305,326]" pageId="26" pageNumber="541">
<taxonomicNameLabel id="A21757F5FF95E81908948D262679FE8C" box="[1207,1308,305,326]" pageId="26" pageNumber="541" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF369CFF95E8190F058D4A2747FDEF" blockId="26.[806,1421,344,549]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190F058D4A2009FEA4" box="[806,876,345,366]" italics="true" pageId="26" pageNumber="541">Types:</emphasis>
<materialsCitation id="3B383CC1FF95E8190F548D4B277BFDEF" collectingDate="2003-10" collectionCode="BMNH" country="Malaysia" county="State" elevation="2900" latitude="6.05" location="Villosa Shelter'" longLatPrecision="1305" longitude="116.55" municipality="Kota Belud District" pageId="26" pageNumber="541" specimenCode="BMNH 20080634" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="F347760CFF95E8190F548D4B2765FEA7" box="[887,1024,344,365]" name="Malaysia" pageId="26" pageNumber="541">MALAYSIA</collectingCountry>
:
<collectingCounty id="628E4E10FF95E819082D8D4B272EFEA7" box="[1038,1099,344,365]" pageId="26" pageNumber="541">State</collectingCounty>
of
<collectingRegion id="4994F87EFF95E81908568D4B27A5FEA4" box="[1141,1216,344,366]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">Sabah</collectingRegion>
:
<collectingMunicipality id="6B8BACE6FF95E81908EC8D4B203FFE46" pageId="26" pageNumber="541">Kota Belud District</collectingMunicipality>
. Southern part of Kinabalu Park. Near
<location id="8E8F6047FF95E819091C8D6520E5FE61" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8180FB48D2121F7FA53:8E8F6047FF95E819091C8D6520E5FE61" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.55" municipality="Kota Belud District" name="Villosa Shelter'" pageId="26" pageNumber="541" stateProvince="Sabah">Villosa Shelter</location>
at
<quantity id="4CA89B79FF95E8190F978D86276DFE61" box="[948,1032,405,427]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="26" pageNumber="541" unit="m" value="2900.0">
<elevation id="007DD1AFFF95E8190F978D86276DFE61" box="[948,1032,405,427]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="26" pageNumber="541" unit="m" value="2900.0">2900 m</elevation>
</quantity>
alt. along the
<location id="8E8F6047FF95E81908EE8D86264CFE61" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8180FB48D2121F7FA53:8E8F6047FF95E81908EE8D86264CFE61" box="[1229,1321,405,427]" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.55" municipality="Kota Belud District" name="Summit" pageId="26" pageNumber="541" stateProvince="Sabah">Summit</location>
trail of
<location id="8E8F6047FF95E8190F058DA72094FE00" LSID="urn:lsid:plazi:treatment:03F9878AFF95E8180FB48D2121F7FA53:8E8F6047FF95E8190F058DA72094FE00" box="[806,1009,436,458]" country="Malaysia" county="State" latitude="6.05" longLatPrecision="1305" longitude="116.55" municipality="Kota Belud District" name="Mount Kinabalu" pageId="26" pageNumber="541" stateProvince="Sabah">Mount Kinabalu</location>
(
<geoCoordinate id="EE64505BFF95E819082C8DA72735FE03" box="[1039,1104,434,458]" degrees="6" direction="north" minutes="3" orientation="latitude" pageId="26" pageNumber="541" precision="925" value="6.05">
6°3
<emphasis id="B924EA8EFF95E81908178DA1275FFE03" box="[1076,1082,434,457]" italics="true" pageId="26" pageNumber="541"></emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE64505BFF95E81908498DA727B4FE03" box="[1130,1233,434,458]" degrees="116" direction="east" minutes="33" orientation="longitude" pageId="26" pageNumber="541" precision="925" value="116.55">
116°33
<emphasis id="B924EA8EFF95E81908948DA127D8FE03" box="[1207,1213,434,457]" italics="true" pageId="26" pageNumber="541"></emphasis>
E
</geoCoordinate>
). Collected by Tachani. Date:
<date id="FFEE105CFF95E8190FCA8DC12753FE2D" box="[1001,1078,466,487]" pageId="26" pageNumber="541" value="2003-10">
<collectingDate id="EFAAE9B4FF95E8190FCA8DC12753FE2D" box="[1001,1078,466,487]" pageId="26" pageNumber="541" value="2003-10">x.2003</collectingDate>
</date>
,
<typeStatus id="54EB883EFF95E819086E8DC127D5FE22" box="[1101,1200,466,488]" pageId="26" pageNumber="541" type="holotype">holotype</typeStatus>
, BOR/MOL 2691,
<figureCitation id="136B2A19FF95E8190F058DE220CDFDCC" box="[806,936,497,519]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="26" pageNumber="541">Figure 12B</figureCitation>
;
<specimenCount id="9D56FD15FF95E8190F978DE22709FDCC" box="[948,1132,497,519]" pageId="26" pageNumber="541" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
, two in
<specimenCode id="DBF69EE7FF95E81908E58DE226E8FDCD" box="[1222,1421,497,519]" collectionCode="BMNH" pageId="26" pageNumber="541">BMNH 20080634</specimenCode>
and one in SP 13066
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6517FF95E8180F058E5C21F7FA53" lastPageId="27" lastPageNumber="542" pageId="26" pageNumber="541" type="etymology">
<paragraph id="8BEF369CFF95E8190F058E5C2747FD4E" blockId="26.[806,1421,591,644]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190F058E5C20C1FDAE" box="[806,932,591,612]" italics="true" pageId="26" pageNumber="541">Etymology:</emphasis>
This species is named
<taxonomicName id="4C504D1FFF95E81908E48E5C2652FDAE" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[1223,1335,591,612]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="541" phylum="Mollusca" rank="species" species="monticola">
<emphasis id="B924EA8EFF95E81908E48E5C2652FDAE" box="[1223,1335,591,612]" italics="true" pageId="26" pageNumber="541">monticola</emphasis>
</taxonomicName>
for its montane distribution.
</paragraph>
<paragraph id="8BEF369CFF95E8190F058EBE20ACF919" blockId="26.[806,1421,685,1747]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190F058EBE2766FD08" box="[806,1027,685,706]" italics="true" pageId="26" pageNumber="541">Material examined:</emphasis>
<collectingRegion id="4994F87EFF95E81908288EBE2701FD08" box="[1035,1124,685,706]" country="Malaysia" name="Sabah" pageId="26" pageNumber="541">SABAH</collectingRegion>
Mount Trusmadi: BOR/ MOL 1310,
<quantity id="4CA89B79FF95E8190F9D8EDF2774FD2B" box="[958,1041,716,738]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="541" unit="m" value="1600.0">1600 m</quantity>
alt.,
<date id="FFEE105CFF95E819087D8EDF27D0FD28" box="[1118,1205,716,738]" pageId="26" pageNumber="541" value="2002-03">iii.2002</date>
; BOR/MOL
<quantity id="4CA89B79FF95E81909728EDF2009FCCA" metricMagnitude="6" metricUnit="m" metricValue="1.13888" pageId="26" pageNumber="541" unit="m" value="1138880.0">1138, 880 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F938EF9209BFD35" box="[944,1022,746,767]" pageId="26" pageNumber="541" value="2001-10">x.2001</date>
. Crocker Range: BOR/MOL 1312, Ulu kimanis,
<quantity id="4CA89B79FF95E8190FEE8F1A2745FCD4" box="[973,1056,777,798]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="26" pageNumber="541" unit="m" value="1200.0">1200 m</quantity>
alt.,
<date id="FFEE105CFF95E81908468F1A27DFFCD5" box="[1125,1210,777,799]" pageId="26" pageNumber="541" value="2003-06">vi.2003</date>
; BOR/MOL 1313, Mahua,
<quantity id="4CA89B79FF95E8190FA78F3B20BDFCF7" box="[900,984,808,829]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="541" unit="m" value="1000.0">1000 m</quantity>
alt.,
<date id="FFEE105CFF95E81908318F34270FFCF7" box="[1042,1130,807,829]" pageId="26" pageNumber="541" value="2002-03">iii.2002</date>
; BOR/MOL 1131, Mahua,
<quantity id="4CA89B79FF95E8190F058F55201FFC96" box="[806,890,838,860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="541" unit="m" value="1000.0">1000 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F9A8F552761FC91" box="[953,1028,838,859]" pageId="26" pageNumber="541" value="2001-05">v.2001</date>
; BOR/MOL 1136, Mahua,
<quantity id="4CA89B79FF95E819091A8F5526E8FC96" box="[1337,1421,838,860]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="541" unit="m" value="1000.0">1000 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F438F7620DBFCB1" box="[864,958,869,891]" pageId="26" pageNumber="541" value="2001-07">vii.2001</date>
. Kinabatangan: BOR/MOL 1411, Batu Tomanggong kecil,
<date id="FFEE105CFF95E81908278F902732FC53" box="[1028,1111,899,921]" pageId="26" pageNumber="541" value="2002-04">iv.2002</date>
. Meliau Range: BOR/MOL
<quantity id="4CA89B79FF95E8190F058FB120CAFC7D" box="[806,943,930,952]" metricMagnitude="6" metricUnit="m" metricValue="3.21464" pageId="26" pageNumber="541" unit="m" value="3214640.0">3214, 640 m</quantity>
alt.,
<date id="FFEE105CFF95E8190FC48FB1272EFC72" box="[999,1099,930,952]" pageId="26" pageNumber="541" value="2004-08">viii.2004</date>
; BOR/MOL
<quantity id="4CA89B79FF95E81908F18FB1263EFC7D" box="[1234,1371,930,952]" metricMagnitude="6" metricUnit="m" metricValue="3.21565" pageId="26" pageNumber="541" unit="m" value="3215650.0">3215, 650 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F058FD320EEFC1C" box="[806,907,960,982]" pageId="26" pageNumber="541" value="2004-08">viii.2004</date>
. Mount Kinabalu: BOR/MOL 5361, southern slope,
<quantity id="4CA89B79FF95E8190F4C8FCC20A6FC3F" box="[879,963,991,1013]" metricMagnitude="3" metricUnit="m" metricValue="1.552" pageId="26" pageNumber="541" unit="m" value="1552.0">1552 m</quantity>
alt.,
<date id="FFEE105CFF95E81908238FCC2730FC3F" box="[1024,1109,991,1013]" pageId="26" pageNumber="541" value="2005-09">ix.2005</date>
; BOR/MOL 5364, southern slope,
<date id="FFEE105CFF95E8190F5E8FED20B1FBDE" box="[893,980,1022,1044]" pageId="26" pageNumber="541" value="2005-03">iii.2005</date>
; BOR/MOL 5368, southern slope,
<quantity id="4CA89B79FF95E8190F05880F201FFBF8" box="[806,890,1052,1074]" metricMagnitude="3" metricUnit="m" metricValue="2.1559999999999997" pageId="26" pageNumber="541" unit="m" value="2156.0">2156 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F9B880F276FFBF8" box="[952,1034,1052,1074]" pageId="26" pageNumber="541" value="2005-04">iv.2005</date>
; BOR/MOL 5369, southern slope,
<quantity id="4CA89B79FF95E8190F058828201FFB9A" box="[806,890,1083,1104]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="26" pageNumber="541" unit="m" value="1784.0">1784 m</quantity>
alt.,
<date id="FFEE105CFF95E8190FF78828274CFB9B" box="[980,1065,1083,1105]" pageId="26" pageNumber="541" value="2005-09">ix.2005</date>
; BOR/MOL 5374, southern slope,
<quantity id="4CA89B79FF95E8190F53884920A1FBA5" box="[880,964,1114,1135]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="26" pageNumber="541" unit="m" value="1700.0">1700 m</quantity>
alt.,
<date id="FFEE105CFF95E8190820884A2737FBA5" box="[1027,1106,1113,1135]" pageId="26" pageNumber="541" value="2005-02">ii.2005</date>
; BOR/MOL 5376, southern slope,
<date id="FFEE105CFF95E8190F5E886B20B1FB44" box="[893,980,1144,1166]" pageId="26" pageNumber="541" value="2005-03">iii.2005</date>
; BOR/MOL 5378, southern slope,
<quantity id="4CA89B79FF95E8190F058884201DFB66" box="[806,888,1175,1196]" metricMagnitude="3" metricUnit="m" metricValue="2.1119999999999997" pageId="26" pageNumber="541" unit="m" value="2112.0">2112 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F9988842768FB67" box="[954,1037,1175,1197]" pageId="26" pageNumber="541" value="2005-04">iv.2005</date>
; BOR/MOL 5365, western slope,
<quantity id="4CA89B79FF95E8190F0588A6201FFB01" box="[806,890,1205,1227]" metricMagnitude="3" metricUnit="m" metricValue="1.7759999999999998" pageId="26" pageNumber="541" unit="m" value="1776.0">1776 m</quantity>
alt.,
<date id="FFEE105CFF95E8190FFB88A62748FB01" box="[984,1069,1205,1227]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5366, western slope,
<quantity id="4CA89B79FF95E8190F5288C720A6FB23" box="[881,963,1236,1258]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="26" pageNumber="541" unit="m" value="1152.0">1152 m</quantity>
alt.,
<date id="FFEE105CFF95E819082688C7273FFB20" box="[1029,1114,1236,1258]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5370, western slope,
<quantity id="4CA89B79FF95E8190FA688E020BCFAC2" box="[901,985,1267,1288]" metricMagnitude="3" metricUnit="m" metricValue="2.48" pageId="26" pageNumber="541" unit="m" value="2480.0">2480 m</quantity>
alt.,
<date id="FFEE105CFF95E819086188E127F2FAC2" box="[1090,1175,1266,1288]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5371, western slope,
<quantity id="4CA89B79FF95E8190FD089022720FAED" box="[1011,1093,1297,1319]" metricMagnitude="3" metricUnit="m" metricValue="1.152" pageId="26" pageNumber="541" unit="m" value="1152.0">1152 m</quantity>
alt.,
<date id="FFEE105CFF95E81908808902279DFAED" box="[1187,1272,1297,1319]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5377, western slope,
<quantity id="4CA89B79FF95E8190812892327E1FA8F" box="[1073,1156,1328,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.496" pageId="26" pageNumber="541" unit="m" value="1496.0">1496 m</quantity>
alt.,
<date id="FFEE105CFF95E81908F189232643FA8C" box="[1234,1318,1328,1350]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BMNH 20080205, southern slope, Mesilau,
<date id="FFEE105CFF95E81908F6895D264EFAAE" box="[1237,1323,1358,1380]" pageId="26" pageNumber="541" value="1967-06">vi.1967</date>
; BMNH 20080206, southern slope, Mesilau,
<date id="FFEE105CFF95E8190917897E26ECFA49" box="[1332,1417,1389,1411]" pageId="26" pageNumber="541" value="1967-06">vi.1967</date>
; BMNH 20080207, southern slope, Mesilau,
<date id="FFEE105CFF95E8190917899826ECFA6B" box="[1332,1417,1419,1441]" pageId="26" pageNumber="541" value="1967-06">vi.1967</date>
. Mount Tambuyukon: BOR/MOL 5372, eastern slope,
<quantity id="4CA89B79FF95E8190F0589DA201DFA14" box="[806,888,1481,1502]" metricMagnitude="3" metricUnit="m" metricValue="1.144" pageId="26" pageNumber="541" unit="m" value="1144.0">1144 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F9889DB2774FA14" box="[955,1041,1480,1502]" pageId="26" pageNumber="541" value="2005-09">ix.2005</date>
; BOR/MOL 5373, eastern slope,
<quantity id="4CA89B79FF95E8190F0589F4201FFA37" box="[806,890,1511,1533]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="26" pageNumber="541" unit="m" value="1880.0">1880 m</quantity>
alt.,
<date id="FFEE105CFF95E8190FF889F42755FA37" box="[987,1072,1511,1533]" pageId="26" pageNumber="541" value="2005-09">ix.2005</date>
; BOR/MOL 5375, eastern slope,
<quantity id="4CA89B79FF95E8190F4E8A1520A5F9D1" box="[877,960,1542,1564]" metricMagnitude="3" metricUnit="m" metricValue="1.584" pageId="26" pageNumber="541" unit="m" value="1584.0">1584 m</quantity>
alt.,
<date id="FFEE105CFF95E8190FDA8A15272AF9D6" box="[1017,1103,1542,1564]" pageId="26" pageNumber="541" value="2005-09">ix.2005</date>
. Mount Nombuyukon: BOR/ MOL 5360, north-eastern slope,
<quantity id="4CA89B79FF95E81908828A362791F9F0" box="[1185,1268,1573,1594]" metricMagnitude="3" metricUnit="m" metricValue="1.204" pageId="26" pageNumber="541" unit="m" value="1204.0">1204 m</quantity>
alt.,
<date id="FFEE105CFF95E81909178A3726ECF9F0" box="[1332,1417,1572,1594]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5362, north-eastern slope,
<quantity id="4CA89B79FF95E81908DF8A502635F993" box="[1276,1360,1603,1625]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="26" pageNumber="541" unit="m" value="1070.0">1070 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F058A72201EF9BD" box="[806,891,1633,1655]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5363, north-eastern slope,
<quantity id="4CA89B79FF95E81909198A7126E8F9BD" box="[1338,1421,1634,1656]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="26" pageNumber="541" unit="m" value="1680.0">1680 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F408A9320D2F95C" box="[867,951,1664,1686]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5367, north-eastern slope,
<quantity id="4CA89B79FF95E8190F058A8C201FF97E" box="[806,890,1695,1717]" metricMagnitude="3" metricUnit="m" metricValue="1.68" pageId="26" pageNumber="541" unit="m" value="1680.0">1680 m</quantity>
alt.,
<date id="FFEE105CFF95E8190F9D8A8C2776F97F" box="[958,1043,1695,1717]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
; BOR/MOL 5379, north-eastern slope,
<date id="FFEE105CFF95E8190F4C8AAE20A0F919" box="[879,965,1725,1747]" pageId="26" pageNumber="541" value="2005-11">xi.2005</date>
.
</paragraph>
<paragraph id="8BEF369CFF95E8190F058AED27F8F8A5" blockId="26.[806,1421,1789,1903]" pageId="26" pageNumber="541">
<emphasis id="B924EA8EFF95E8190F058AED2734F8D8" box="[806,1105,1789,1811]" italics="true" pageId="26" pageNumber="541">Diagnostic characteristics:</emphasis>
This species is without distinct characteristics on the shell, except for the moderately elevated spire, in which it differs from other
<taxonomicName id="4C504D1FFF95E8190F058B4A2082F8A5" box="[806,999,1881,1903]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="541" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EA8EFF95E8190F058B4A20E8F8A4" box="[806,909,1881,1902]" italics="true" pageId="26" pageNumber="541">Everettia</emphasis>
species
</taxonomicName>
of similar size.
</paragraph>
<paragraph id="8BEF369CFF94E8180C878CD521C2FB99" blockId="27.[164,779,198,1107]" pageId="27" pageNumber="542">
<emphasis id="B924EA8EFF94E8180C878CD52249FF11" box="[164,300,198,219]" italics="true" pageId="27" pageNumber="542">Description:</emphasis>
Shell (
<figureCitation id="136B2A19FF94E8180DBF8CD52292FF11" box="[412,503,198,220]" captionStart="Figure 3" captionStartId="8.[144,223,1547,1566]" captionTargetBox="[305,1263,196,1518]" captionTargetId="figure-68@8.[303,1263,196,1518]" captionTargetPageId="8" captionText="Figure 3. Shell surface sculptures (scanning electron micrograph). A, upper surface of Everettia interior sp. nov.; B, upper surface of Everettia jucunda; C, upper surface of Everettia layanglayang sp. nov.; D, protoconch of E. layanglayang sp. nov.; E, underside of E. layanglayang sp. nov.; F, upper surface of Everettia dominiki sp. nov.; G, upper surface of Everettia monticola sp. nov.; H, upper surface of Everettia planispira sp. nov. Scale bar = 200 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443029" httpUri="https://zenodo.org/record/5443029/files/figure.png" pageId="27" pageNumber="542">Figs 3G</figureCitation>
,
<figureCitation id="136B2A19FF94E8180E318CD52127FF16" box="[530,578,198,220]" captionStart="Figure 12" captionStartId="24.[144,223,696,715]" captionTargetBox="[146,1423,199,666]" captionTargetId="figure-241@24.[144,1425,196,666]" captionTargetPageId="24" captionText="Figure 12. Shells. A, Everettia jasilini sp. nov.; B, Everettia monticola sp. nov.; C, Everettia dominiki sp. nov.; D, Everettia safriei sp. nov.; E, Everettia planispira sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443047" httpUri="https://zenodo.org/record/5443047/files/figure.png" pageId="27" pageNumber="542">12B</figureCitation>
): medium-sized, rather thin, brownish, spire moderately elevated, outer whorls shouldered below suture. Periphery rounded. Above the periphery, smooth but sometimes with irregularly placed oblique wrinkling that is slightly stronger towards the suture. Below the periphery, shell is with or without fine, shallow spiral grooves. Height up to 7.0 mm; width up to
<quantity id="4CA89B79FF94E8180E808D8F2063FE78" box="[675,774,412,434]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="27" pageNumber="542" unit="mm" value="11.9">11.9 mm</quantity>
; diameter of the first three whorls 0.7, 0.5, and 1.0 mm, respectively; number of whorls up to five; height aperture up to
<quantity id="4CA89B79FF94E8180D8E8DEB2161FDC7" box="[429,516,504,526]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="27" pageNumber="542" unit="mm" value="5.4">5.4 mm</quantity>
; width aperture up to
<quantity id="4CA89B79FF94E8180C878E052399FDE6" box="[164,252,534,556]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="27" pageNumber="542" unit="mm" value="6.5">6.5 mm</quantity>
. Genitalia (
<figureCitation id="136B2A19FF94E8180DAB8E052289FDE7" box="[392,492,534,557]" captionStart="Figure 13" captionStartId="24.[144,223,1326,1345]" captionTargetBox="[148,1422,795,1294]" captionTargetId="figure-279@24.[144,1424,792,1296]" captionTargetPageId="24" captionText="Figure 13. Genitalia. A, Everettia jasilini sp. nov.; B, Everettia dominiki sp. nov.; C, Everettia safriei sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443049" httpUri="https://zenodo.org/record/5443049/files/figure.png" pageId="27" pageNumber="542">Fig. 13B</figureCitation>
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
<quantity id="4CA89B79FF94E8180E268E47212CFDA3" box="[517,585,596,617]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="27" pageNumber="542" unit="mm" value="7.0">7 mm</quantity>
. The penis and dart-sac have thin muscular walls, and vagina has very thin transparent wall. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the openings of each of the genital structures. BC long, almost same length as DS. Animal (
<figureCitation id="136B2A19FF94E8180D638F1822F0FCEB" box="[320,405,779,801]" captionStart="Figure 8" captionStartId="14.[144,223,1760,1779]" captionTargetBox="[147,1419,200,1728]" captionTargetId="figure-0@14.[144,1424,195,1730]" captionTargetPageId="14" captionText="Figure 8. Animal and mantle pigmentation. A, Everettia subconsul west coast lowland; B, Everettia layanglayang sp. nov.; C, Everettia monticola sp. nov.; D, Everettia safriei sp. nov.; E, Everettia interior sp. nov.; F, Everettia subconsul east coast lowland; G, Everettia corrugata corrugata; H, Everettia jucundior sp. nov.; I, Everettia klemmantanica; J, Everettia lapidini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443039" httpUri="https://zenodo.org/record/5443039/files/figure.png" pageId="27" pageNumber="542">Fig. 8C</figureCitation>
): black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles and extends backward to the end of the mantle and downward to the foot fringe, although in some animals, the head around the tentacles is uniformly black but a white band appears near the end of mantle. Other parts of the animal are covered by regularly spaced oblique dotted belts or a grey smear. The mantle is brownish or greyish and in the last half whorl with sparse irregular black and bright markings.
</paragraph>
<paragraph id="8BEF369CFF94E8180C8788972116FB3C" blockId="27.[164,779,1156,1270]" pageId="27" pageNumber="542">
<emphasis id="B924EA8EFF94E8180C8788972286FB53" box="[164,483,1156,1177]" italics="true" pageId="27" pageNumber="542">Distribution and habitat:</emphasis>
Tropical lowland and montane primary forest
<quantity id="4CA89B79FF94E8180DDE88B021E2FB72" box="[509,647,1187,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="2.5" metricValueMin="0.7" pageId="27" pageNumber="542" unit="m" value="1600.0" valueMax="2500.0" valueMin="700.0">7002500 m</quantity>
(common between
<quantity id="4CA89B79FF94E8180D2888D222C3FB1D" box="[267,422,1217,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="27" pageNumber="542" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">10001500 m</quantity>
).
<collectingRegion id="4994F87EFF94E8180D9888D12163FB12" box="[443,518,1218,1240]" country="Malaysia" name="Sabah" pageId="27" pageNumber="542">Sabah</collectingRegion>
: Crocker Range, Trusmadi Range, Mount Kinabalu (
<figureCitation id="136B2A19FF94E8180E3288F32102FB3C" box="[529,615,1248,1270]" captionStart="Figure 9" captionStartId="15.[164,243,1622,1641]" captionTargetBox="[325,1281,196,1590]" captionTargetId="figure-58@15.[323,1283,194,1592]" captionTargetPageId="15" captionText="Figure 9. Distribution maps (Mount Kinabalu and Mount Tambuyukon). A, Everettia monticola sp. nov. and Everettia safriei sp. nov.; B, Everettia planispira sp. nov., Everettia jasilini sp. nov., and Everettia corrugata corrugata; C, Everettia dominiki sp. nov.; D, Everettia themis, Everettia subconsul, Everettia corrugata williamsi ssp. nov. and Everettia layanglayang sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443041" httpUri="https://zenodo.org/record/5443041/files/figure.png" pageId="27" pageNumber="542">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BEF369CFF94E8180C87893421F7FA53" blockId="27.[164,779,1319,1433]" pageId="27" pageNumber="542">
<emphasis id="B924EA8EFF94E8180C878934226AFAF6" box="[164,271,1319,1340]" italics="true" pageId="27" pageNumber="542">Remarks:</emphasis>
The shell of this species is similar to
<taxonomicName id="4C504D1FFF94E8180C8789542279FA91" baseAuthorityName="Gude" baseAuthorityYear="1917" box="[164,284,1350,1372]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="542" phylum="Mollusca" rank="species" species="robusta">
<emphasis id="B924EA8EFF94E8180C8789542279FA91" box="[164,284,1350,1372]" italics="true" pageId="27" pageNumber="542">E. robusta</emphasis>
</taxonomicName>
, which was described from Borneo (
<bibRefCitation id="EFC14B6DFF94E8180EE4895523BBFAB1" author="Gude GK" pageId="27" pageNumber="542" pagination="315 - 317" refId="ref25247" refString="Gude GK. 1917. Diagnoses of four undescribed Helicoids from Borneo and Guam (Ladrones). Proceedings of the Malacological Society 12: 315 - 317." type="journal article" year="1917">Gude, 1917</bibRefCitation>
). However, it is impossible to determine where in
<collectingRegion id="4994F87EFF94E8180CE789902269FA53" box="[196,268,1411,1433]" country="Malaysia" name="Sabah" pageId="27" pageNumber="542">Sabah</collectingRegion>
(if at all) the
<typeStatus id="54EB883EFF94E8180D948997228DFA53" box="[439,488,1412,1433]" pageId="27" pageNumber="542">type</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>